Sud-Sacaco West (SAS) horizon: Messinian, Peru

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Amantis sp.
Stucchi and Urbina 2004
Gastropoda - Neogastropoda - Muricidae
Concholepas kieneri Hupé 1854
Stucchi and Urbina 2004
Acanthina triangularis DeVries 2003
Stucchi and Urbina 2004
Acanthina obesa DeVries 2003
Stucchi and Urbina 2004
Herminespina sp. DeVries and Vermeij 1997
Stucchi and Urbina 2004
Bivalvia - Arcida - Arcidae
Anadara sp. Gray 1847
Stucchi and Urbina 2004
Bivalvia - Cardiida - Veneridae
Dosinia sp. Scopoli 1777
Stucchi and Urbina 2004
Chondrichthyes - Myliobatiformes - Myliobatidae
Myliobatis sp. Cuvier 1816
Muizon 1984
Chondrichthyes - Carcharhiniformes - Carcharhinidae
Carcharhinus sp. de Blainville 1816
Muizon and DeVries 1985
Chondrichthyes - Lamniformes - Carchariidae
Odontaspis aff. taurus (Rafinesque 1810)
Muizon 1984
original and current combination Carcharias taurus
Chondrichthyes - Lamniformes - Lamnidae
Carcharodon carcharias (Linnaeus 1758)
Marocco and Muizon 1988
Carcharodon hubbelli n. sp. Ehret et al. 2012
Ehret et al. 2012 2 specimens
Chondrichthyes - Lamniformes - Otodontidae
Procarcharodon megalodon (Agassiz 1835)
Muizon 1984
recombined as Otodus megalodon
Carcharocles megalodon (Agassiz 1835)
Stucchi and Urbina 2004
recombined as Otodus megalodon
Mammalia - Cetacea - Ziphiidae
Ninoziphius platyrostris n. gen., n. sp. Muizon 1983
Muizon 1983
Mammalia - Cetacea - Kogiidae
Scaphokogia sp. Muizon 1988
Lambert and Muizon 2013 1 specimen
sp. nov.
Mammalia - Cetacea - Pontoporiidae
Pliopontos littoralis n. gen., n. sp. Muizon 1983
Mammalia - Cetacea
Delphinoidea indet. Flower 1865
Muizon 1984
Mammalia - Cetacea - Delphinidae
Globicephalinae indet. (Gray 1866)
Muizon 1984
Tursiops oligodon n. sp. Pilleri and Siber 1989
Pilleri and Siber 1989 1 specimen
recombined as Hemisyntrachelus oligodon
Mammalia - Cetacea - Phocoenidae
Phocoenidae indet. (Gray 1825)
Marocco and Muizon 1988
Piscolithax longirostris n. gen., n. sp. Muizon 1983
Muizon 1983
Mammalia - Cetacea - Odobenocetopsidae
Odobenocetops leptodon Muizon et al. 1999
Muizon et al. 1999 1 specimen
Odobenocetops peruvianus n. gen., n. sp. Muizon 1993
Muizon 1993
(1 measurement)
Mammalia - Cetacea - Cetotheriidae
Cetotheriidae indet. (Brandt 1872)
Marocco and Muizon 1988
Piscobalaena nana n. gen., n. sp. Pilleri and Siber 1989
Pilleri and Siber 1989 1 specimen
(11 measurements)
Mammalia - Carnivora - Procyonidae
Cyonasua sp. Ameghino 1885
Tarquini et al. 2020
MNHN SAS 1625
Mammalia - Carnivora - Otariidae
Arctocephalus lomasiensis (Muizon 1978)
Pilleri 1990
recombined as Hydrarctos lomasiensis
Mammalia - Carnivora - Phocidae
Phocidae indet. (Gray 1821)
Marocco and Muizon 1988
original and current combination Phocadae
Monachinae indet. Trouessart 1897
Muizon 1984
form 2
Hadrokirus martini n. gen., n. sp. Amson and Muizon 2013
Amson and Muizon 2013 2 specimens
Piscophoca pacifica n. gen., n. sp. Muizon 1981
Muizon 1981
Acrophoca longirostris n. gen., n. sp. Muizon 1981
Muizon 1981
Mammalia - Megatheriidae
Megatheriidae indet. (Gray 1821)
Muizon 1984
misspelling Megatheriadae
Mammalia - Nothrotheriidae
Thalassocnus littoralis n. sp. McDonald and Muizon 2002
McDonald and Muizon 2002 4 specimens
Reptilia
Eusuchia sp. Huxley 1875
Muizon 1984
    = Piscogavialis jugaliperforatus Kraus 1998
Stucchi and Urbina 2004
MUSM 165
Sacacosuchus cordovai n. gen., n. sp. Salas-Gismondi et al. 2022
Salas-Gismondi et al. 2022
MUSM 162 - holotype; referred specimen: MUSM 160
Reptilia - Odontopterygiformes - Pelagornithidae
cf. Pelagornithidae indet. Fürbringer 1888
Muizon 1984
Pelagornis sp. Lartet 1857
Stucchi and Urbina 2004
Reptilia - Spheniscidae
Spheniscidae indet. Bonaparte 1831
Marocco and Muizon 1988
form 2
Spheniscus humboldti Meyen 1834
Stucchi and Urbina 2004
Spheniscus urbinai n. sp. Stucchi 2002
Stucchi 2002 4 specimens
Reptilia - Procellariidae
Procellariidae indet. Vigors 1825
Muizon 1984
Fulmarus sp. Stephens 1826
Cheneval 1993
Reptilia - Suliformes - Sulidae
cf. Sula sp. Brisson 1760
Muizon 1984
Sula sp. Brisson 1760
Stucchi and Urbina 2004
Ramphastosula ramirezi n. gen., n. sp. Stucchi and Urbina 2004
Stucchi and Urbina 2004 5 specimens
Ramphastosula aguierrei Stucchi et al. 2016
Stucchi et al. 2016 1 specimen
Reptilia - Suliformes - Phalacrocoracidae
cf. Phalacrocorax sp. Brisson 1760
Muizon 1984
Reptilia - Cathartiformes - Cathartidae
Vulturidae indet. Illiger 1811
Stucchi and Urbina 2004
spelled with current rank as Vulturinae
Actinopteri - Tetraodontiformes
Tetraodontiformes indet. Berg 1940
Muizon and DeVries 1985
Actinopteri - Tetraodontiformes - Balistidae
Balistidae indet.
Muizon 1984
Actinopteri - Scombriformes - Scombridae
Scombridae indet. (Rafinesque 1815)
Muizon 1984
Actinopteri - Scombriformes - Cybiidae
Cybiidae indet.
Muizon 1984
Actinopteri - Istiophoriformes - Xiphiidae
Xiphiidae indet. Swainson 1839
Muizon 1984
Actinopteri - Centropomidae
Centropomidae indet. Poey 1868
Muizon and DeVries 1985
Actinopteri - Siluriformes - Ariidae
Ariidae indet. Berg 1958
Muizon 1984
cf. Ariidae indet. Berg 1958
Muizon and DeVries 1985
Actinopteri - Clupeiformes - Clupeoidei
Clupeidae indet. Cuvier 1817
Stucchi and Urbina 2004
Alosinae indet.
Muizon 1984
Actinopteri - Acanthopteri - Sphyraenidae
Sphyraenidae indet.
Muizon 1984
see common names

Geography
Country:Peru
Coordinates: 15.6° South, 74.7° West (view map)
Paleocoordinates:15.8° South, 73.3° West
Basis of coordinate:stated in text
Geographic resolution:local area
Time
Period:Neogene Epoch:Miocene
Stage:Messinian 10 m.y. bin:Cenozoic 6
*Period:Tertiary *Epoch:Early/Lower Pliocene
Key time interval:Messinian
Age range of interval:7.24600 - 5.33300 m.y. ago
Age estimate:6.35 to 5.47 Ma (Sr isotope)
* legacy (obsolete) database fields
Stratigraphy
Formation:Pisco
Stratigraphic resolution:bed
Stratigraphy comments: See Muizon and de Vries (1985). Basal most Pliocene. A tuff layer from the Sacaco level, above the Sud-Sacaco level, has been dated radiometrically (K/Ar) to 3.9 Myr (de Muizon & Bellon, 1980, 1986). Ehret et al (2012) updated the age to Messinian (5.93 Sr isotopes) rather than Zanclean.
Lithology and environment
Primary lithology:diatomaceous siltstone
Includes fossils?Y
Environment:foreshore
Geology comments: SAS (Montemar) horizon. Shore environment with beaches and reefs exposed to marine currents. High sedimentation rate, shallow-to-deep shelf, upwelling
Taphonomy
Modes of preservation:body,original phosphate
Size of fossils:macrofossils
Preservation of anatomical detail:medium
Associated major elements:some
Collection methods and comments
Collection methods:field collection,observed (not collected)
Reason for describing collection:taxonomic analysis
Metadata
Also known as:SASw
Database number:13079
Authorizer:M. Uhen, P. Mannion Enterer:M. Uhen, N. Pyenson, P. Mannion
Modifier:P. Mannion Research group:vertebrate
Created:2002-01-18 12:36:08 Last modified:2022-05-12 06:48:06
Access level:the public Released:2002-01-18 12:36:08
Creative Commons license:CC BY
Reference information

Primary reference:

11891. C. Muizon. 1983. Pliopontos littoralis un nouveau Platanistidae Cetacea du Pliocene de la cote peruvienne. Comptes Rendus de l'Academie des Sciences Paris Serie II (296)1101-1104 [M. Uhen/N. Pyenson/N. Pyenson]

Secondary references:

48158 E. Amson and C. Muizon. 2013. A new durophagous phocid (Mammalia: Carnivora) from the late Neogene of Peru and considerations on monachine seals phylogeny. Journal of Systematic Palaeontology 12(5):523-548 [M. Uhen/V. Mason/M. Uhen]
40374 J. Cheneval. 1993. L'avifaune Mio-Pliocene de la Formation Pisco (Perou) etude prliminaire. Document Labratorie Geologique Lyon 125:85-95 [M. Uhen/M. Uhen]
30017 D. J. Ehret, B. J. MacFadden, and R. Salas-Gismondi. 2009. Caught in the act: Trophic interactions between a 4-million-year-old white shark (Charcharodon) and mysticete whale from Peru. Palaios 24:329-333 [M. Uhen/M. Uhen]
46232 D. J. Ehret, B. J. MacFadden, D. S. Jones, T. J. DeVries, D. A. Foster and R. Salas-Gismondi. 2012. Origin of the white shark Carcharodon (Lamniformes: Lamnidae) based on recalibration of the upper Neogene Pisco Formation of Peru. Palaeontology 55(6):1139-1153 [M. Uhen/M. Uhen]
27969 O. Lambert, G. Bianucci, and C. Muizon. 2008. A new stem-sperm whale (Cetacea, Odontoceti, Physeteroidea) from the Latest Miocene of Peru. Comptes Rendus Palevol 7(6):361-369 [M. Uhen/M. Uhen/J. Alroy]
46791 O. Lambert and C. Muizon. 2013. A new long-snouted species of the Miocene pontoporiid dolphin Brachydelphis and a review of the Mio-Pliocene marine mammal levels in the Sacaco Basin, Peru. Journal of Vertebrate Paleontology 33(3):709-721 [M. Uhen/M. Uhen]
12182 R. Marocco and C. d.e. Muizon. 1988. Los vertebrados del Neogeno de La Costa Sur del Perú: Ambiente sedimentario y condiciones de fosilización. Bulletin de l'Institut Frances d'Etudes Andines 17(2):105-117 [M. Uhen/M. Uhen/M. Uhen]
16792 H. G. McDonald and C. Muizon. 2002. The cranial anatomy of Thalassocnus (Xenarthra, Mammalia), a derived nothrothere from the Neogene of the Pisco Formation (Peru). Journal of Vertebrate Paleontology 22(2):349-365 [M. Uhen/M. Uhen/J. Alroy]
11888 C. Muizon. 1983. Un nouveau Phocoenidae (Cetacea) du Pliocene inferieur du Perou. Comptes Rendus de l'Academie des Sciences Paris Serie II (296)1203-1206 [M. Uhen/N. Pyenson/N. Pyenson]
11893 C. Muizon. 1993. Walrus-like feeding adaptation in a new cetacean from the Pliocene of Peru. Nature 365:745-748 [M. Uhen/N. Pyenson/N. Pyenson]
16910 C. Muizon. 1981. Les vertebres fossiles de la Formation Pisco (Perou). Premiere parti: Deux nouveaux Monachina (Phocidae, Mammalia) du Pliocene de Sud-Sacaco. Recherche sur les grandes civilisations Memoire 6:1-150 [M. Uhen/M. Uhen/M. Uhen]
11880 C. Muizon. 1983. Un Ziphiidae nouveau du Pliocene inferieur de Perou. Comptes rendus de l'Academic des Sciences Paris Serie II 297:85-88 [M. Uhen/N. Pyenson/M. Uhen]
15398 C. Muizon. 1984. Les vertebres fossiles de la Formation Pisco (Perou) II: Les Odontocetes (Cetacea, Mammalia) du Pliocene inferieur de Sud-Sacaco. Institut Francais d'Etudes Andines Editions Recherche sur les Civilizations Memoire 50:1-188 [M. Uhen/M. Uhen]
11895 C. Muizon. 1988. Les vertebres fossiles de la Formation Pisco (Perou). Troisieme partie: Les Odontocetes (Cetacea, Mammalia) du Miocene. Editions Recherche sur les Civilisations (78)1-244 [M. Uhen/N. Pyenson/M. Uhen]
16797 C. Muizon and T. J. DeVries. 1985. Geology and paleontology of late Cenozoic marine deposits in the Sacaco area (Peru). Geologische Rundschau 74(3):547-563 [M. Uhen/M. Uhen/M. Uhen]
15397 C. Muizon, D. P. Domning, and M. Parrish. 1999. Dimorphic tusks and adaptive strategies in a new species of walrus-like dolphin (Odobenocetopsidae) from the Pliocene of Peru. C. R. Academie des Sciences, Paris, Sciences de la terre et des planets 329:449-455 [M. Uhen/M. Uhen]
16788 C. Muizon and H. G. McDonald. 1995. An aquatic sloth from the Pliocene of Peru. Nature 375:224-227 [M. Uhen/M. Uhen]
81296 D. Ochoa, T. J. DeVries, K. Quispe, A. Barbosa-Espitia, R. Salas-Gismondi, D. A. Foster, R. Gonzales, S. Revillon, R. Berrospi, L. Pairazamán, J. Cardich, A. Perez, P. Romero, M. Urbina, and M. Carré. 2022. Age and provenance of the Mio-Pleistocene sediments from the Sacaco area, Peruvian continental margin. Journal of South American Earth Sciences 116:103799 [M. Uhen/M. Uhen/M. Uhen]
26378 G. Pilleri. 1990. Zweiter fund von Arctocephalus lomasiensis und Acrophoca longirostris (Pinnipedia) in der Pisco Formation Perus. Beiträge zur Paläontologie der Cetaceen und Pinnipedier der Pisco Formation Perus II:237-240 [M. Uhen/M. Uhen/J. Alroy]
43585% 10880G. Pilleri and O. Pilleri. 1989. Bartenwale aus der Pisco-Formation Perus. Beitrage zur Palaontologie der cetaceen Perus 11-38 [M. Uhen/M. Uhen/J. Alroy]
12814 G. Pilleri and H. J. Siber. 1989. Neuer Spattertiarer cetotherid (Cetacea, Mysticeti) aus der Pisco Formation Perus. Beitrage zur Palaontologie der Cetaceen Perus 109-115 [M. Uhen/M. Uhen/M. Uhen]
12842 G. Pilleri and H. J. Siber. 1989. Neuer delphinid (Cetacea, Odontoceti) aus der Pisco-Formation Perus. Beitrage zur Palaontologie der cetaceen Perus 167-175 [M. Uhen/M. Uhen/M. Uhen]
81650 R. Salas-Gismondi, D. Ochoa, S. Jouve, P. E. Romero, J. Cardich, A. Perez, T. DeVries, P. Baby, M. Urbina and M. Carré. 2022. Miocene fossils from the southeastern Pacific shed light on the last radiation of marine crocodylians. Proceedings of the Royal Society B 289:20220380:1-10 [P. Mannion/P. Mannion]
34468 M. Stucchi. 2002. Una nueva especie de Spheniscus (Aves: Spheniscidae) de la Formacion Pisco, Peru. Boletim de la Sociedad Geologica del Peru 94:17-24 [M. Uhen/M. Uhen/M. Uhen]
18990 M. Stucchi and M. Urbina. 2004. Ramphastosula (aves, Sulidae): a new genus from the early Pliocene of the Pisco Formation, Peru. Journal of Vertebrate Paleontology 24(4):974-978 [M. Uhen/M. Uhen]
59281 M. Stucchi, R. M. Varas-Malca, and M. Urbina-Schmitt. 2016. New Miocene sulid birds from Peru and considerations on their Neogene fossil record in the Eastern Pacific Ocean. Acta Palaeontologica Polonica 61(2):417-427 [M. Uhen/M. Uhen]
74387 J. Tarquini, L. H. Soibelzon, R. Salas-Gismondi and C. Muizon. 2020. Cyonasua (Carnivora, Procyonidae) from late Miocene of Peru shed light on the early dispersal of carnivorans in South America. Journal of Vertebrate Paleontology e1834406:1-9 [P. Mannion/P. Mannion]