Laguna del Hunco: quarry LH2: Ypresian, Argentina
collected by P. Wilf, N.R. Cuneo, K.R. Johnson, P. Puerta, M. A. Gandolfo 1999, 2002, 2004

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Polypodiopsida - Osmundaceae
Filicopsida informal TY001 leaf
1 specimen
    = Todea amissa Carvalho et al. 2013
Carvalho et al. 2013
Pteridopsida
Filicopsida informal TY003 leaf
2 specimens
synonym of Pteridopsida
Angiospermae - Laurales - Monimiaceae
Magnoliopsida informal TY027 leaf
1 specimen
    = Monimiophyllum callidentatum Knight and Wilf 2013
Knight and Wilf 2013
Angiospermae - Laurales - Lauraceae
Lauraceae informal TY084 leaf
40 specimens
Angiospermae - Proteales - Proteaceae
Lomatia occidentalis leaf Frenguelli 1943
9 specimens
(Berry) Frenguelli
Angiospermae - Fabales - Fabaceae
Fabaceae informal TY091 seed/fruit
2 specimens
Angiospermae - Fagales - Casuarinaceae
Gymnostoma informal TY051 seed/fruit
1 specimen
    = Gymnostoma spp. seed/fruit
Zamaloa et al. 2006
Angiospermae - Fagales - Fagaceae
Magnoliopsida informal "Tetracera" patagonica leaf
44 specimens
Berry
    = Castaneophyllum patagonicum Wilf et al. 2019
Wilf et al. 2019
Angiospermae - Malpighiales - Euphorbiaceae
Macaranga kirkjohnsonii Wilf et al. 2023
Wilf et al. 2023 2 specimens
MPEF‐Pb 7990a,b and 7991a,b
Angiospermae - Malvales
Malvales informal cf. Sterculiaceae TY023 leaf
13 specimens
Angiospermae - Brassicales - Akaniaceae
Akania patagonica leaf Gandolfo et al. 1988
3 specimens
Gandolfo, Dibbern and Romero
Angiospermae - Myrtales - Myrtaceae
Myrtaceae informal "Myrcia" deltoidea leaf
7 specimens
Engelhardt
Myrtaceae informal TY157 leaf
1 specimen
"Myrcia" chubutensis leaf
21 specimens
Berry
    = Eucalyptus frenguelliana leaf
Hermsen et al. 2012
Gandolfo and Zamaloa
Angiospermae
Monocotyledonae informal TY130 leaf
2 specimens
spelled with current rank as Monocotyledoneae
Magnoliopsida
Magnoliopsida informal ? Fabaceae TY017 leaf
3 specimens
Magnoliopsida informal "Schmidelia" proedulis leaf
66 specimens
Berry
Magnoliopsida informal "Celtis" ameghenoi leaf
141 specimens
Berry
Magnoliopsida informal TY022 leaf
9 specimens
Magnoliopsida informal "Rhamnidium" leaf
6 specimens
Magnoliopsida informal TY028 leaf
4 specimens
Magnoliopsida informal TY030 leaf
3 specimens
Magnoliopsida informal "Banara" prehernandiensis leaf
4 specimens
as figured by Berry (1938)
Magnoliopsida informal TY036 leaf
1 specimen
Magnoliopsida informal TY037 leaf
1 specimen
Magnoliopsida informal TY038 leaf
13 specimens
Magnoliopsida informal ?Escallonia TY043 leaf
10 specimens
Magnoliopsida informal TY045 leaf
3 specimens
Magnoliopsida informal cf. Macaranga or Mallotus leaf
2 specimens
Magnoliopsida informal TY049 fertile axis
1 specimen
Magnoliopsida informal TY050 leaf
2 specimens
Magnoliopsida informal TY054 leaf
3 specimens
Magnoliopsida informal TY056 leaf
17 specimens
Magnoliopsida informal "Myrica" mira leaf
4 specimens
Berry
Magnoliopsida informal TY067 leaf
1 specimen
Magnoliopsida informal TY070 leaf
2 specimens
Magnoliopsida informal "Sterculia" patagonica leaf
1 specimen
Berry
Magnoliopsida informal ?Myrtaceae TY075 leaf
3 specimens
Magnoliopsida informal TY076 leaf
1 specimen
Magnoliopsida informal TY077 leaf
1 specimen
Magnoliopsida informal TY078 leaf
1 specimen
Magnoliopsida informal TY083 leaf
1 specimen
Magnoliopsida informal TY085 fertile axis
1 specimen
Magnoliopsida informal TY093 seed/fruit
3 specimens
Magnoliopsida informal TY100 seed/fruit
10 specimens
Magnoliopsida informal TY102 leaf
1 specimen
Magnoliopsida informal TY103 leaf
1 specimen
Magnoliopsida informal "Cupania" grosseserrata leaf
7 specimens
(Engelhardt) Berry
Magnoliopsida informal "Coprosma" incerta leaf
7 specimens
Berry
Magnoliopsida informal TY108 leaf
1 specimen
Magnoliopsida informal TY111 leaf
1 specimen
Magnoliopsida informal TY113 leaf
5 specimens
Magnoliopsida informal "Cupania" latifolioides leaf
11 specimens
Berry
Magnoliopsida informal "Cassia" argentinensis leaf
98 specimens
Berry
Magnoliopsida informal TY122 leaf
3 specimens
Magnoliopsida informal TY126 leaf
2 specimens
Magnoliopsida informal TY135 leaf
1 specimen
Magnoliopsida informal TY136 leaf
1 specimen
Magnoliopsida informal ?Melastomataceae TY138 leaf
2 specimens
Magnoliopsida informal TY153 leaf
2 specimens
Magnoliopsida informal TY159 leaf
2 specimens
Magnoliopsida informal TY167 leaf
2 specimens
Magnoliopsida informal ? Lauraceae TY190 leaf
6 specimens
Magnoliopsida informal TY191 leaf
3 specimens
Magnoliopsida informal TY192 leaf
5 specimens
Magnoliopsida informal ?Menispermaceae TY193 leaf
1 specimen
Magnoliopsida informal cf. Sterculiaceae TY194 leaf
1 specimen
Magnoliopsida informal TY195 leaf
1 specimen
Magnoliopsida informal TY196 leaf
1 specimen
Magnoliopsida informal TY197 leaf
1 specimen
Magnoliopsida informal TY198 leaf
1 specimen
Coniferales - Cupressaceae
"Libocedrus" prechilensis leaf
2 specimens
Berry
    = Papuacedrus prechilensis leaf
Wilf et al. 2009
(Berry) Wilf et al.
Coniferales - Podocarpaceae
Podocarpaceae informal TY121 leaf
4 specimens
(small planated leaves)
    = Dacrycarpus puertae leaf
Wilf 2012
Wilf 2012
Podocarpus andiniformis leaf
4 specimens
Berry
Equisetopsida - Araucariaceae
"Zamia" tertiaria leaf
2 specimens
Engelhardt
    = Agathis zamunerae Wilf et al. 2014
Wilf et al. 2014
Araucaria informal TY013 seed/fruit
3 specimens
(seed and cone scale)
see common names

Geography
Country:Argentina State/province:Chubut
Coordinates: 42.5° South, 70.0° West (view map)
Paleocoordinates:45.0° South, 60.0° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Paleogene Epoch:Eocene
Stage:Ypresian 10 m.y. bin:Cenozoic 2
Key time interval:Ypresian
Age range of interval:56.00000 - 47.80000 m.y. ago
Age estimate:52.22 ± 0.22 Ma (Ar/Ar)
Stratigraphy
Formation:Huitrera
Stratigraphic resolution:bed
Stratigraphy comments: See P. Wilf et al. 2003, Science suppl., for detailed lithostratigraphy, paleomagnetic stratigraphy, and first Ar-Ar ages. See P. Wilf et al. 2012 (American Journal of Botany) for the revised Ar-Ar age given above.
Lithology and environment
Primary lithology:fine,tuffaceous,gray lithified silty,sandy "siliciclastic"
Includes fossils?Y
Lithology description: See Wilf et al. 2003, Science suppl., for complete lithologic summary.
Environment:crater lake Tectonic setting:volcanic basin
Geology comments: Large caldera-lake, embedded in the Eocene middle Chubut River volcanic pyroclastic complex. See E. Aragon and M.M. Mazzon, Rev. Asoc. Geol. Argent. 52(3):243-256 (1997).
Taphonomy
Modes of preservation:adpression,original silica,original carbon,replaced with silica
Size of fossils:macrofossils,mesofossils,microfossils
Spatial orientation:random
Preservation of anatomical detail:very poor
Abundance in sediment:abundant
Size sorting:medium
Fragmentation:frequent
Bioerosion:none
Feeding/predation traces:punctures,external foliage feeding,arthropod mining,arthropod galling,arthropod boring,piercing/sucking,oviposition
Temporal resolution:time-averaged
Spatial resolution:allochthonous
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:bulk,selective quarrying,mechanical,field collection
Reason for describing collection:paleoecologic analysis
Museum repositories:MEF
Collectors:P. Wilf, N.R. Cuneo, K.R. Johnson, P. Puerta, M. A. Gandolfo Collection dates:1999, 2002, 2004
Metadata
Also known as:LH2
Database number:133043
Authorizer:P. Wilf, E. Vlachos Enterer:M. Donovan, E. Stiles, F. Aspromonte
Modifier:E. Stiles Research group:paleobotany
Created:2012-09-01 01:02:16 Last modified:2018-11-02 09:16:15
Access level:the public Released:2012-09-01 01:02:16
Creative Commons license:CC BY
Reference information

Primary reference:

42725. P. Wilf, K. R. Johnson, N. R. Cúneo, M. E. Smith, B. S. Singer and M. A. Gandolfo. 2005. Eocene Plant Diversity at Laguna del Hunco and Río Pichileufú, Patagonia, Argentina. The American Naturalist 165(6):634-650 [P. Wilf/M. Donovan]

Secondary references:

55786 M. R. Carvalho, P. Wilf, E. J. Hermsen, M. A. Gandolfo, N. R. Cúneo and K. R. Johnson. 2013. First record of Todea (Osmundaceae) in South America, from the early Eocene paleorainforests of Laguna del Hunco (Patagonia, Argentina). American Journal of Botany 100:1831-1848 [C. Jaramillo/C. Jaramillo]
43002 E. J. Hermsen, M. A. Gandolfo, and M. C. Zamaloa. 2012. The fossil record of Eucalyptus in Patagonia. American Journal of Botany 99:1356-1374 [P. Wilf/M. Donovan]
56100 C. Knight and P. Wilf. 2013. Rare leaf fossils of Monimiaceae and Atherospermataceae (Laurales) from Eocene Patagonian rainforests and their biogeographic significance. Palaeontologia Electronica 16(3):26A-39 p. [C. Jaramillo/C. Jaramillo]
43004 P. Wilf. 2012. Rainforest conifers of Eocene Patagonia: attached cones and foliage of the extant southeast-Asian and Australasian genus Dacrycarpus (Podocarpaceae). American Journal of Botany 99:562-584 [P. Wilf/M. Donovan]
55856 P. Wilf, I. H. Escapa, N. R. Cúneo, R. M. Kooyman, K. R. Johnson and A. Iglesias. 2014. First South American Agathis (Araucariaceae), Eocene of Patagonia. American Journal of Botany 101(1):156-179 [C. Jaramillo/C. Jaramillo]
87948 P. Wilf, A. Iglesias, and M. A. Gandolfo. 2023. The first Gondwanan Euphorbiaceae fossils reset the biogeographic history of the Macaranga‐Mallotus clade. American Journal of Botany 110(5):1-20 [E. Vlachos/F. Aspromonte]
43000 P. Wilf, S. A. Little, A. Iglesias, M. C. Zamaloa, M. A. Gandolfo, N. R. Cúneo, and K. R. Johnson. 2009. Papuacedrus (Cupressaceae) in Eocene Patagonia, a new fossil link to Australasian rainforests. American Journal of Botany 96:2031-2047 [P. Wilf/M. Donovan]
69525 P. Wilf, K. C. Nixon, M. A. Gandolfo and N. R. Cúneo. 2019. Eocene Fagaceae from Patagonia and Gondwanan legacy in Asian rainforests. Science 364(6444):eaaw5139 [P. Wilf/E. Stiles]
43003 M. C. Zamaloa, M. A. Gandolfo, C. C. González, E. J. Romero, N. R. Cúneo and P. Wilf. 2006. Casuarinaceae from the Eocene of Patagonia, Argentina. International Journal of Plant Sciences 167:1279-1289 [P. Wilf/M. Donovan]