Gran Barranca - southern cliff of lake Colhué Huapí: Colhuehuapian, Argentina
collected by Tournouër and others

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Rodentia - Chinchillidae
Eoviscaccia australis Vucetich 1989
Vucetich et al. 2010
Mammalia - Rodentia
Chinchilloidea indet. Bennett 1833
Vucetich et al. 2010
Mammalia - Rodentia - Neoepiblemidae
Neoepiblemidae indet. Kraglievich 1926
Vera et al. 2015 1 specimen
MGP-PD 29099, mandible p4-m2
Perimys incavatus Ameghino 1902
Vucetich et al. 2010
Perimys transversus Ameghino 1902
Vucetich et al. 2010
Perimys dissimilis Ameghino 1902
Vucetich et al. 2010
Mammalia - Rodentia
Saremmys ligcura Busker et al. 2019
Busker et al. 2019
Luantus initialis (Ameghino 1902)
Vucetich et al. 2010
original and current combination Luanthus initialis
Soriamys sp. Kramarz 2001
Vucetich et al. 2010
? Eosteiromys sp. Ameghino 1902
Vucetich et al. 2010
Eosteiromys segregatus Ameghino 1902
Vucetich et al. 2010
Eosteiromys homogenidens Ameghino 1902
Vucetich et al. 2010
Mammalia - Rodentia - Erethizontidae
Parasteiromys friantae n. sp. Candela 1999
Candela 1999 1 specimen
MPEF 5501, isolated upper right molariform
Parasteiromys uniformis n. sp. Ameghino 1904
Candela 1999 1 specimen
MACN A 52-178, maxillary with DP4-M1, holotype
Hypsosteiromys sp. Patterson 1958
Dozo et al. 2004 1 specimen
MPEF-PV 6029, skull
Steiromys axiculus n. sp. Ameghino 1902
Patterson 1958 1 specimen
recombined as Hypsosteiromys axiculus
MACN A 52-171, dentary m1-3, holotype
Steiromys nectus n. sp. Ameghino 1902
Candela and Vucetich 2002 4 specimens
recombined as Hypsosteiromys nectus
MACN A 52-177, dentary dp4-m2, holotype; MACN A 52-175A, p4; MNHN col. 50, p4; MLP 85-VII-3-4, m1 or m2
Steiromys tabulatus n. sp. Ameghino 1902
Patterson 1958 1 specimen
synonym of Hypsosteiromys nectus
MACN A 52-172, dentary p4-m3, lectotype
Branisamyopsis australis Candela 2003
Vucetich et al. 2010
Branisamyopsis australis n. gen., n. sp. Candela 2003
Candela 2003
Holotype: MMP 937
Mammalia - Rodentia - Cephalomyidae
? Cephalomyidae indet. Ameghino 1897
Vucetich et al. 2010
Mammalia - Rodentia
Octodontoidea informal Gen. nov. sp. large
Vucetich et al. 2010
Mammalia - Rodentia - Acaremyidae
Acaremys cf. murinus Ameghino 1887
(6 measurements)
Galileomys antelucanus n. gen., n. sp. Vucetich and Kramarz 2003
(5 measurements)
Mammalia - Rodentia
Caviocricetus lucasi Vucetich and Verzi 1996
Vucetich et al. 2010
Acarechimys pulchellus n. sp. (Ameghino 1902)
Vucetich et al. 2010
synonym of Acarechimys gracilis
Holotype. MACN A 52-128
Acarechimys cf. minutissimus Ameghino 1887
Vucetich et al. 2010
Prospaniomys informal sp. 1 Ameghino 1902
Vucetich et al. 2010
Prospaniomys priscus Ameghino 1902
Vucetich et al. 2010
Protadelphomys latus Ameghino 1902
Vucetich et al. 2010
Mammalia - Rodentia - Echimyidae
Paradelphomys fissus n. gen., n. sp. Patterson and Pascual 1968
Patterson and Pascual 1968 1 specimen
MLP 125, dentary dp4-m1, type
Mammalia - Rodentia
? Protacaremys sp. Ameghino 1902
Vucetich et al. 2010
Protacaremys avunculus Ameghino 1902
Vucetich et al. 2010
Protacaremys prior Ameghino 1902
Vucetich et al. 2010
Mammalia - Primates - Homunculidae
Mazzonicebus almendrae n. gen., n. sp. Kay 2010
Kay 2010 33 specimens
Teeth, maxillaries, and mandibles in MLP and MPEF-PV. Several stratigraphic levels.
Mazzonicebus almendrae Kay 2010
Novo et al. 2018
MNHN.F.COL93a and b. The specimen label reads (Fig. 3G): “Primate, col 93 // Colhué Huapi B // M.A. Tournouër, 1899-7”. Both specimens were found in the box with the same label
Homunculus sp. Ameghino 1891
Hershkovitz 1981
No. 69-111-12-1, lower jaw
Mammalia - Chiroptera - Molossidae
Mormopterus indet. Peters 1865
Czaplewski 2010 1 specimen
MLP 93-XI-18-19, right dentary fragment with m2
Mormopterus barrancae n. sp. Czaplewski 2010
Czaplewski 2010 2 specimens
MLP 93-XI-18-15, mandible with p2, p4, m1, m2, holotype, from "nivel 'C' (gris)"; MLP 93-XI-18-18, left m1
Mammalia - Chiroptera - Phyllostomidae
Phyllostominae indet. Gray 1825
Czaplewski 2010 1 specimen
MLP 93-XI-18-16, right m3
Mammalia - Panameriungulata - Adianthidae
Adianthus patagonicus n. sp. Ameghino 1904
Cifelli and Soria 1983 1 specimen
recombined as Proheptaconus patagonicus
MACN A52-218, M3, holotype
Mammalia - Panameriungulata - Proterotheriidae
Prolicaphrium spectabile Ameghino 1902
Vera et al. 2015 1 specimen
MGP-PD 29096, upper molar
Prolicaphrium festinum Ameghino 1902
Vera et al. 2015 2 specimens
recombined as Licaphrops festinus
MGP-PD 29024, mandible with two molars; MGP-PD 26591: mandible with left m1-3
Mammalia - Panameriungulata - Macraucheniidae
Pternoconius tournoueri n. sp. Soria and Hoffstetter 1985
Soria and Hoffstetter 1985
Holotype: MNHN COL 121
Cramauchenia indet. Ameghino 1902
Vera et al. 2015 3 specimens
MGP-PD 29040 A, M3; MGP-PD 29060 A, lower molar, and B, astragalus
Cramauchenia normalis n. gen., n. sp. Ameghino 1902
Dozo and Vera 2010
MACN A-52-119 - holotype
Mammalia - Cingulata - Glyptodontidae
Glyptodontidae indet. Burmeister 1879
Vera et al. 2015 1 specimen
MGP-PD 31693, proximal ulna
Mammalia - Cingulata - Propalaehoplophoridae
Propalaehoplophorinae indet. (Ameghino 1891)
Carlini et al. 2005
original and current combination Propalaehoplophoridae
Mammalia - Cingulata - Peltephilidae
Peltephilidae indet. Ameghino 1894
Carlini et al. 2005
Mammalia - Cingulata - Dasypodidae
Dasypodidae indet. (Gray 1821)
Vera et al. 2015 2 specimens
MGP-PD 29101, bony external plates; MGP-PD 29076 A, astragalus
Pseudostegotherium glangeaudi Ameghino 1902
Carlini et al. 2010 16 specimens
MACN A-12679, osteroderms of the carapace and left mandibular fragment with two teeth; osteoderms in MPEF PV
Mammalia - Cingulata - Chlamyphoridae
Prozaedyus humilis Ameghino 1902
Carlini et al. 2010
Prozaedyus cf. humilis Ameghino 1902
Carlini et al. 2005
Proeutatus sp. Ameghino 1891
Carlini et al. 2010
Proeutatus cf. robustus Scott 1903
Carlini et al. 2005
Proeutatus oenophorus (Ameghino 1887)
Carlini et al. 2005
recombined as Proeutatus oenophorum
Mammalia - Megalonychidae
Megalonychidae indet. Gervais 1855
Vera et al. 2015 1 specimen
MGP-PD 31692, Mc? II or III
Proschismotherium sp. Ameghino 1902
Carlini et al. 2005
Hapaloides sp. Ameghino 1902
Carlini et al. 2005
Holomegalonyx sp. Scillato-Yané 1979
Carlini et al. 2005
Mammalia
Stegotherium pascuali n. sp. Fernicola and Vizcaino 2008
Fernicola and Vizcaino 2008
Holotipo. MACN A-12680d. Found in level 5
Stegotherium caroloameghinoi n. sp. Fernicola and Vizcaino 2008
Fernicola and Vizcaino 2008
Holotipo. MACN-A 10443a. Found in level 5
Stegotherium variegatum Ameghino 1902
Carlini et al. 2010
Stegotherium variegatum Ameghino 1902
Carlini et al. 2005
Mammalia - Notoungulata
Argyrohippus fraterculus Ameghino 1902
Hernández Del Pino et al. 2019
MACN A 52-17 (lectotype), right mandible and symphysis with left i2–i3–c and right i1–i3–c, alveoli of p1–p2 and p3–m3; MACN A 52-18, right maxillary fragment with P4–M3 (left) and left maxillary fragment (reversed; right) with alveoli of P1 and P2–P4
Argyrohippus cf. boulei Ameghino 1902
López et al. 2010 3 specimens
synonym of Argyrohippus fraterculus
MPEF-PV 5388, mandible with left and right i1-3, c, p2-m3; MPEF-PV 5922, fragment of upper molar and left m1; MPEF-PV 7325, left lower molar
Argyrohippus boulei Ameghino 1902
Hernández Del Pino et al. 2019
synonym of Argyrohippus fraterculus
MACN A 52-12 (lectotype)
Mammalia - Notoungulata - Leontiniidae
Leontiniidae indet. Ameghino 1894
Vera et al. 2015 3 specimens
"MGP-PD 29107: A, fragment of mandible with left m3; B, left lower molar; C, left lower molar"
Colpodon sp. Burmeister 1885
Vera et al. 2015 1 specimen
"MGP-PD 29076B: mandible with left m1-2"
Colpodon distinctus n. sp. Ameghino 1902
Ribeiro et al. 2010 21 specimens
Maxilla, dentary, skull and tooth fragments in MACN, MPEF-PV, AMNH, and FMNH. From localities GBV-8, GBV-9, GBV-10, GBV-38, and GBV-43.
Mammalia - Notoungulata
Pachyrukhos politus Ameghino 1901
Seoane and Cerdeño 2019
MLP 82-V-2-107, right mandibular fragment with p2–m3; and MLP 85VII-3-28, right mandibular fragment with p2–p3
Mammalia - Notoungulata - Interatheriidae
Cochilius volvens Ameghino 1902
Simpson 1932
AMNH 29651, 29686, 29652, 29653, 29658, 29654, 29655, 29657
Mammalia - Astrapotheria - Astrapotheriidae
Astrapotheriidae informal gen. et sp. nov.
Kramarz and Bond 2010 1 specimen
MACN A 52-522b, isolated canine, syntype of _Parastrapotherium crassum_. Species appears to also occur at Cerro Bandera and may be uruguaytheriine.
Astrapotheriidae indet. Ameghino 1887
Vera et al. 2015 9 specimens
"MGP-PD 29049 A, mandible with left m3; B, right P2; C, right P1; D, right upper molar, E, canine?; F, lingual fragment of left upper molar; G, fragments of tooth; MGP-PD 29075: two teeth; MGP-PD 29108: teeth"
? Astrapotherium ruderarium n. sp. (Ameghino 1902)
Kramarz and Bond 2010
Parastrapotherium symmetrum n. sp. (Ameghino 1902)
Kramarz and Bond 2010 2 specimens
MACN A 52-507a, holotype, isolated lower incisor, from "_Colpodon_ beds at Gran Barranca". MPEF PV 7923, isolated lower incisor, from Gran Barranca Colhuehuapian West locality, level B
Mammalia
Marsupialia indet. (Illiger 1811)
Goin et al. 2007 2 specimens
"Didelphimorphia o Sparassodonta gen. et sp. indet." MLP 77-VI-13-8, trigonid; MLP 92-X-10-11, M4
Mammalia - Didelphimorphia
Didelphimorphia indet. Gill 1872
Goin et al. 2007 1 specimen
MLP 77-VI-13-25, left dentary fragment with c, p2, half of p3, fragments of m2-3, and edentulous right dentary
Didelphoidea indet. Gray 1821
Goin and Abello 2013 1 specimen
MLP 77-VI-13-26, left maxillary fragment with part of the plate, roots of P1 and P3, and complete P2 and M1-4
Mammalia - Didelphimorphia - Didelphidae
Didelphidae indet. Gray 1821
Goin et al. 2007 4 specimens
MPEF-PV 4774, m2 or m3; MPEF-PV 4709, m?1; MPEF-PV 4707, Mx; MPEF-PV 4773, m?1 fragment
Mammalia - Didelphimorphia - Caluromyidae
? Caluromyidae indet. Kirsch and Reig 1977
Goin et al. 2007 1 specimen
spelled with current rank as Caluromyinae
MPEF-PV 4708, mx
Mammalia - Paucituberculata - Palaeothentidae
Parabderites bicrispatus Ameghino 1902
Marshall et al. 1984 1 specimen
MNHN COL 124, dentary m2
Mammalia - Paucituberculata - Caenolestidae
Stilotherium parvum n. sp. Abello et al. 2021
Abello et al. 2021
Type: MPEF-PV 5830 - was in Abello, 2013; Rincón et al., 2015; Engelman et al., 2016; Abello
Caenolestoides miocaenicus n. gen., n. sp. Abello et al. 2021
Abello et al. 2021
Type: MPEF-PV 533 - was in Abello, 2013; Rincón et al., 2015; Engelman et al., 2016; Abello
Mammalia - Microbiotheria - Microbiotheriidae
Microbiotheriidae indet. (Ameghino 1887)
Goin and Abello 2013 50 specimens
Likely either _Clenia_ or _Oligobiotherium_. "Fragmentos mandibulares MPEF-PV 4727, MPEF-PV 5831, MPEF-PV 5866, MPEF-PV 4946, MPEF-PV 4947, MPEF-PV 4950, MPEF-PV 4970, MPEF- PV 6018 y MPEF-PV 6019; premolares aislados MPEF-PV 6020 y MPEF-PV 6021; fragmentos de molares inferiores MPEF-PV 4724, MPEF-PV 4726, MPEF-PV 4904, MPEF- PV 4924, MPEF-PV 4925, MPEF-PV 4926, MPEF-PV 4927, MPEF-PV 4928, MPEF-PV 4929, MPEF-PV 4936, MPEF-PV 4956, MPEF-PV 4958, MPEF-PV 4980, MPEF- PV 4981, MPEF-PV 4983, MPEF-PV 4984, MPEF-PV 4985 y MPEF-PV 6025; molares superiores (completos o fragmentarios) MPEF-PV 4911, MPEF-PV 4725, MPEF- PV 4978, MPEF-PV 4723, MPEF-PV 4722, MPEF-PV 4721, MPEF-PV 4720, MPEF-PV 4719, MPEF-PV 4718, MPEF-PV 4913, MPEF-PV 4979, MPEF-PV 4909, MPEF- PV 4953, MPEF-PV 4921, MPEF-PV 4977, MPEF-PV 4906, MPEF-PV 4907, MPEF-PV 4908, MPEF-PV 4905, MPEF-PV 4954 y MPEF-PV 4910."
Clenia minuscula n. gen., n. sp. Ameghino 1904
Goin and Abello 2013 34 specimens
MACN 52-381, dentary m1-2, type. Referred: "Fragmento mandibular izquierdo con el p2, MLP 82-V-2-88; fragmento mandibular derecho con m2–m3, MLP 83-III-I-204b; fragmento mandibular iz- quierdo con los alvéolos de los p3 y m1 completo, MPEF- PV 4746; fragmento mandibular izquierdo con p3-m2, MPEF-PV 4750; fragmento mandibular derecho sin dien- tes, MPEF-PV 4751; fragmento mandibular izquierdo con p3-m1, MPEF-PV 5677; fragmento mandibular derecho con el m4, MPEF-PV 5859; fragmento mandibular izquier- do con alvéolos y raíces de los p1–p3 y m4 y los m1–m3 completos (Fig. 3.1–2), MPEF-PV 6017; m1 izquierdo, MPEF-PV 4738; m1 izquierdo, MPEF-PV 4747; m1 iz- quierdo, MPEF-PV 4748; m1 izquierdo, MPEF-PV 4749; m1 izquierdo, MPEF-PV 4933; m1 derecho, MPEF-PV 4968; m1 izquierdo, MPEF-PV 6023; m2 derecho, MPEF- PV 4743; m2 izquierdo, MPEF-PV 4744; m2 izquierdo, MPEF-PV 4745; m2 izquierdo, MPEF-PV 4965; m2 iz- quierdo, MPEF-PV 4998; m3 izquierdo, MPEF-PV 4935; m3 izquierdo, MPEF-PV 4937; m3 derecho, MPEF-PV 4742; m3 derecho incompleto, MPEF-PV 4957; m3 iz- quierdo, MPEF-PV 5285; m4 derecho, MPEF-PV 4942; fragmento de maxilar izquierdo con P3–M1 (Fig. 3.3–4), MPEF-PV 4912; M1 derecho, MPEF-PV 4914; M1 dere- cho incompleto, MPEF-PV 4955; M2 izquierdo, MPEF-PV 4915; M2 izquierdo (Fig. 3.5–6), MPEF-PV 4974; M3 iz- quierdo, MPEF-PV 4916; MPEF-PV 6028, M3 izquierdo."
Microbiotherium sp. Ameghino 1887
Goin and Abello 2013 11 specimens
"Rama mandibular derecha con p3–m4 muy gastados, MLP 77-VI-13-13 (Fig. 3.16–17); fragmento de rama mandibular izquierda con m2 y alvéolos del m3, MPEF-PV 5302 (Fig. 3.18); m1 izquierdo, MPEF-PV 4941; m2 izquierdo, MPEF-PV 4728; m3 derecho, MPEF- PV 4923; m2 izquierdo, MPEF-PV 4966; m2 izquierdo, MPEF-PV 4967; m2 izquierdo, MPEF-PV 4987; m2 de- recho MPEF-PV 4988; m3 derecho, MPEF-PV 4991; m3 derecho, MPEF-PV 4992."
Eomicrobiotherium mykerum n. sp. Goin and Abello 2013
Goin and Abello 2013 20 specimens
"Holotipo. Fragmento mandibular derecho con raíces de los p2–p3, m1 completo, raíces del m2 y m3–m4 completos, MPEF-PV 5658 (Fig. 3.11). Material referido. m1 izquierdo, MPEF-PV 4944; m1 derecho, MPEF-PV 4964; m1 derecho MPEF-PV 4993; m2 izquierdo, MPEF- PV 4734; m2 izquierdo, MPEF-PV 4939; m2 izquierdo, MPEF-PV 4959; m2 izquierdo, MPEF-PV 4990; m2 dere- cho, MPEF-PV 6024; m3 izquierdo, MPEF-PV 4729; m3 izquierdo, MPEF-PV 4733; m3 izquierdo, MPEF-PV 4960; m4 izquierdo, MPEF-PV 4986; M2 izquierdo, MPEF-PV 4731; M1 derecho, MPEF-PV 4732 (Fig. 3.12–13); M2 de- recho, MPEF-PV 4917; M2 derecho, MPEF-PV 4972 (Fig. 3.14–15); M3 derecho, MPEF-PV 4973; M3 izquierdo, MPEF-PV 6027; M4 izquierdo MPEF-PV 4730."
Pachybiotherium acclinum n. gen., n. sp. Ameghino 1902
Marshall 1982 7 specimens
From ref. 57760 (Marshall 1982): MACN 52-370a, dentary p2-m1, m3, complete, type; 52-370b, dentary p3. From ref. 57989 (Marshall et al. 1984): MNHN COL 23, edentulous dentary. From ref. 58272 (Goin and Abello 2010): MLP 82-V-2-111, dentary p2-m3; 92-X-10-10, dentary m1-3; MPEF-PV 551, dentary p3-m1; MPEF-PV 6534, dentary m1-2
Pachybiotherium cf. acclinum Ameghino 1902
Goin and Abello 2013 2 specimens
MPEF-PV 5676b, edentulous dentary; MPEF-PV 5839, edentulous dentary
Oligobiotherium divisus n. gen., n. sp. Ameghino 1902
Marshall 1982 37 specimens
corrected as Oligobiotherium divisum
Described by ref. 57760: MACN 52-374, dentary m2-3, type; AMNH 29684, dentary p3-m2; 29708, dentary p3-m2; MMP M-217, dentary p1-3, m1; M-942, dentary p1-3; M-943, dentary p3-m2; M-951a, edentulous dentary; M-951b, edentulous dentary. Described by ref. 58272 (Goin and Abello 2010): "Fragmento mandibular izquierdo con al- véolos del i2–p3 y m1 completo, MLP 83-III-I-204a; frag- mento mandibular izquierdo con p3–m2 (Fig. 3.7–8), MLP 92-X-10-1,; fragmento mandibular derecho con m2, MPEF- PV 4962; fragmento mandibular derecho con m1, MPEF-PV 4989; m1 derecho, MPEF-PV 4741; m1 izquierdo, MPEF- PV 4931; m1 izquierdo, MPEF-PV 4940; m1 izquierdo, MPEF-PV 5284; m2 izquierdo, MPEF-PV 4739; m2 izquier- do, MPEF-PV 4934; m2 izquierdo, MPEF-PV 4943; m2 iz- quierdo, MPEF-PV 4961; m3 derecho, MPEF-PV 4932; m3 derecho, MPEF-PV 4938; m3 izquierdo, MPEF-PV 4963; m4 derecho, MPEF-PV 4930; m4 derecho, MPEF-PV 5281; fragmento de maxilar derecho con M2–M3 (Fig. 3.9–10), MPEF-PV 5661; fragmento maxilar derecho con M3–M4, MPEF-PV 6026; M1 derecho, MPEF-PV 4737; M1 derecho, MPEF-PV 4919; M1 derecho, MPEF-PV 4920; M1 derecho, MPEF-PV 4971; M1 izquierdo, MPEF-PV 4975; M2 iz- quierdo, MPEF-PV 4736; M2 derecho, MPEF-PV 5282; M2 izquierdo, MPEF-PV 5283; M3 derecho, MPEF-PV 4918; M3 izquierdo, MPEF-PV 4922."
Microbiotherium hernandezi n. sp. Simpson 1932
Marshall 1982 1 specimen
synonym of Oligobiotherium divisum
AMNH 29664, dentary p2-m4, type
Mammalia - Sparassodonta - Thylacosmilidae
Thylacosmilidae indet. Riggs 1933
Goin et al. 2007 1 specimen
"MLP 92-X-10-6 un molar superior derecho (M?3)."
Mammalia - Sparassodonta - Hathliacynidae
Hathliacynidae indet. Ameghino 1894
Goin et al. 2007 4 specimens
corrected as Hathlyacininae
"MPEF-PV 4771, un premolar inferior aislado." "MPEF-PV 4767, un talónido derecho aislado." "MPEF-PV 4766 (figura 6.K), un molar superior derecho (M?1)." "MPEF-PV 4770, una hemimandíbula casi completa con los i3-m4 (figura 6.L)."
Mammalia - Sparassodonta
Cladosictis sp. Ameghino 1887
Goin et al. 2007 2 specimens
"MLP 82-V-2-109, un fragmento mandibular derecho con los m1-2 completos." "MPEF-PV 4768, un molar inferior derecho aislado (m?3)."
Cladosictis centralis n. sp. Ameghino 1902
Marshall et al. 1984 5 specimens
From ref. 58306 (Marshall, 1981): MACN 11639, partial skull, type. From ref. 57989 (Marshall et al. 1984): MNHN COL 4, mandible with right c-m4 and left p2-m4; MNHN COL 122, m3. From ref. 58052 (Goin et al. 2007): MPEF-PV 4772, M2 or M3 fragment; MPEF-PV 5854, M1
Sipalocyon sp. Ameghino 1887
Goin et al. 2007 2 specimens
"MPEF-PV 4769 (figuras 6.E-F), un molar inferior derecho aislado (m4)." "MPEF-PV 4545 (figura 6.G), un M3 izquierdo aislado."
Sipalocyon externus n. sp. (Ameghino 1902)
Goin et al. 2007 5 specimens
Reported by ref. 58306 (Marshall, 1981): MACN 52-383, partial skull, holotype. Reported by ref. 58052: "MPEF-PV 5840, un fragmento mandibular iz- quierdo con los m1-2 completos (figura 6.A); MPEF-PV 5832, un fragmento de maxilar izquierdo con el P2, un M3 sin el protocono y el M4 completo (figuras 6.B-C), todos probablemente asociados; MLP 77-VI-13-7, un fragmento de M2 aislado; MLP 82-V-2-85, fragmento de maxilar derecho con el M1 completo (figura 6.D)."
Anatherium herrerae n. sp. Marshall 1981
Marshall 1981 1 specimen
synonym of Acyon tricuspidatus
FMNH P13521, left and right dentaries, holotype
Borhyaeninae indet. Ameghino 1894
Goin et al. 2007 6 specimens
"MLP 85-VII-3-46, un fragmento de maxilar izquierdo con parte del alvéolo del canino, el P1 completo y las raíces de los P2-3; MLP 82-V-2-4b, un incisivo inferior aislado; MPEF-PV 5637a, un incisivo inferior aislado; MPEF-PV 5637c, un fragmento man- dibular derecho con los alvéolos del p2; MPEF-PV 5637b, un frag- mento de m2 derecho; MPEF-PV 5846b, un P3 izquierdo aislado."
Arctodictis sinclairi Marshall 1978
Goin et al. 2007 9 specimens
"MLP 85-VII-3-1, un cráneo y mandíbulas casi completos con buena parte del esqueleto postcraneano asociado; MLP 93-XI-18-9, un canino superior derecho aislado; MPEF-PV 5846c, un M1 izquierdo aislado, fragmentario (figura 7.H); MPEF- PV 4752, un fragmento mandibular izquierdo con los m3-4 com- pletos (figuras 7.I, J); MLP 82-V-2-4c, un canino superior izquierdo aislado; MLP 77-VI-13-27, un m2 izquierdo aislado; MLP 82-V-2- 4a, un m2 derecho aislado; MPEF-PV 5846d, un P2 derecho aislado; MLP 82-V-2-116, un P3 derecho aislado."
cf. Arctodictis sinclairi Marshall 1978
Goin et al. 2007 3 specimens
"MLP 77-VI-13-1, una hemimandíbula izquierda casi completa con los i1-m4 completos, una hemimandíbula dere- cha con los i1 e i3, y los c-m1 completos, y un canino superior ais- lado, probablemente asociado; MLP 84-III-15-1, fragmento de ra- ma mandibular derecha con los m2-4 completos y un M1 derecho aislado asociado; MPEF-PV 5846a, un p2 izquierdo aislado."
Pseudoborhyaena macrodonta n. sp. Ameghino 1902
Goin et al. 2007 4 specimens
recombined as Borhyaena macrodonta
"MACN 52-390 (tipo de Pseudoborhyaena macrodonta Ameghino 1902), un resto craneano con los P2-M4 izquierdos, aso- ciado a un fragmento mandibular derecho con la raíz del p1, los p2-m3 completos y un fragmento del m4." "MLP 82-V-4-3, fragmento de maxi- lar derecho con los P2-3 completos (figuras 7.D-E); MLP 92-X-10-8, un m2 izquierdo aislado (figura 7.F); MPEF-PV 1467, fragmento de maxilar izquierdo con los P3-M3 (figura 7.G, detalle del M1)."
Pseudoborhyaena longaeva n. sp. Ameghino 1902
Goin et al. 2007 1 specimen
synonym of Borhyaena macrodonta
"MACN 52-366 (tipo de Pseudoborhyaena longeva Ameghino, 1902), un fragmento mandibular derecho con los c-p1 completos, parte del p2, raíces del p3, m1 completo, raíces del m2 y los m3-4 parcialmente rotos"
cf. Acrocyon riggsi (Sinclair 1930)
Marshall et al. 1984 1 specimen
MNHN COL 125, P3
Pseudothylacynus rectus n. gen., n. sp. Ameghino 1902
Goin et al. 2007 6 specimens
MNHN COL 5, dentary p3-m4, isolated m2, isolated m4; MPEF-PV 5838, incisor, p1, p?3, premolar; MLP 92-X-10-7, maxillary M1-2; MPEF-PV 4765, dentary m3-4; MLP 92-X-10-5, dentary p2-3. Described by ref. 58324 (Marshall, 1979): MACN 52-369, dentary p1-m4, holotype
Mammalia - Argyrolagidae
Proargyrolagus argentinus n. sp. Goin and Abello 2013
Goin and Abello 2013 24 specimens
"Holotipo. M2 derecho, MPEF-PV 4711 (Fig. 4.9–10). Material referido. M1 derecho, MPEF-PV 4715; M1 dere- cho, MPEF-PV 4714 (Fig. 4.7–8); M1 derecho, MPEF-PV 4786; M2 derecho MPEF-PV 4781; M2 izquierdo MPEF- PV 4547; M3 izquierdo, MPEF-PV 4713 (Fig. 4.11–12); M3 derecho, MPEF 4783-PV; M?4 derecho, MPEF-PV 4782; Mx izquierdo, MPEF-PV 4784; Mx izquierdo, MPEF-PV 4785; m1 izquierdo, MPEF-PV 4712 (Fig. 4.1– 2); m2 izquierdo, MPEF-PV 4778 (Fig. 4.3–4); m2 izquier- do, MPEF-PV 4789; m2 derecho, MPEF-PV 4776; m3 izquierdo, MPEF-PV 4716; m3 derecho, MPEF-PV 4710 (Fig. 4.5–6); m3 derecho, MPEF-PV 4788; m3 derecho, MPEF-PV 4790; m4 derecho, MPEF-PV 4717; mx dere- cho, MPEF-PV 4780; mx izquierdo, MPEF-PV 4787; mx derecho, MPEF-PV 4775."
Mammalia - Meridiolestida
Necrolestes mirabilis n. sp. Goin et al. 2007
Goin et al. 2007 6 specimens
MLP 92-X-10-14, mandibular fragment with p?4 and dp?4, holotype; MPEF-PV 4706, P?4; MPEF-PV 6030, m?1; MPEF-PV 4548, Mx; MPEF-PV 4549, upper premolariform; MPEF-PV 4546, upper premolariform
see common names

Geography
Country:Argentina State/province:Chubut
Coordinates: 45.7° South, 68.7° West (view map)
Paleocoordinates:46.4° South, 62.8° West
Basis of coordinate:estimated from map
Altitude:257 meters
Geographic resolution:local area
Time
Period:Neogene Epoch:Miocene
10 m.y. bin:Cenozoic 5
Key time interval:Colhuehuapian Mammal zone: Colhuehuapian
Age range of interval:21.00000 - 17.50000 m.y. ago
Age estimate:20.2 to 20 Ma (Ar/Ar)
Stratigraphy
Formation:Sarmiento Member:Colhué Huapí
Stratigraphic resolution:member
Stratigraphy comments: Busker, et al. 2019: Colhuehuapian SALMA (early Miocene) (Flynn and Swisher, 1995; Simpson, 1940; Vucetich et al., 2010b).

Novo et al 2018: The Colhue-Huapi Member of the Sarmiento Formation is, early Miocene (Burdigalian Age) (see Simpson 1964; Madden & Scarano 2010 and references therein), approximately 20.0-20.2 Ma (Ré et al. 2010).

Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:paleoecologic analysis
Museum repositories:MLP,MNHN
Collectors:Tournouër and others
Collection method comments: Busker, et al. 2019: Only MPEF-PV 5421 comes from Gran Barranca locality, Chubut Province
Metadata
Database number:141368
Authorizer:C. Jaramillo, J. Zijlstra, J. Carrillo, P. Mannion Enterer:J. Carrillo, P. Mannion, K. Pino, M. Kouvari, G. Varnham, J. Zijlstra
Modifier:M. Kouvari Research group:vertebrate
Collections that are a subset of this one:176236, 176237, 176238, 176239, 176240
Created:2013-03-26 08:17:32 Last modified:2020-11-30 05:46:45
Access level:database members Released:2014-03-26 08:17:32
Creative Commons license:CC BY
Reference information

Primary reference:

45974. M. G. Vucetich and A. G. Kramarz. 2003. New Miocene rodents from Patagonia (Argentina) and their bearing on the early radiation of the Octodontoids (Hystricognathi). Journal of Vertebrate Paleontology 23(2):435-444 [C. Jaramillo/J. Carrillo]

Secondary references:

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69887 F. Busker, M. E. Pérez, and M. T. Dozo. 2019. A new chinchilloid (Rodentia, Hystricognathi) from the early Miocene of the localities of Bryn Gwyn and Gran Barranca (Patagonia, Argentina). Comptes Rendus Palevol 18(5):525-540 [P. Mannion/M. Kouvari/E. Vlachos]
58217 A. M. Candela. 1999. The evolution of the molar pattern of the Erethizontidae (Rodentia, Hystricognathi) and the validity of Parasteiromys Ameghino, 1904. Palaeovertebrata 28(1):53-73 [J. Zijlstra/J. Zijlstra]
71055 A. M. Candela. 2003. A new porcupine (Rodentia, Hystricognathi, Erethizontidae) from the Early and Middle Miocene of Patagonia. Ameghiniana 40(3):483-493 [P. Mannion/M. Kouvari]
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69517 S. Hernández Del Pino, M. G. Fernández, E. Cerdeño and J. C. Fernicola. 2019. Anatomy and systematics of Notohippus toxodontoides Ameghino, 1891 (Mammalia, Notoungulata), from the Miocene of Santa Cruz Province, Argentina. Journal of Vertebrate Paleontology e1577870 [P. Mannion/G. Varnham]
73094 P. Hershkovitz. 1981. Comparative Anatomy of Platyrrhine Mandibular Cheek Teeth dpm4, pm4, m1 with Particular Reference to Those of Homunculus (Cebidae), and Comments on Platyrrhine Origins. Folia Primatologica 35:179-217 [P. Mannion/G. Varnham]
57939 R. F. Kay. 2010. A new primate from the early Miocene of Gran Barranca, Chubut Province, Argentina: paleoecological implications. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 220-239 [J. Zijlstra/J. Zijlstra]
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57896 G. M. López, A. M. Ribeiro, and M. Bond. 2010. The Notohippidae (Mammalia, Notoungulata) from Gran Barranca: preliminary considerations. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 143-151 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
57760 L. Marshall. 1982. Systematics of the South American marsupial family Microbiotheriidae. Fieldiana Geology 10:1-75 [J. Zijlstra/J. Zijlstra]
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57989 L. G. Marshall, C. de Muizon, and R. Hoffstetter. 1984. Fossil Marsupialia in the Muséum national d'Histoire naturelle collected by André Tournouër from Patagonia, southern Argentina. Bulletin du Muséum national d'histoire naturelle 6(1):33-58 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
72535 N. M. Novo, M. F. Tejedor, and L. R. González-Ruiz. 2018. Previously unknown fossil platyrrhines (Primates) of Patagonia from the Tournouër collection at the Muséum national d'Histoire naturelle, Paris. Geodiversitas 40(22):529-535 [P. Mannion/M. Kouvari]
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57933 M. G. Vucetich, A. G. Kramarz, and A. M. Candela. 2010. Colhuehuapian rodents from Gran Barranca and other Patagonian localities: the state of the art. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 206-219 [J. Zijlstra/J. Zijlstra]