PPP# 2172 - Escudo de Veraguas Island [Escudo de Veraguas Fm]: Piacenzian, Panama
collected by Jung et al 1995

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gastropoda - Naticidae
Sinum sp. Röding 1798
Jung 1989 1 specimen
Polinices sp. Monfort 1810
Jung 1989 19 specimens
Natica (Naticarius) sp. (Dumeril 1806)
Jung 1989 5 specimens
original and current combination Naticarius
Stigmaulax sp. Morch 1852
Jung 1989 1 specimen
Gastropoda - Triviidae
Trivia sp. Broderip 1837
Jung 1989 1 specimen
Gastropoda - Strombidae
Strombus (Strombus) sp. Linnaeus 1758
Jung 1989 6 specimens
Gastropoda - Personidae
Distorsio clathrata (Lamarck 1816)
Jung 1989 2 specimens
Gastropoda - Neogastropoda - Volutidae
Voluta sp. Linnaeaus 1758
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Cystiscidae
Granulina sp. Jousseaume 1888
Jung 1989 3 specimens
Persicula sp. Schumacher 1817
Jung 1989 7 specimens
Gastropoda - Neogastropoda - Muricidae
Siphonochelus (Pilsbrytyphis) sp. (Woodring 1959)
Jung 1989 7 specimens
Haustellum sp. Schumacher 1817
Jung 1989 19 specimens
Gastropoda - Neogastropoda - Marginellidae
Dentimargo sp. Cossmann 1899
Jung 1989 29 specimens
Volvarina sp. (Hinds 1844)
Jung 1989 111 specimens
Prunum sp. Swainson 1840
Jung 1989 6 specimens
Gastropoda - Neogastropoda - Costellariidae
Knefastia sp. Dall 1919
Jung 1989 2 specimens
Gastropoda - Neogastropoda - Ancillariidae
Ancilla (Eburna) sp. (Lamarck 1801)
Jung 1989 9 specimens
original and current combination Eburna
Gastropoda - Neogastropoda - Olividae
Olivella sp. Swainson 1831
Jung 1989 1 specimen
Olivella (Minioliva) sp. Olsson 1956
Jung 1989 6 specimens
Olivella (Macgintiella) sp. Olsson 1956
Jung 1989 102 specimens
Olivella (Niteoliva) sp. Olsson 1956
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Clathurellidae
Glyphostoma (Glyphostoma) sp. Gabb 1872
Jung 1989 2 specimens
Gastropoda - Neogastropoda - Terebridae
Strioterebrum sp. Sacco 1891
Jung 1989 8 specimens
Gastropoda - Neogastropoda - Turridae
Globidrillia sp. Woodring 1928
Jung 1989 2 specimens
Polystira sp. Woodring 1928
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Raphitomidae
Pleurotomella sp. Verrill 1872
Jung 1989 9 specimens
Gastropoda - Neogastropoda - Mangeliidae
Ithycythara sp. Woodring 1928
Jung 1989 2 specimens
Granoturris sp. Fargo 1953
Jung 1989 1 specimen
Saccharoturris sp. Woodring 1928
Jung 1989 1 specimen
Vitricythara sp. Fargo 1953
Jung 1989 8 specimens
Gastropoda - Neogastropoda - Conidae
Conus sp. Linnaeus 1758
Jung 1989 8 specimens
Gastropoda - Neogastropoda - Drilliidae
Syntomodrillia sp. Woodring 1928
Jung 1989 3 specimens
Hindsiclava sp. Hertlein and Strong 1955
Jung 1989 9 specimens
Leptadrillia sp. Woodring 1928
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Columbellidae
Mitrella sp. Risso 1826
Jung 1989 7 specimens
Cotonopsis (Cotonopsis) sp. Olsson 1942
Jung 1989 3 specimens
Cosmioconcha sp. (Dall 1913)
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Nassariidae
Nassarius sp. Duméril 1806
Jung 1989 31 specimens
Gastropoda - Neogastropoda - Fasciolariidae
Fusinus sp. Rafinesque 1815
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Buccinidae
Metula sp. Adams and Adams 1853
Jung 1989 2 specimens
Antillophos (Antillophos) sp. Woodring 1928
Jung 1989 105 specimens
Gastropoda - Neogastropoda - Mitridae
Subcancilla sp. Olsson and Harbison 1953
Jung 1989 1 specimen
Gastropoda - Neogastropoda - Cancellariidae
Aphera cf. lindae Petuch 1987
Landau et al. 2012 4 specimens
Gastropoda - Xenophoridae
Onustus sp. (Swainson 1840)
Jung 1989 1 specimen
Xenophora sp. Fischer von Waldheim 1807
Jung 1989 2 specimens
Gastropoda - Vermetidae
Serpulorbis sp. Sasso 1827
Jung 1989 5 specimens
Gastropoda - Rissoidae
Rissoina (Rissoina) sp. d'Orbigny 1840
Jung 1989 1 specimen
Folinia sp. Crosse 1868
Jung 1989 2 specimens
Zebina (Zebina) sp. Adams and Adams 1854
Jung 1989 44 specimens
Gastropoda - Eulimidae
Niso sp. Risso 1826
Jung 1989 1 specimen
Gastropoda - Pseudomelaniidae
Miraclathurella sp. Woodring 1928
Jung 1989 6 specimens
Lioglyphostoma sp. Woodring 1928
Jung 1989 1 specimen
Gastropoda - Opisthobranchia - Cavoliniidae
Clio sp. Linnaeus 1767
Jung 1989 9 specimens
Creseis sp. Rang 1828
Jung 1989 1 specimen
Cavolinia sp. Avilgaard 1791
Jung 1989 59 specimens
Styliola sp. Gray 1850
Jung 1989 4 specimens
Gastropoda - Opisthobranchia - Bullidae
Bulla sp. Linnaeus 1758
Jung 1989 2 specimens
Gastropoda - Opisthobranchia - Retusidae
Volvulella (Volvulella) sp. Newton 1891
Jung 1989 1 specimen
Gastropoda - Opisthobranchia - Cylichnidae
Scaphander sp. Monfort 1810
Jung 1989 5 specimens
Acteocina sp. Gray 1847
Jung 1989 1 specimen
Cylichnella sp. Gabb 1873
Jung 1989 1 specimen
Gastropoda - Opisthobranchia - Haminoeidae
Atys (Aliculastrum) sp. (Pilsbry 1896)
Jung 1989 1 specimen
original and current combination Aliculastrum
Gastropoda - Heterostropha - Acteonidae
Acteon sp. Montfort 1810
Jung 1989 4 specimens
Gastropoda - Heterostropha - Ringiculidae
Ringicula (Ringiculella) sp. (Sacco 1892)
Jung 1989 16 specimens
Gastropoda - Heterostropha - Architectonicidae
Architectonica sp. Röding 1798
Jung 1989 1 specimen
Gastropoda - Heterostropha - Pyramidellidae
Chemnitzia sp. d'Orbigny 1840
Jung 1989 1 specimen
Pyrgiscus sp. Philippi 1841
Jung 1989 2 specimens
spelled with current rank as Turbonilla (Pyrgiscus)
Callolongchaeus sp. Dall 1903
Jung 1989 3 specimens
Gastropoda - Trochidae
Calliostoma sp. Swainson 1840
Jung 1989 1 specimen
Gastropoda - Lepetellida - Fissurellidae
Diodora sp. Gray 1821
Jung 1989 1 specimen
Bivalvia - Nuculanida - Nuculanidae
Costelloleda sp. Hertlein and Strong 1940
Jung 1989 1 specimen
Saccella sp. Woodring 1925
Jung 1989 6 specimens
Bivalvia - Nuculida - Nuculidae
Leionucula sp. Quenstedt 1930
Jung 1989 6 specimens
Nucula (Lamellinucula) sp. (Schenck 1944)
Jung 1989 2 specimens
Bivalvia - Carditida - Carditidae
Cardites sp. Link 1807
Jung 1989 68 specimens
Bivalvia - Cardiida - Veneridae
Gouldia sp. Adams 1847
Jung 1989 45 specimens
Ventricolaria sp. Keen 1954
Jung 1989 1 specimen
synonym of Globivenus
Bivalvia - Cardiida - Chamidae
Chama sp. Linnaeus 1758
Jung 1989 12 specimens
Bivalvia - Cardiida - Tellinidae
Eurytellina sp. Fischer 1887
Jung 1989 2 specimens
spelled with current rank as Tellina (Eurytellina)
Bivalvia - Cardiida - Semelidae
Abra sp. Lamarck 1818
Jung 1989 54 specimens
Bivalvia - Cardiida - Cardiidae
Laevicardium sp. Swainson 1840
Jung 1989 6 specimens
Bivalvia - Pholadida - Corbulidae
Varicorbula sp. Grant and Gale 1931
Jung 1989 3 specimens
spelled with current rank as Corbula (Varicorbula)
Bothrocorbula (Hexacorbula) sp. (Olsson 1932)
Jung 1989 5 specimens
spelled with current rank as Hexacorbula
Caryocorbula sp. (Gardner 1926)
Jung 1989 82 specimens
original and current combination Corbula (Caryocorbula)
Bivalvia - Pectinida - Pectinidae
Argopecten sp. Monterosato 1889
Jung 1989 32 specimens
Spathochlamys sp. Waller 1993
Jung 1989 16 specimens
Leopecten coraliphilla
Jung 1989 3 specimens
Bivalvia - Pectinida - Propeamussiidae
Cyclopecten sp. Verrill 1897
Jung 1989 1 specimen
Bivalvia - Arcida - Arcidae
Barbatia (Barbatia) sp. Gray 1842
Jung 1989 1 specimen
Arca (Arca) sp. Linnaeus 1758
Jung 1989 1 specimen
Anadara (Sectiarca) sp. (Olsson 1961)
Jung 1989 66 specimens
Scaphopoda - Dentaliida - Dentaliidae
Dentalium sp. Linnaeus 1758
Jung 1989 43 specimens
Scaphopoda - Gadilida - Gadilidae
Gadilopsis sp. (Woodring 1925)
Jung 1989 131 specimens
see common names

Geography
Country:Panama State/province:Bocas
Coordinates: 9.1° North, 81.6° West (view map)
Paleocoordinates:9.0° North, 81.1° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Pliocene
Stage:Piacenzian 10 m.y. bin:Cenozoic 6
Key time interval:Piacenzian
Age range of interval:3.60000 - 2.58000 m.y. ago
Age estimate:2.9 Ma (other)3.6 to 2.1 Ma (other)
Stratigraphy
Geological group:Bocas del Toro Formation:Escudo de Veraguas
Local section:Escudo de Veraguas, Northern Coast Local bed:78.5 m
Local order:bottom to top
Stratigraphic resolution:bed
Stratigraphy comments: STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date extrapolated from Cheetham et al. (1999); age range also provided by Bybell (1999). STRATIGRAPHIC POSITION: From a height of 78.5 m in section 10 of Collins et al. (1999).
Lithology and environment
Primary lithology:burrows,bioturbation,fine poorly lithified argillaceous sandstone
Secondary lithology:burrows,bioturbation poorly lithified argillaceous siltstone
Includes fossils?Y
Includes fossils?Y
Lithology description: GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.
Environment:deep-water indet.
Geology comments: ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:bulk,sieve,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:BMNH,NMB,USNM
Collectors:Jung et al Collection dates:1995
Collection method comments: COLLECTORS:coll. P. Jung, A. Heitz, J. Todd et al., 2/22/95. REPOSITORY: Naturhistorisches Museum Basel (NMB).
Taxonomic list comments:COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Metadata
Also known as:NMB Loc. 18671; 95-05-11; mislabelled as PPP# 2127 in Landau et al (2012)
Database number:135885
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2012-11-12 04:03:18 Last modified:2013-11-25 13:37:23
Access level:authorizer only Released:2014-11-12 04:03:18
Creative Commons license:CC BY
Reference information

Primary reference:
17667. L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

42438 P. Jung. 1989. Unpublished notes on Caribbean collection of the Naturhistorisches Museum Basel. [A. Hendy/A. Hendy]
43469 B. D. Landau, R. E. Petit, and C. M. Silva. 2012. The family Cancellariidae (Mollusca: Gastropoda) in the Neogene of the Bocas del Toro region, Panama, with the description of seven new species. Journal of Paleontology 86(2):311-339 [A. Hendy/A. Hendy]