PPP# 2187 - Escudo de Veraguas Island [Escudo de Veraguas Fm]: Piacenzian, Panama
collected by Jung et al 1996

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Umbilibalcis sp.
Jung 1989 1 specimen
Scaphopoda - Gadilida - Gadilidae
Gadilopsis sp. (Woodring 1925)
Jung 1989 87 specimens
Scaphopoda - Dentaliida - Dentaliidae
Dentalium sp. Linnaeus 1758
Jung 1989 4 specimens
Scaphopoda - Dentaliida - Gadilinidae
Episiphon sp. Pilsbry 1897
Jung 1989 11 specimens
Gastropoda - Heterostropha - Pyramidellidae
Eulimella (Ebalina) sp. (Thiele 1929)
Jung 1989 6 specimens
Chemnitzia sp. d'Orbigny 1840
Jung 1989 6 specimens
Callolongchaeus sp. Dall 1903
Jung 1989 2 specimens
Gastropoda - Heterostropha - Architectonicidae
Pseudotorinia sp. Sacco 1892
Jung 1989 1 specimen
Architectonica sp. Röding 1798
Jung 1989 3 specimens
Gastropoda - Heterostropha - Ringiculidae
Ringicula (Ringiculella) sp. (Sacco 1892)
Jung 1989 4 specimens
Gastropoda - Heterostropha - Acteonidae
Acteon sp. Montfort 1810
Jung 1989 3 specimens
Gastropoda - Pseudomelaniidae
Carinodrillia sp. (Dall 1919)
Jung 1989 2 specimens
Miraclathurella sp. Woodring 1928
Jung 1989 9 specimens
Gastropoda - Modulidae
Modulus sp. Gray 1842
Jung 1989 1 specimen
Gastropoda - Potamididae
Potamides sp. Brongniart 1810
Jung 1989 1 specimen
Gastropoda - Eulimidae
Niso sp. Risso 1826
Jung 1989 5 specimens
Eulima sp. Risso 1826
Jung 1989 1 specimen
Gastropoda - Cassidae
Sconsia striata (Lamarck 1816)
Jung 1989 3 specimens
replaced by Sconsia grayi
Gastropoda - Personidae
Distorsio clathrata (Lamarck 1816)
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Marginellidae
Volvarina sp. (Hinds 1844)
Jung 1989 4 specimens
Prunum sp. Swainson 1840
Jung 1989 8 specimens
Dentimargo sp. Cossmann 1899
Jung 1989 38 specimens
Gastropoda - Neogastropoda - Muricidae
Haustellum sp. Schumacher 1817
Jung 1989 2 specimens
Murexiella (Subpterynotus) sp. Olsson and Harbison 1953
Jung 1989 1 specimen
Siphonochelus (Laevityphis) sp. (Cossmann 1903)
Jung 1989 3 specimens
Gastropoda - Neogastropoda - Cystiscidae
Granulina sp. Jousseaume 1888
Jung 1989 54 specimens
Gastropoda - Neogastropoda - Mitridae
Subcancilla sp. Olsson and Harbison 1953
Jung 1989 9 specimens
Gastropoda - Neogastropoda - Olividae
Oliva (Oliva) sp. Bruguière 1789
Jung 1989 2 specimens
Olivella (Macgintiella) sp. Olsson 1956
Jung 1989 8 specimens
Olivella (Minioliva) sp. Olsson 1956
Jung 1989 84 specimens
Gastropoda - Neogastropoda - Nassariidae
Nassarius sp. Duméril 1806
Jung 1989 37 specimens
Gastropoda - Neogastropoda - Buccinidae
Antillophos (Antillophos) sp. Woodring 1928
Jung 1989 50 specimens
Gastropoda - Neogastropoda - Columbellidae
Sincola (Sinaxila) sp. Jung 1989
Jung 1989 14 specimens
Mitrella sp. Risso 1826
Jung 1989 2 specimens
Decipifus sp. Olsson and McGinty 1958
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Drilliidae
Syntomodrillia sp. Woodring 1928
Jung 1989 7 specimens
Agladrillia (Agladrillia) sp. Woodring 1928
Jung 1989 9 specimens
Agladrillia (Eumetadrillia) sp. (Woodring 1928)
Jung 1989 1 specimen
Clathrodrillia sp. Dall 1918
Jung 1989 1 specimen
Hindsiclava sp. Hertlein and Strong 1955
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Mangeliidae
Bactrocythara sp. Woodring 1928
Jung 1989 3 specimens
Ithycythara sp. Woodring 1928
Jung 1989 4 specimens
Vitricythara sp. Fargo 1953
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Turridae
Globidrillia sp. Woodring 1928
Jung 1989 8 specimens
Scobinella sp. Conrad 1848
Jung 1989 3 specimens
Lepicythara sp. Olsson 1964
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Terebridae
Strioterebrum sp. Sacco 1891
Jung 1989 13 specimens
Gastropoda - Neogastropoda - Clathurellidae
Glyphostoma (Glyphostoma) sp. Gabb 1872
Jung 1989 2 specimens
Nannodiella sp. (Dall 1919)
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Conidae
Conus sp. Linnaeus 1758
Jung 1989 10 specimens
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria stri n. sp. Landau et al. 2012
Landau et al. 2012 1 specimen
Agatrix epomis (Woodring 1928)
Jung 1989 1 specimen
Gastropoda - Naticidae
Natica (Naticarius) sp. (Dumeril 1806)
Jung 1989 1 specimen
original and current combination Naticarius
Stigmaulax sp. Morch 1852
Jung 1989 5 specimens
Natica (Natica) sp. Scopoli 1777
Jung 1989 1 specimen
Polinices sp. Monfort 1810
Jung 1989 3 specimens
Gastropoda - Rissoidae
Zebina (Zebina) sp. Adams and Adams 1854
Jung 1989 92 specimens
Gastropoda - Trochidae
Microgaza sp. Dall 1881
Jung 1989 7 specimens
Gastropoda - Opisthobranchia - Retusidae
Volvulella (Volvulella) sp. Newton 1891
Jung 1989 3 specimens
Gastropoda - Opisthobranchia - Cylichnidae
Cylichnella sp. Gabb 1873
Jung 1989 1 specimen
Scaphander sp. Monfort 1810
Jung 1989 1 specimen
Acteocina sp. Gray 1847
Jung 1989 3 specimens
Gastropoda - Opisthobranchia - Cavoliniidae
Diacria sp. Gray 1847
Jung 1989 1 specimen
Cavolinia sp. Avilgaard 1791
Jung 1989 1 specimen
Styliola sp. Gray 1850
Jung 1989 5 specimens
Bivalvia - Nuculanida - Nuculanidae
Costelloleda sp. Hertlein and Strong 1940
Jung 1989 4 specimens
Saccella sp. Woodring 1925
Jung 1989 76 specimens
Bivalvia - Nuculida - Nuculidae
Leionucula sp. Quenstedt 1930
Jung 1989 1 specimen
Nucula (Lamellinucula) sp. (Schenck 1944)
Jung 1989 1 specimen
Bivalvia - Pectinida - Pectinidae
Argopecten sp. Monterosato 1889
Jung 1989 1 specimen
Bivalvia - Arcida - Glycymerididae
Tucetona sp. Iredale 1939
Jung 1989 4 specimens
Bivalvia - Arcida - Arcidae
Anadara (Sectiarca) sp. (Olsson 1961)
Jung 1989 7 specimens
Bivalvia - Lucinida - Lucinidae
Myrtea sp. Turton 1822
Jung 1989 2 specimens
Bivalvia - Poromyida - Verticordiidae
Haliris (Haliris) sp. Dall 1886
Jung 1989 1 specimen
Trigonulina sp. d'Orbigny 1946
Jung 1989 1 specimen
Bivalvia - Pholadida - Corbulidae
Caryocorbula sp. (Gardner 1926)
Jung 1989 2 specimens
original and current combination Corbula (Caryocorbula)
Bivalvia - Cardiida - Semelidae
Abra sp. Lamarck 1818
Jung 1989 1 specimen
Actinopteri - Myctophiformes - Myctophidae
Diaphus splendidus (Prochazka 1893)
Schwarzhans and Aguilera 2013
recombined as Scopelus splendidus
Schwarzhans and Aguilera 2013
see common names

Geography
Country:Panama State/province:Bocas
Coordinates: 9.1° North, 81.6° West (view map)
Paleocoordinates:9.8° North, 81.1° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Pliocene
Stage:Piacenzian 10 m.y. bin:Cenozoic 6
Key time interval:Piacenzian
Age range of interval:3.60000 - 2.58000 m.y. ago
Age estimate:2.9 Ma (other)3.6 to 2.1 Ma (other)
Stratigraphy
Geological group:Bocas del Toro Formation:Escudo de Veraguas
Local section:Escudo de Veraguas, Southeastern Coast Local bed:10 m
Local order:bottom to top
Stratigraphic resolution:bed
Stratigraphy comments: STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date extrapolated from Cheetham et al. (1999); age range also provided by Bybell (1999). STRATIGRAPHIC POSITION: From a height of 10 m in section 10 of Collins et al. (1999).
Lithology and environment
Primary lithology:burrows,bioturbation,fine poorly lithified argillaceous sandstone
Secondary lithology:burrows,bioturbation poorly lithified argillaceous siltstone
Includes fossils?Y
Includes fossils?Y
Lithology description: GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.
Environment:deep-water indet.
Geology comments: ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:bulk,sieve,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:BMNH,NMB,USNM
Collectors:Jung et al Collection dates:1996
Collection method comments: COLLECTORS: Peter Jung, A. Heitz, J. Todd et al., 14.9.19959/14/95. REPOSITORY: Naturhistorisches Museum Basel (NMB).
Taxonomic list comments:COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Metadata
Also known as:NMB Loc. 18686
Database number:135876
Authorizer:A. Hendy, M. Uhen Enterer:A. Hendy, M. Uhen
Modifier:A. Hendy Research group:marine invertebrate
Created:2012-11-11 13:33:42 Last modified:2013-11-25 13:37:23
Access level:authorizer only Released:2014-11-11 13:33:42
Creative Commons license:CC BY
Reference information

Primary reference:
17667. L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

42438 P. Jung. 1989. Unpublished notes on Caribbean collection of the Naturhistorisches Museum Basel. [A. Hendy/A. Hendy]
43469 B. D. Landau, R. E. Petit, and C. M. Silva. 2012. The family Cancellariidae (Mollusca: Gastropoda) in the Neogene of the Bocas del Toro region, Panama, with the description of seven new species. Journal of Paleontology 86(2):311-339 [A. Hendy/A. Hendy]
76158 W. Schwarzhans and O. Aguilera. 2013. Otoliths of the Myctophidae from the Neogene of tropical America. Paleo Ichthyologica 13:83-150 [M. Uhen/M. Uhen]