PBDB Collection

Eckfeld Maar (NHMN 2000 collection): MP 13, Germany
collected 2000

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Insecta - Diptera - Limoniidae

	Dicranoptycha cf. megaphallus Alexander 1926	Wappler 2002	1 specimen
	NHMM PE 2000/843 (2 measurements)
Insecta - Odonata

	Lestinoidea indet. Calvert 1901	Wappler 2003	1 specimen
	NHMM PE2000/545a,b LS
Insecta - Odonata - Priscalestidae

	Priscalestes germanica n. gen., n. sp. Wappler and Petrulevičius 2007	Wappler and Petrulevičius 2007	1 specimen
	PE 2000/45a+b (1 measurement)
Insecta - Odonata - Megapodagrionidae

	Euarchistigma superstes n. sp. Wappler 2003	Wappler 2003	1 specimen
	recombined as Eckfeldia superstes
	NHMM PE 2000/44a+b (1 measurement)
Insecta - Hemiptera - Fulgoridae

	Lycorma sp. Koch 1882	Wappler 2003	1 specimen
	NHMM PE2000/831 a+b LS
Insecta - Hemiptera - Ricaniidae

	Ricaniidae informal indet. 1 (Amyot and Audinet-Serville 1843)	Wappler 2003	2 specimens
	NHMM PE2000/832, 833 a+b LS

	Ricaniidae informal indet. 2 (Amyot and Audinet-Serville 1843)	Wappler 2003	2 specimens
	NHMM PE2000/829 a+b, PE2000/830 LS
Insecta - Hemiptera - Lygaeidae

	Lygaeidae informal indet. 1 Schilling 1829	Wappler 2003	1 specimen
	NHMM PE2000/1047 LS

	Lygaeidae informal indet. 2 Schilling 1829	Wappler 2003	1 specimen
	NHMM PE2000/1052 LS
Insecta - Hemiptera - Aradidae

	Mezira crassifemur n. sp. Wappler and Heiss 2006	Wappler and Heiss 2006	2 specimens
	NHMM PE 2000/1048, 1050 (1 measurement)

	Aradidae indet. (Brullé 1836)	Wappler 2003	2 specimens
	NHMM PE2000/1049, 1064
	= Mezira eckfeldensis Wappler and Heiss 2006	Wappler and Heiss 2006
Insecta - Hemiptera - Acanthosomatidae

	Acanthosomatidae indet. (Signoret 1863)	Wappler 2003	1 specimen
	NHMM PE2000/1051 a+b
Insecta - Hemiptera - Pentatomidae

	Pentatomidae informal indet. 1 Leach 1815	Wappler 2003	1 specimen
	NHMM PE2000/1046 a+b
Insecta - Hemiptera - Gerridae

	Lutetiabates eckfeldensis Wappler and Andersen 2004	Wappler 2003	4 specimens
	NHMM PE2000/1, 542, 543, 823

	Gerris sp. Fabricius 1794	Wappler and Andersen 2004	1 specimen
	NHMM PE2000/2-LS
Insecta - Hemiptera

	Cercopoidea informal indet. 1 Leach 1815	Wappler 2003	3 specimens
	NHMM PE2000/824, 825, 1983

	Cercopoidea informal indet. 2 Leach 1815	Wappler 2003	5 specimens
	NHMM PE2000/826, 827, 835, 1039, 1983

	Cercopoidea informal indet. 3 Leach 1815	Wappler 2003	1 specimen
	NHMM PE2000/828 a+b

	Membracoidea indet. Rafinesque 1815	Wappler 2003	1 specimen
	NHMM PE2000/836
Insecta - Blattaria

	Blattaria indet. (Latreille 1810)	Wappler 2003	2 specimens
	NHMM PE2000/1067 a+b LS, PE2000/1066 a+b LS
Insecta - Dermaptera

	Neodermaptera informal indet. 2 Engel 2003	Wappler et al. 2005	1 specimen
	NHMM PE 2000/1062-LS
Insecta - Dermaptera - Forficulidae

	Forficulina indet.	Wappler 2003	2 specimens
	NHMM PE2000/1061 LS, PE2000/1063 a+b LS. (2 measurements)
	= Forficula kotejai n. sp. Wappler et al. 2005	Wappler et al. 2005
Insecta - Blattodea - Blattellidae

	Blattellidae informal indet. 1 Karny 1908	Wappler 2003	1 specimen
	NHMM PE 2000/1070

	Blattellidae informal indet. 2 Karny 1908	Wappler 2003	2 specimens
	NHMM PE 2000/1069, PE2000/1068
Insecta - Blattodea - Blattidae

	Blattidae indet. Latreille 1810	Wappler 2003	1 specimen
	NHMM PE2000/1065 ab
Insecta - Hymenoptera - Formicidae

	Oecophylla eckfeldiana n. sp. Dlussky et al. 2008	Dlussky et al. 2008	7 specimens
	NHMM PE-2000/8, 1948, 1947, 1951, 9, 11, 12 (1 measurement)

	Gesomyrmex flavescens n. sp. Dlussky et al. 2009		1 specimen
	PE2000/14 (1 measurement)
Insecta - Hymenoptera - Apidae

	Electrapis informal sp.1 Cockerell 1908	Wappler and Engel 2003	1 specimen
	NHMM PE2000/853a,b LS (2 measurements)

	Electrapis informal sp.2 Cockerell 1908	Wappler and Engel 2003	1 specimen
	NHMM PE2000/846a,b LS (1 measurement)

	Electrapis informal sp.3 Cockerell 1908	Wappler and Engel 2003	1 specimen
	NHMM PE2000/849a,b LS (1 measurement)

	Electrapis micheneri Wappler and Engel 2003	Wappler and Engel 2003	1 specimen
	recombined as Electrapis (Mesomelissa) micheneri
	NHMM PE2000/852a,b LS (2 measurements)

	Electrapis electrapoides (Lutz 1993)	Wappler and Engel 2003	1 specimen
	recombined as Electrapis (Eckfeldapis) electrapoides
	PE2000/850 LS (2 measurements)

	Electrapis prolata Wappler and Engel 2003	Wappler and Engel 2003	2 specimens
	recombined as Electrapis (Eckfeldapis) prolata
	NHMM PE2000/847 LS, PE2000/848a,b LS (2 measurements)

	Protobombus informal sp. 1 Cockerell 1908	Wappler and Engel 2003	1 specimen
	NHMM PE2000/851a,b LS (1 measurement)

	Protobombus pristinus Wappler and Engel 2003	Wappler and Engel 2003	1 specimen
	recombined as Protobombus (Thnetobombus) pristinus
	NHMM PE2000/863a,b LS (1 measurement)
Insecta - Hymenoptera - Crabronidae

	cf. Crabronidae indet. (Latreille 1802)	Wappler 2003	1 specimen
	original and current combination Crabronites
	NHMM PE2000/865
Insecta - Coleoptera - Carabidae

	Carabidae informal indet. 1 (Latreille 1802)	Wappler 2003	1 specimen
	NHMM PE2000?1076 a+b

	Carabidae informal indet. 3 (Latreille 1802)	Wappler 2003	1 specimen
	NHMM PE2000/1078 a+b

	Pleuarthropterus (Blaticarthropterus) nonsucineus n. sp. Wappler 2003	Wappler 2003	1 specimen
	NHMM PE2000/544; stratigraphischer Bereich ca. 24 cm unter Leithorizont S8.
Insecta - Coleoptera - Dytiscidae

	? Dytiscidae indet. (Leach 1815)	Wappler 2003	1 specimen
	NHMM PE2000/990
Insecta - Coleoptera - Lucanidae

	Lucanidae indet. (Latreille 1804)	Wappler 2003	2 specimens
	NHMM PE2000/1176, 1175
Insecta - Coleoptera - Scarabaeidae

	Aphodiinae informal indet. 1 (Leach 1815)	Wappler 2003	55 specimens
	NHMM PE2000/578

	Aphodiinae informal indet. 2 (Leach 1815)	Wappler 2003	39 specimens
	NHMM PE2000/605

	Aphodiinae informal indet. 3 (Leach 1815)	Wappler 2003	2 specimens
	NHMM PE2000/593, 592

	Rutelinae indet. (MacLeay 1819)	Wappler 2003	2 specimens
	NHMM PE2000/591, 590
Insecta - Coleoptera - Staphylinidae

	Pselaphinae indet. (Latreille 1802)	Wappler 2003	4 specimens
	Pselaphii
	NHMM PE2000/987, 989, 988, 1984
Insecta - Coleoptera - Dermestidae

	? Attagenus sp. Latreille 1802	Wappler 2003	3 specimens
	NHMM PE2000/995, 996, 997 (2 measurements)
	= Eckfeldattagenus eocenicus n. gen., n. sp. Háva and Wappler 2014	Háva and Wappler 2014
Insecta - Coleoptera - Lophocateridae

	Cassidinae informal indet. 1 Gyllenhal 1813	Wappler 2003	1 specimen
	NHMM 2000/475 (2 measurements)
	= Ancyrona eocenica n. sp. Schmied et al. 2009	Schmied et al. 2009
Insecta - Coleoptera - Cleridae

	Cleridae indet. (Latreille 1802)	Wappler 2003	2 specimens
	original and current combination Clerii
	NHMM PE2000/991, 998
Insecta - Coleoptera - Tenebrionidae

	Tenebrionidae indet. (Latreille 1802)	Wappler 2003	91 specimens
	original and current combination Tenebrionites
	NHMM PE2000/1179, 713
Insecta - Coleoptera - Zopheridae

	Colydiidae indet. (Billberg 1820)	Wappler 2003	3 specimens
	replaced by Zopheridae
	NHMM PE2000/992, 993, 994
Insecta - Coleoptera - Curculionidae

	Curculionini indet. Latreille 1802	Wappler 2003	2 specimens
	NHMM PE2000/1493, 1492
Insecta - Coleoptera - Brentidae

	Brentinae informal indet. 2 Billberg 1820	Wappler 2003	1 specimen
	NHMM PE2000/1248

	? Trachelizini informal indet. 1 Lacordaire 1865	Wappler 2003	3 specimens
	NHMM PE2000/1279, 1247, 1246

	Eckfelderolispa manderschieta n. sp. Legalov and Wappler 2019	Legalov and Wappler 2019	1 specimen
	NHMM PE 2000/2166 (1 measurement)
Insecta - Coleoptera - Chrysomelidae

	Lamprosomatinae indet. Lacordaire 1848	Wappler 2003	1 specimen
	NHMM PE2000/474

	Donaciinae indet. (Kirby 1837)	Wappler 2003	1 specimen
	NHMM PE2000/477

	Chrysomelinae indet. Latreille 1802	Wappler 2003	1 specimen
	NHMM 2000

	Cassidinae informal indet. 2 Gyllenhal 1813	Wappler 2003	1 specimen
	NHMM 2000/476
Insecta - Coleoptera - Cerambycidae

	Cerambycidae informal indet. 1 (Latreille 1802)	Wappler 2003	1 specimen
	Cerambicini
	NHMM PE2000/1186
Insecta - Coleoptera - Buprestidae

	Buprestinae informal indet. 1 Leach 1815	Wappler 2003	1 specimen
	NHMM PE2000/842

	Buprestinae informal indet. 2 Leach 1815	Wappler 2003	2 specimens
	NHMM PE2000/708, 698

	Buprestinae informal indet. 3 Leach 1815	Wappler 2003	1 specimen
	NHMM PE2000/692 a+b

	Buprestinae informal indet. 4 Leach 1815	Wappler 2003	1 specimen
	NHMM PE2000/665

	Buprestis sp. Linnaeus 1758	Wappler 2003	5 specimens
	NHMM PE2000/1224, 1225, 691, 690, 689

	Julodinae indet. Lacordaire 1857	Wappler 2003	2 specimens
	NHMM PE2000/839, 671

	? Agrilini informal indet. 1 Laporte 1835	Wappler 2003	2 specimens
	NHMM PE2000/668, 667

	? Agrilini informal indet. 2 Laporte 1835	Wappler 2003	1 specimen
	NHMM PE2000/666

	Coroebini informal indet. 2 Bedel 1921	Wappler 2003	2 specimens
	corrected as Coraebini
	NHMM PE2000/662, 661

	Coroebini informal indet. 3 Bedel 1921	Wappler 2003	1 specimen
	corrected as Coraebini
	NHMM PE2000/663

	Coroebini informal indet. 5 Bedel 1921	Wappler 2003	1 specimen
	corrected as Coraebini
	NHMM PE2000/664
Insecta - Coleoptera - Throscidae

	Throscidae indet. Laporte 1840	Wappler 2003	1 specimen
	NHMM PE2000/353
Insecta - Coleoptera - Elateridae

	? Athous sp. Eschscholtz 1829	Wappler 2003	2 specimens
	NHMM PE2000/2213, 347

	Lanelater nicoleae n. sp. Wappler 2003	Wappler 2003	1 specimen
	NHMM PE2000/349 stratigraphischer Bereich 64 cm unter Leithorizont S8.

	? Lacon sp. Laporte 1838	Wappler 2003	1 specimen
	NHMM PE2000/345

	Macropunctum densipunctum n. sp. Wappler 2003	Wappler 2003	2 specimens
	NHMM PE2000/955, 744 stratigraphischer Bereich ca. 30 cm unter Leithorizont KaLH.
Insecta - Coleoptera - Cupedidae

	Cupedidae informal indet. 1 Laporte 1836	Wappler 2003	1 specimen
	NHMM PE2000/1145 a+b

	Cupedidae informal indet. 2 Laporte 1836	Wappler 2003	1 specimen
	NHMM PE2000/1146 a+b

	? Tenomerga eckfeldense Tröster 1993	Wappler 2003	8 specimens
	recombined as Cupes eckfeldensis
	NHMM PE2000/1173, 1138, 1139, 1140, 1141, 1142, 1143, 1144
see common names

Geography

Country:	Germany	State/province:	Eifel	County:	Eckfeld
Coordinates:	50.1° North, 6.8° East (view map)
Paleocoordinates:	47.0° North, 2.9° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	small collection

Time

Period:	Paleogene	Epoch:	Eocene
Stage:	Lutetian	10 m.y. bin:	Cenozoic 2
Key time interval:	MP 13
Age range of interval:	47.80000 - 41.20000 m.y. ago

Stratigraphy

Formation:	Eifel
Stratigraphic resolution:	bed
Stratigraphy comments: "MP 13... recently dated at 44.3 +/- 0.4 million years BP (Mertz et al. 2000)" (Franzen 2004). Insects come mostly from horizons near marker horizon HT & KaLH.

Lithology and environment

Primary lithology:	planar lamination siltstone
Secondary lithology:	planar lamination "shale"
Includes fossils?	Y
Includes fossils?	Y
Lithology description: ...laminitic "oilshales" and siltstones... with frequent turbiditic intercalations... plant macrofossils are ... distributed throughout the sequence under investigation. ... coarse and robust plant debris such as wood fragments and large fruits/seeds is concentrated in the turbiditic layers. (Wilde and Frankenhauser 1998).
Environment:	crater lake	Tectonic setting:	volcanic basin
Geology comments: "a small freshwater [crater] lake a few hundred meters in diameter" (Franzen 2004)

Taphonomy

Modes of preservation:	mold/impression,adpression,original carbon
Size of fossils:	macrofossils,mesofossils

Collection methods and comments

Collection methods:	selective quarrying,surface (in situ),field collection,survey of museum collection
Reason for describing collection:	taxonomic analysis
Collection dates:	2000
Collection method comments: Repository: Naturhistorisches Museum Mainz

Metadata

Database number:	138912
Authorizer:	M. Clapham	Enterer:	M. Clapham, J. Karr, J. Sandlin
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	2013-02-01 15:25:51	Last modified:	2017-09-19 10:50:06
Access level:	the public	Released:	2013-02-01 15:25:51
Creative Commons license:	CC BY

Reference information

Primary reference:

44599.

G. M. Dlussky, T. Wappler, and S. Wedmann. 2009. Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities. Zootaxa 2031:1-20 [M. Clapham/M. Clapham/M. Clapham]

Secondary references:

37721		G. M. Dlussky, T. Wappler, and S. Wedmann. 2008. New midle Eocene formicid species from Germany and the evolution of weaver ants. Acta Palaeontologica Polonica 53(4):615-626 [M. Clapham/M. Clapham]
49552		J. Háva and T. Wappler. 2014. A new genus and species of Dermestidae (Coleoptera) from the Eckfeld Maar crater (Middle Eocene, Germany). Bulletin of Geosciences 89:67-74 [M. Clapham/M. Clapham]
77928		A. A. Legalov and T. Wappler. 2019. The oldest record of straight-snouted weevils (Coleoptera: Curculionoidea: Brentidae: Brentinae) from the Eocene of Germany. Historical Biology 33:1464-1472 [M. Clapham/M. Clapham]
46916		H. Schmied, T. Wappler, and J. Kolibáč. 2009. A new bark-gnawing beetle (Coleoptera, Trogossitidae) from the middle Eocene of Europe, with a checklist of fossil Trogossitidae. Zootaxa 1993:17-26 [M. Clapham/J. Karr/M. Clapham]
52179		T. Wappler. 2002. Dicranoptycha (Insecta: Nematocera: Limoniidae) aus der mitteleozänen Fossillagerstätte Eckfelder Maar bei Manderscheid/Eifel (Deutschland). Mainzer Naturwissenschaftliches Archiv 40:129-134 [M. Clapham/M. Clapham]
43376		T. Wappler. 2003. Systematik, Phylogenie, Taphonomie und Paläoökologie der Insekten aus dem Mittel-Eozän des Eckfelder Maares, Vulkaneifel. Clausthaler Geowissenschaften 2:1-241 [M. Clapham/J. Karr]
44718		T. Wappler and N. M. Andersen. 2004. Fossil water striders from the Middle Eocene fossil sites of Eckfeld and Messel, Germany (Hemiptera, Gerromorpha). Paläontologische Zeitschrift 78:41-52 [M. Clapham/M. Clapham/M. Clapham]
50146		T. Wappler, M. S. Engel, and F. Haas. 2005. The earwigs (Dermaptera: Forficulidae) from the middle Eocene Eckfeld maar, Germany. Polskie Pismo Entomologiczne 74:227-250 [M. Clapham/M. Clapham/M. Clapham]
51777		T. Wappler and E. Heiss. 2006. Flatbugs from Paleogene limnic sediments. II. Eckfeld maar (Heteroptera: Aradidae). Mainzer Naturwissenschaftliches Archiv 44:53-60 [M. Clapham/M. Clapham/M. Clapham]
57309		T. Wappler, C. C. Labandeira, M. S. Engel, R. Zetter, and F. Grímsson. 2015. Specialized and generalized pollen-collection strategies in an ancient bee lineage. Current Biology 25:3092-3098 [M. Clapham/M. Clapham]