Gurilin Tsav [HMNS-GIN]: Maastrichtian, Mongolia
collected by HMNS-GIN 1993–1995, 1998

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia
Reptilia indet. Laurenti 1768
Watabe et al. 2010 1 individual
"an enigmatic giant vertebrate with very porous bone histology"
Testudinata indet. (Oppel 1811)
Watabe et al. 2010
Reptilia - Testudines - Lindholmemydidae
Suzuki and Narmandakh 2004 2 specimens
Reptilia - Testudines - Trionychidae
Watabe and Suzuki 2000
    = Trionychidae indet. Gray 1825
Danilov et al. 2014
Incomplete postcranium of one individual, including nuchal, proximal part of costal 1, fragment of right hyo- and hypoplastron, fragment of xiphiplastron, and assorted non-shell bones (IPGAS 11-5-1, holotype of Amyda menneri)
Amyda menneri n. sp. Chkhikvadze 1988
Danilov and Vitek 2013
nomen dubium belonging to Trionychidae
Reptilia
Mongolochelys sp. Khosatzky 1997
8 specimens
field no. 940829 GT SZK
    = Mongolochelys efremovi Khosatzky 1997
Suzuki and Tsogtbaatar 2010
MPC 25/106: shell fragments, R & L distal end of humeri (= HMNS 94-10-131); MPC 25/107: shell fragments (= HMNS 94-10-132); MPC 25/113: R humerus (= HMNS 94-10-135); MPC 25/114: shell fragments, R proximal end humerus, R ischium proximal, L femur proximal, R tibia proximal & distal, R fibula proximal (= HMNS 94-10-136); MPC 25/115: skull w/lower jaws (= HMNS 94-10-134); MPC 25/116: almost complete carapace & plastron, R & L pectoral girdles (= HMNS 94-10-25); MPC 25/141: skull fragments, R scapula & distal end of humerus (= HMNS 2006-04-489); MPC 25/142: peripheral, L epiplastron, 3rd & 7th cervicals, 1 caudal, R distal end of radius & proximal end of ulna & 5 phalanges (= HMNS 2006-04-490); MPC 25/143: braincase, 3-4th neurals, pelvic girdle (= HMNS 2006-04-491); MPC 25/144: braincase, 5 caudals, 12 phalanges (= HMNS 2006-04-492); MPC 25/146: lower jaw fragments, 1 caudal (= HMNS 2006-04-495)
Crocodylia indet. (Owen 1842)
Watabe et al. 2010
Dinosauria indet. (Owen 1842)
Suzuki and Watabe 2000
dinosaur eggs and shells; field no. 980808 GT SDV
Reptilia - Ankylosauridae
Ankylosauridae indet. Brown 1908
Watabe and Suzuki 2000 1 individual
Pelvic parts
Reptilia
Pachycephalosauria indet. (Maryańska and Osmólska 1974)
Watabe and Suzuki 2000 1 specimen
skull, MPC-D 100/1205 (Field no. 940828 GT WTB)
    = Prenocephale prenes Maryańska and Osmólska 1974
Evans et al. 2018
Reptilia - Hadrosauridae
Hadrosauridae indet. Cope 1869
Watabe and Suzuki 2000 6 specimens
isolated maxilla, skull fragments; skin impression; field no. 980808 GT WTB
    = ? Saurolophus sp. Brown 1912
Suzuki and Watabe 2000
    = Saurolophus angustirostris Rozhdestvensky 1952
Tsogtbaatar and Chinzorig 2007
incl. isolated bones; 060820 GT BD M-20 to M-23; 940829 GT TSOGT
Reptilia
Theropoda indet. (Marsh 1881)
Watabe and Suzuki 2000 2 individuals
"small theropod digit", skull and vertebrae
Reptilia - Oviraptoridae
Oviraptorosauria indet. Barsbold 1976
Suzuki and Watabe 2000 2 specimens
two new oviraptorosaurians, same species and similar to that found at Bugin Tsav in 1994; field no. 980808 GT COY Oviraptr, SZK Oviraptr
    = Heyuanninae indet. Yun 2019
Funston 2020
    = Oksoko avarsan Funston et al. 2020
Funston et al. 2020
Reptilia - Ornithomimidae
Ornithomimidae indet. Marsh 1890
1 specimen
pubis + ilium
    = Gallimimus sp. Osmólska et al. 1972
Matsumoto et al. 2010
980809 GT GU
? Gallimimus sp. Osmólska et al. 1972
Watabe and Suzuki 2000 1 specimen
long tibia
Reptilia - Tyrannosauridae
Tarbosaurus sp. Maleev 1955
Watabe and Suzuki 2000 1 specimen
partial skeleton with gastroliths; field no. 980808 GT ULZ Tarbo
Reptilia
Sauropoda indet. (Marsh 1878)
Suzuki and Watabe 2000 1 individual
ilium and associated vertebrae
? Pterosauria indet. (Kaup 1834)
Matsumoto et al. 2010 1 specimen
20060820 GT MB
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 43.9° North, 100.1° East (view map)
Paleocoordinates:41.4° North, 89.1° East
Basis of coordinate:stated in text
Altitude:1033
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
Key time interval:Maastrichtian
Age range of interval:72.10000 - 66.00000 m.y. ago
Stratigraphy
Formation:Nemegt
Stratigraphic resolution:bed
Lithology and environment
Primary lithology:current ripples,fine,white,blue sandstone
Secondary lithology:paleosol/pedogenic,red mudstone
Includes fossils?Y
Lithology description: some fossils in "the bluish white fine sandstone layers, overlying the red mudstone.""Alternation of sandstone and mudstone layers is its dominant lithology. The paleosol horizons are very frequently observed in the beds."
Environment:fluvial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:surface (in situ),mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:HMNS-GIN Collection dates:1993–1995, 1998
Collection method comments: HMNS-Hayashibara Museum of Natural Sciences
Metadata
Also known as:Gurlin Tsav, Guriliin Tsav
Database number:127670
Authorizer:R. Benson, P. Holroyd, P. Mannion, M. Carrano Enterer:R. Benson, P. Holroyd, J. Tennant, M. Carrano
Modifier:M. Carrano Research group:vertebrate
Created:2012-05-21 22:47:51 Last modified:2023-10-12 15:22:35
Access level:the public Released:2012-05-21 22:47:51
Creative Commons license:CC BY
Reference information

Primary reference:

42289. M. Watabe and S. Suzuki. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1993. Hayashibara Museum of Natural Sciences Research Bulletin 1:17-29 [R. Benson/R. Benson/M. Carrano]

Secondary references:

50209 I. G. Danilov, R. Hirayama, V. B. Sukhanov, S. Suzuki, M. Watabe and N. S. Vitek. 2014. Cretaceous soft-shelled turtles (Trionychidae) of Mongolia: new diversity, records, and a revision. Journal of Systematic Palaeontology 12(7):799-832 [P. Mannion/J. Tennant/P. Mannion]
64230 I. G. Danilov and N. S. Vitek. 2013. Cretaceous Trionychids of Asia: An Expanded Review of Their Record and Biogeography. In D. Brinkman, P. A. Holroyd, J. Gardner (eds.), Morphology and Evolution of Turtles 419-438 [P. Holroyd/P. Holroyd/P. Holroyd]
86067 D. C. Evans, S. Hayashi, K. Chiba, M. Watabe, M. J. Ryan, Y.-N. Lee, P. J. Currie, K. Tsogtbaatar, and R. Barsbold. 2018. Morphology and histology of new cranial specimens of Pachycephalosauridae (Dinosauria: Ornithischia) from the Nemegt Formation, Mongolia. Palaeogeography, Palaeoclimatology, Palaeoecology 494:121-134 [M. Carrano/M. Carrano]
73939 G. F. Funston. 2020. Caenagnathids of the Dinosaur Park Formation (Campanian) of Alberta, Canada: anatomy, osteohistology, taxonomy, and evolution. Vertebrate Anatomy Morphology Palaeontology 8:105-153 [M. Carrano/M. Carrano]
74873 G. F. Funston, T. Chinzorig, K. Tsogtbaatar, Y. Kobayashi, C. Sullivan and P. J. Currie. 2020. A new two-fingered dinosaur sheds light on the radiation of Oviraptorosauria. Royal Society Open Science 7(104643):201184:1-15 [M. Carrano/M. Carrano/M. Carrano]
82301 Y. Matsumoto, R. Hashimoto, and T. Sonoda. 2000. Report of preparation works for Mongolian specimens in HMNS from March 1994 to December 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:113-127 [M. Carrano/M. Carrano]
82304 Y. Matsumoto, R. Hashimoto, T. Sonoda, Y. Fujiyama, B. Mifune, Y. Kawahara, and M. Saneyoshi. 2010. Report of the preparation works for Mongolian specimens in Hayashibara Museum of Natural Sciences: 1999–2008. Hayashibara Museum of Natural Sciences Research Bulletin 3:167-185 [M. Carrano/M. Carrano/M. Carrano]
41670 S. Suzuki and P. Narmandakh. 2004. Change of the Cretaceous turtle faunas in Mongolia. Hayashibara Museum of Natural Sciences Research Bulletin 2:7-14 [R. Benson/R. Benson]
44038 S. Suzuki and C. Tsogtbaatar. 2010. A catalog of Mongolochelys collected by the HMNS-MPC Joint Paleontological Expedition. Hayashibara Museum of Natural Sciences Research Bulletin 3:119-131 [R. Benson/R. Benson]
42293 S. Suzuki and M. Watabe. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:83-98 [R. Benson/R. Benson/M. Carrano]
80572 K. Tsogtbaatar and T. Chinzorig. 2007. Mongolian hadrosaurids: paleobiogeography, dispersal and environmental relationships. The Korean Journal of Quaternary Research 21(2):49-53 [M. Carrano/M. Carrano]
80151 K. Tsogtbaatar and T. Chinzorig. 2010. Fossil specimens prepared in Mongolian Paleontological Center: 2002–2008. Hayashibara Museum of Natural Sciences Research Bulletin 3:155-166 [M. Carrano/M. Carrano]
42154 M. Watabe and S. Suzuki. 2000. Cretaceous fossil localities and a list of fossils collected by the Hayashibara Museum of Natural Sciences and Mongolian Paleontological Center Joint Paleontological Expedition (JMJPE) from 1993 through 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:99-108 [R. Benson/R. Benson]
55060 M. Watabe, S. Suzuki, K. Tsogtbaatar, T. Tsubamoto, and M. Saneyoshi. 2010. Report of the HMNS-MPC Joint Paleontological Expedition in 2006. Hayashibara Museum of Natural Sciences Research Bulletin 3:11-18 [M. Carrano/M. Carrano]
55298 M. Watabe, K. Tsogtbaatar, S. Suzuki and M. Saneyoshi. 2010. Geology of dinosaur-fossil-bearing localities (Jurassic and Cretaceous: Mesozoic) in the Gobi Desert: Results of the HMNS-MPC Joint Paleontological Expedition. Hayashibara Museum of Natural Sciences Research Bulletin 3:41-118 [M. Carrano/M. Carrano]
42291 W. Watabe and S. Suzuki. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1994. Hayashibara Museum of Natural Sciences Research Bulletin 1:30-44 [R. Benson/R. Benson/M. Carrano]