Gran Barranca - southern cliff of lake Colhué Huapí: Colhuehuapian, Argentina
collected by Tournouër and others
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia
- Rodentia
- Chinchillidae
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Eoviscaccia australis
Vucetich 1989
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Vucetich et al. 2010 | |||||||||
Mammalia
- Rodentia
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Chinchilloidea indet.
Bennett 1833
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Vucetich et al. 2010 | |||||||||
Mammalia
- Rodentia
- Neoepiblemidae
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Neoepiblemidae indet.
Kraglievich 1926
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Vera et al. 2015 | 1 specimen | ||||||||
MGP-PD 29099, mandible p4-m2 | ||||||||||
Perimys incavatus
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Perimys transversus
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Perimys dissimilis
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Mammalia
- Rodentia
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Saremmys ligcura
Busker et al. 2019
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Busker et al. 2019 | |||||||||
Luantus initialis
(Ameghino 1902)
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Vucetich et al. 2010 | |||||||||
original and current combination Luanthus initialis | ||||||||||
Soriamys sp.
Kramarz 2001
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Vucetich et al. 2010 | |||||||||
? Eosteiromys sp.
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Eosteiromys segregatus
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Eosteiromys homogenidens
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Mammalia
- Rodentia
- Erethizontidae
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Parasteiromys friantae n. sp.
Candela 1999
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Candela 1999 | 1 specimen | ||||||||
MPEF 5501, isolated upper right molariform | ||||||||||
Parasteiromys uniformis n. sp.
Ameghino 1904
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Candela 1999 | 1 specimen | ||||||||
MACN A 52-178, maxillary with DP4-M1, holotype | ||||||||||
Hypsosteiromys sp.
Patterson 1958
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Dozo et al. 2004 | 1 specimen | ||||||||
MPEF-PV 6029, skull | ||||||||||
Steiromys axiculus n. sp.
Ameghino 1902
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Patterson 1958 | 1 specimen | ||||||||
recombined as Hypsosteiromys axiculus | ||||||||||
MACN A 52-171, dentary m1-3, holotype | ||||||||||
Steiromys nectus n. sp.
Ameghino 1902
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Candela and Vucetich 2002 | 4 specimens | ||||||||
recombined as Hypsosteiromys nectus | ||||||||||
MACN A 52-177, dentary dp4-m2, holotype; MACN A 52-175A, p4; MNHN col. 50, p4; MLP 85-VII-3-4, m1 or m2 | ||||||||||
Steiromys tabulatus n. sp.
Ameghino 1902
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Patterson 1958 | 1 specimen | ||||||||
synonym of Hypsosteiromys nectus | ||||||||||
MACN A 52-172, dentary p4-m3, lectotype | ||||||||||
Branisamyopsis australis
Candela 2003
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Vucetich et al. 2010 | |||||||||
Branisamyopsis australis n. gen., n. sp.
Candela 2003
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Candela 2003 | |||||||||
Holotype: MMP 937 | ||||||||||
Mammalia
- Rodentia
- Cephalomyidae
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? Cephalomyidae indet.
Ameghino 1897
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Vucetich et al. 2010 | |||||||||
Mammalia
- Rodentia
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Octodontoidea informal Gen. nov. sp. large
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Vucetich et al. 2010 | |||||||||
Mammalia
- Rodentia
- Acaremyidae
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Acaremys cf. murinus
Ameghino 1887
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(6 measurements) | ||||||||||
Galileomys antelucanus n. gen., n. sp.
Vucetich and Kramarz 2003
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(5 measurements) | ||||||||||
Mammalia
- Rodentia
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Caviocricetus lucasi
Vucetich and Verzi 1996
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Vucetich et al. 2010 | |||||||||
Acarechimys pulchellus n. sp.
(Ameghino 1902)
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Vucetich et al. 2010 | |||||||||
synonym of Acarechimys gracilis | ||||||||||
Holotype. MACN A 52-128 | ||||||||||
Acarechimys cf. minutissimus
Ameghino 1887
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Vucetich et al. 2010 | |||||||||
Prospaniomys informal sp. 1
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Prospaniomys priscus
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Protadelphomys latus
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Mammalia
- Rodentia
- Echimyidae
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Paradelphomys fissus n. gen., n. sp.
Patterson and Pascual 1968
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Patterson and Pascual 1968 | 1 specimen | ||||||||
MLP 125, dentary dp4-m1, type | ||||||||||
Mammalia
- Rodentia
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? Protacaremys sp.
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Protacaremys avunculus
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Protacaremys prior
Ameghino 1902
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Vucetich et al. 2010 | |||||||||
Mammalia
- Primates
- Homunculidae
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Mazzonicebus almendrae n. gen., n. sp.
Kay 2010
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Kay 2010 | 33 specimens | ||||||||
Teeth, maxillaries, and mandibles in MLP and MPEF-PV. Several stratigraphic levels. | ||||||||||
Mazzonicebus almendrae
Kay 2010
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Novo et al. 2018 | |||||||||
MNHN.F.COL93a and b. The specimen label reads (Fig. 3G): “Primate, col 93 // Colhué Huapi B // M.A. Tournouër, 1899-7”. Both specimens were found in the box with the same label | ||||||||||
Homunculus sp.
Ameghino 1891
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Hershkovitz 1981 | |||||||||
No. 69-111-12-1, lower jaw | ||||||||||
Mammalia
- Chiroptera
- Molossidae
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Mormopterus indet.
Peters 1865
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Czaplewski 2010 | 1 specimen | ||||||||
MLP 93-XI-18-19, right dentary fragment with m2 | ||||||||||
Mormopterus barrancae n. sp.
Czaplewski 2010
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Czaplewski 2010 | 2 specimens | ||||||||
MLP 93-XI-18-15, mandible with p2, p4, m1, m2, holotype, from "nivel 'C' (gris)"; MLP 93-XI-18-18, left m1 | ||||||||||
Mammalia
- Chiroptera
- Phyllostomidae
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Phyllostominae indet.
Gray 1825
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Czaplewski 2010 | 1 specimen | ||||||||
MLP 93-XI-18-16, right m3 | ||||||||||
Mammalia
- Panameriungulata
- Adianthidae
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Adianthus patagonicus n. sp.
Ameghino 1904
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Cifelli and Soria 1983 | 1 specimen | ||||||||
recombined as Proheptaconus patagonicus | ||||||||||
MACN A52-218, M3, holotype | ||||||||||
Mammalia
- Panameriungulata
- Proterotheriidae
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Prolicaphrium spectabile
Ameghino 1902
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Vera et al. 2015 | 1 specimen | ||||||||
MGP-PD 29096, upper molar | ||||||||||
Prolicaphrium festinum
Ameghino 1902
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Vera et al. 2015 | 2 specimens | ||||||||
recombined as Licaphrops festinus | ||||||||||
MGP-PD 29024, mandible with two molars; MGP-PD 26591: mandible with left m1-3 | ||||||||||
Mammalia
- Panameriungulata
- Macraucheniidae
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Pternoconius tournoueri n. sp.
Soria and Hoffstetter 1985
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Soria and Hoffstetter 1985 | |||||||||
Holotype: MNHN COL 121 | ||||||||||
Cramauchenia indet.
Ameghino 1902
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Vera et al. 2015 | 3 specimens | ||||||||
MGP-PD 29040 A, M3; MGP-PD 29060 A, lower molar, and B, astragalus | ||||||||||
Cramauchenia normalis n. gen., n. sp.
Ameghino 1902
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Dozo and Vera 2010 | |||||||||
MACN A-52-119 - holotype | ||||||||||
Mammalia
- Cingulata
- Glyptodontidae
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Glyptodontidae indet.
Burmeister 1879
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Vera et al. 2015 | 1 specimen | ||||||||
MGP-PD 31693, proximal ulna | ||||||||||
Mammalia
- Cingulata
- Propalaehoplophoridae
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Propalaehoplophorinae indet.
(Ameghino 1891)
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Carlini et al. 2005 | |||||||||
original and current combination Propalaehoplophoridae | ||||||||||
Mammalia
- Cingulata
- Peltephilidae
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Peltephilidae indet.
Ameghino 1894
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Carlini et al. 2005 | |||||||||
Mammalia
- Cingulata
- Dasypodidae
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Dasypodidae indet.
(Gray 1821)
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Vera et al. 2015 | 2 specimens | ||||||||
MGP-PD 29101, bony external plates; MGP-PD 29076 A, astragalus | ||||||||||
Pseudostegotherium glangeaudi
Ameghino 1902
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Carlini et al. 2010 | 16 specimens | ||||||||
MACN A-12679, osteroderms of the carapace and left mandibular fragment with two teeth; osteoderms in MPEF PV | ||||||||||
Mammalia
- Cingulata
- Chlamyphoridae
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Prozaedyus humilis
Ameghino 1902
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Carlini et al. 2010 | |||||||||
Prozaedyus cf. humilis
Ameghino 1902
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Carlini et al. 2005 | |||||||||
Proeutatus sp.
Ameghino 1891
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Carlini et al. 2010 | |||||||||
Proeutatus cf. robustus
Scott 1903
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Carlini et al. 2005 | |||||||||
Proeutatus oenophorus
(Ameghino 1887)
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Carlini et al. 2005 | |||||||||
recombined as Proeutatus oenophorum | ||||||||||
Mammalia
- Megalonychidae
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Megalonychidae indet.
Gervais 1855
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Vera et al. 2015 | 1 specimen | ||||||||
MGP-PD 31692, Mc? II or III | ||||||||||
Proschismotherium sp.
Ameghino 1902
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Carlini et al. 2005 | |||||||||
Hapaloides sp.
Ameghino 1902
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Carlini et al. 2005 | |||||||||
Holomegalonyx sp.
Scillato-Yané 1979
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Carlini et al. 2005 | |||||||||
Mammalia
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Stegotherium pascuali n. sp.
Fernicola and Vizcaino 2008
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Fernicola and Vizcaino 2008 | |||||||||
Holotipo. MACN A-12680d. Found in level 5 | ||||||||||
Stegotherium caroloameghinoi n. sp.
Fernicola and Vizcaino 2008
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Fernicola and Vizcaino 2008 | |||||||||
Holotipo. MACN-A 10443a. Found in level 5 | ||||||||||
Stegotherium variegatum
Ameghino 1902
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Carlini et al. 2010 | |||||||||
Stegotherium variegatum
Ameghino 1902
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Carlini et al. 2005 | |||||||||
Mammalia
- Notoungulata
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Argyrohippus fraterculus
Ameghino 1902
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Hernández Del Pino et al. 2019 | |||||||||
MACN A 52-17 (lectotype), right mandible and symphysis with left i2–i3–c and right i1–i3–c, alveoli of p1–p2 and p3–m3; MACN A 52-18, right maxillary fragment with P4–M3 (left) and left maxillary fragment (reversed; right) with alveoli of P1 and P2–P4 | ||||||||||
Argyrohippus cf. boulei
Ameghino 1902
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López et al. 2010 | 3 specimens | ||||||||
synonym of Argyrohippus fraterculus | ||||||||||
MPEF-PV 5388, mandible with left and right i1-3, c, p2-m3; MPEF-PV 5922, fragment of upper molar and left m1; MPEF-PV 7325, left lower molar | ||||||||||
Argyrohippus boulei
Ameghino 1902
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Hernández Del Pino et al. 2019 | |||||||||
synonym of Argyrohippus fraterculus | ||||||||||
MACN A 52-12 (lectotype) | ||||||||||
Mammalia
- Notoungulata
- Leontiniidae
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Leontiniidae indet.
Ameghino 1894
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Vera et al. 2015 | 3 specimens | ||||||||
"MGP-PD 29107: A, fragment of mandible with left m3; B, left lower molar; C, left lower molar" | ||||||||||
Colpodon sp.
Burmeister 1885
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Vera et al. 2015 | 1 specimen | ||||||||
"MGP-PD 29076B: mandible with left m1-2" | ||||||||||
Colpodon distinctus n. sp.
Ameghino 1902
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Ribeiro et al. 2010 | 21 specimens | ||||||||
Maxilla, dentary, skull and tooth fragments in MACN, MPEF-PV, AMNH, and FMNH. From localities GBV-8, GBV-9, GBV-10, GBV-38, and GBV-43. | ||||||||||
Mammalia
- Notoungulata
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Pachyrukhos politus
Ameghino 1901
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Seoane and Cerdeño 2019 | |||||||||
MLP 82-V-2-107, right mandibular fragment with p2–m3; and MLP 85VII-3-28, right mandibular fragment with p2–p3 | ||||||||||
Mammalia
- Notoungulata
- Interatheriidae
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Cochilius volvens
Ameghino 1902
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Simpson 1932 | |||||||||
AMNH 29651, 29686, 29652, 29653, 29658, 29654, 29655, 29657 | ||||||||||
Mammalia
- Astrapotheria
- Astrapotheriidae
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Astrapotheriidae informal gen. et sp. nov.
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Kramarz and Bond 2010 | 1 specimen | ||||||||
MACN A 52-522b, isolated canine, syntype of _Parastrapotherium crassum_. Species appears to also occur at Cerro Bandera and may be uruguaytheriine. | ||||||||||
Astrapotheriidae indet.
Ameghino 1887
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Vera et al. 2015 | 9 specimens | ||||||||
"MGP-PD 29049 A, mandible with left m3; B, right P2; C, right P1; D, right upper molar, E, canine?; F, lingual fragment of left upper molar; G, fragments of tooth; MGP-PD 29075: two teeth; MGP-PD 29108: teeth" | ||||||||||
? Astrapotherium ruderarium n. sp.
(Ameghino 1902)
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Kramarz and Bond 2010 | |||||||||
Parastrapotherium symmetrum n. sp.
(Ameghino 1902)
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Kramarz and Bond 2010 | 2 specimens | ||||||||
MACN A 52-507a, holotype, isolated lower incisor, from "_Colpodon_ beds at Gran Barranca". MPEF PV 7923, isolated lower incisor, from Gran Barranca Colhuehuapian West locality, level B | ||||||||||
Mammalia
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Marsupialia indet.
(Illiger 1811)
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Goin et al. 2007 | 2 specimens | ||||||||
"Didelphimorphia o Sparassodonta gen. et sp. indet." MLP 77-VI-13-8, trigonid; MLP 92-X-10-11, M4 | ||||||||||
Mammalia
- Didelphimorphia
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Didelphimorphia indet.
Gill 1872
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Goin et al. 2007 | 1 specimen | ||||||||
MLP 77-VI-13-25, left dentary fragment with c, p2, half of p3, fragments of m2-3, and edentulous right dentary | ||||||||||
Didelphoidea indet.
Gray 1821
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Goin and Abello 2013 | 1 specimen | ||||||||
MLP 77-VI-13-26, left maxillary fragment with part of the plate, roots of P1 and P3, and complete P2 and M1-4 | ||||||||||
Mammalia
- Didelphimorphia
- Didelphidae
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Didelphidae indet.
Gray 1821
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Goin et al. 2007 | 4 specimens | ||||||||
MPEF-PV 4774, m2 or m3; MPEF-PV 4709, m?1; MPEF-PV 4707, Mx; MPEF-PV 4773, m?1 fragment | ||||||||||
Mammalia
- Didelphimorphia
- Caluromyidae
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? Caluromyidae indet.
Kirsch and Reig 1977
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Goin et al. 2007 | 1 specimen | ||||||||
spelled with current rank as Caluromyinae | ||||||||||
MPEF-PV 4708, mx | ||||||||||
Mammalia
- Paucituberculata
- Palaeothentidae
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Parabderites bicrispatus
Ameghino 1902
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Marshall et al. 1984 | 1 specimen | ||||||||
MNHN COL 124, dentary m2 | ||||||||||
Mammalia
- Paucituberculata
- Caenolestidae
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Stilotherium parvum n. sp.
Abello et al. 2021
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Abello et al. 2021 | |||||||||
Type: MPEF-PV 5830 - was |
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Caenolestoides miocaenicus n. gen., n. sp.
Abello et al. 2021
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Abello et al. 2021 | |||||||||
Type: MPEF-PV 533 - was |
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Mammalia
- Microbiotheria
- Microbiotheriidae
|
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Microbiotheriidae indet.
(Ameghino 1887)
|
Goin and Abello 2013 | 50 specimens | ||||||||
Likely either _Clenia_ or _Oligobiotherium_. "Fragmentos mandibulares MPEF-PV 4727, MPEF-PV 5831, MPEF-PV 5866, MPEF-PV 4946, MPEF-PV 4947, MPEF-PV 4950, MPEF-PV 4970, MPEF- PV 6018 y MPEF-PV 6019; premolares aislados MPEF-PV 6020 y MPEF-PV 6021; fragmentos de molares inferiores MPEF-PV 4724, MPEF-PV 4726, MPEF-PV 4904, MPEF- PV 4924, MPEF-PV 4925, MPEF-PV 4926, MPEF-PV 4927, MPEF-PV 4928, MPEF-PV 4929, MPEF-PV 4936, MPEF-PV 4956, MPEF-PV 4958, MPEF-PV 4980, MPEF- PV 4981, MPEF-PV 4983, MPEF-PV 4984, MPEF-PV 4985 y MPEF-PV 6025; molares superiores (completos o fragmentarios) MPEF-PV 4911, MPEF-PV 4725, MPEF- PV 4978, MPEF-PV 4723, MPEF-PV 4722, MPEF-PV 4721, MPEF-PV 4720, MPEF-PV 4719, MPEF-PV 4718, MPEF-PV 4913, MPEF-PV 4979, MPEF-PV 4909, MPEF- PV 4953, MPEF-PV 4921, MPEF-PV 4977, MPEF-PV 4906, MPEF-PV 4907, MPEF-PV 4908, MPEF-PV 4905, MPEF-PV 4954 y MPEF-PV 4910." | ||||||||||
Clenia minuscula n. gen., n. sp.
Ameghino 1904
|
Goin and Abello 2013 | 34 specimens | ||||||||
MACN 52-381, dentary m1-2, type. Referred: "Fragmento mandibular izquierdo con el p2, MLP 82-V-2-88; fragmento mandibular derecho con m2–m3, MLP 83-III-I-204b; fragmento mandibular iz- quierdo con los alvéolos de los p3 y m1 completo, MPEF- PV 4746; fragmento mandibular izquierdo con p3-m2, MPEF-PV 4750; fragmento mandibular derecho sin dien- tes, MPEF-PV 4751; fragmento mandibular izquierdo con p3-m1, MPEF-PV 5677; fragmento mandibular derecho con el m4, MPEF-PV 5859; fragmento mandibular izquier- do con alvéolos y raíces de los p1–p3 y m4 y los m1–m3 completos (Fig. 3.1–2), MPEF-PV 6017; m1 izquierdo, MPEF-PV 4738; m1 izquierdo, MPEF-PV 4747; m1 iz- quierdo, MPEF-PV 4748; m1 izquierdo, MPEF-PV 4749; m1 izquierdo, MPEF-PV 4933; m1 derecho, MPEF-PV 4968; m1 izquierdo, MPEF-PV 6023; m2 derecho, MPEF- PV 4743; m2 izquierdo, MPEF-PV 4744; m2 izquierdo, MPEF-PV 4745; m2 izquierdo, MPEF-PV 4965; m2 iz- quierdo, MPEF-PV 4998; m3 izquierdo, MPEF-PV 4935; m3 izquierdo, MPEF-PV 4937; m3 derecho, MPEF-PV 4742; m3 derecho incompleto, MPEF-PV 4957; m3 iz- quierdo, MPEF-PV 5285; m4 derecho, MPEF-PV 4942; fragmento de maxilar izquierdo con P3–M1 (Fig. 3.3–4), MPEF-PV 4912; M1 derecho, MPEF-PV 4914; M1 dere- cho incompleto, MPEF-PV 4955; M2 izquierdo, MPEF-PV 4915; M2 izquierdo (Fig. 3.5–6), MPEF-PV 4974; M3 iz- quierdo, MPEF-PV 4916; MPEF-PV 6028, M3 izquierdo." | ||||||||||
Microbiotherium sp.
Ameghino 1887
|
Goin and Abello 2013 | 11 specimens | ||||||||
"Rama mandibular derecha con p3–m4 muy gastados, MLP 77-VI-13-13 (Fig. 3.16–17); fragmento de rama mandibular izquierda con m2 y alvéolos del m3, MPEF-PV 5302 (Fig. 3.18); m1 izquierdo, MPEF-PV 4941; m2 izquierdo, MPEF-PV 4728; m3 derecho, MPEF- PV 4923; m2 izquierdo, MPEF-PV 4966; m2 izquierdo, MPEF-PV 4967; m2 izquierdo, MPEF-PV 4987; m2 de- recho MPEF-PV 4988; m3 derecho, MPEF-PV 4991; m3 derecho, MPEF-PV 4992." | ||||||||||
Eomicrobiotherium mykerum n. sp.
Goin and Abello 2013
|
Goin and Abello 2013 | 20 specimens | ||||||||
"Holotipo. Fragmento mandibular derecho con raíces de los p2–p3, m1 completo, raíces del m2 y m3–m4 completos, MPEF-PV 5658 (Fig. 3.11). Material referido. m1 izquierdo, MPEF-PV 4944; m1 derecho, MPEF-PV 4964; m1 derecho MPEF-PV 4993; m2 izquierdo, MPEF- PV 4734; m2 izquierdo, MPEF-PV 4939; m2 izquierdo, MPEF-PV 4959; m2 izquierdo, MPEF-PV 4990; m2 dere- cho, MPEF-PV 6024; m3 izquierdo, MPEF-PV 4729; m3 izquierdo, MPEF-PV 4733; m3 izquierdo, MPEF-PV 4960; m4 izquierdo, MPEF-PV 4986; M2 izquierdo, MPEF-PV 4731; M1 derecho, MPEF-PV 4732 (Fig. 3.12–13); M2 de- recho, MPEF-PV 4917; M2 derecho, MPEF-PV 4972 (Fig. 3.14–15); M3 derecho, MPEF-PV 4973; M3 izquierdo, MPEF-PV 6027; M4 izquierdo MPEF-PV 4730." | ||||||||||
Pachybiotherium acclinum n. gen., n. sp.
Ameghino 1902
|
Marshall 1982 | 7 specimens | ||||||||
From ref. 57760 (Marshall 1982): MACN 52-370a, dentary p2-m1, m3, complete, type; 52-370b, dentary p3. From ref. 57989 (Marshall et al. 1984): MNHN COL 23, edentulous dentary. From ref. 58272 (Goin and Abello 2010): MLP 82-V-2-111, dentary p2-m3; 92-X-10-10, dentary m1-3; MPEF-PV 551, dentary p3-m1; MPEF-PV 6534, dentary m1-2 | ||||||||||
Pachybiotherium cf. acclinum
Ameghino 1902
|
Goin and Abello 2013 | 2 specimens | ||||||||
MPEF-PV 5676b, edentulous dentary; MPEF-PV 5839, edentulous dentary | ||||||||||
Oligobiotherium divisus n. gen., n. sp.
Ameghino 1902
|
Marshall 1982 | 37 specimens | ||||||||
corrected as Oligobiotherium divisum | ||||||||||
Described by ref. 57760: MACN 52-374, dentary m2-3, type; AMNH 29684, dentary p3-m2; 29708, dentary p3-m2; MMP M-217, dentary p1-3, m1; M-942, dentary p1-3; M-943, dentary p3-m2; M-951a, edentulous dentary; M-951b, edentulous dentary. Described by ref. 58272 (Goin and Abello 2010): "Fragmento mandibular izquierdo con al- véolos del i2–p3 y m1 completo, MLP 83-III-I-204a; frag- mento mandibular izquierdo con p3–m2 (Fig. 3.7–8), MLP 92-X-10-1,; fragmento mandibular derecho con m2, MPEF- PV 4962; fragmento mandibular derecho con m1, MPEF-PV 4989; m1 derecho, MPEF-PV 4741; m1 izquierdo, MPEF- PV 4931; m1 izquierdo, MPEF-PV 4940; m1 izquierdo, MPEF-PV 5284; m2 izquierdo, MPEF-PV 4739; m2 izquier- do, MPEF-PV 4934; m2 izquierdo, MPEF-PV 4943; m2 iz- quierdo, MPEF-PV 4961; m3 derecho, MPEF-PV 4932; m3 derecho, MPEF-PV 4938; m3 izquierdo, MPEF-PV 4963; m4 derecho, MPEF-PV 4930; m4 derecho, MPEF-PV 5281; fragmento de maxilar derecho con M2–M3 (Fig. 3.9–10), MPEF-PV 5661; fragmento maxilar derecho con M3–M4, MPEF-PV 6026; M1 derecho, MPEF-PV 4737; M1 derecho, MPEF-PV 4919; M1 derecho, MPEF-PV 4920; M1 derecho, MPEF-PV 4971; M1 izquierdo, MPEF-PV 4975; M2 iz- quierdo, MPEF-PV 4736; M2 derecho, MPEF-PV 5282; M2 izquierdo, MPEF-PV 5283; M3 derecho, MPEF-PV 4918; M3 izquierdo, MPEF-PV 4922." | ||||||||||
Microbiotherium hernandezi n. sp.
Simpson 1932
|
Marshall 1982 | 1 specimen | ||||||||
synonym of Oligobiotherium divisum | ||||||||||
AMNH 29664, dentary p2-m4, type | ||||||||||
Mammalia
- Sparassodonta
- Thylacosmilidae
|
||||||||||
Thylacosmilidae indet.
Riggs 1933
|
Goin et al. 2007 | 1 specimen | ||||||||
"MLP 92-X-10-6 un molar superior derecho (M?3)." | ||||||||||
Mammalia
- Sparassodonta
- Hathliacynidae
|
||||||||||
Hathliacynidae indet.
Ameghino 1894
|
Goin et al. 2007 | 4 specimens | ||||||||
corrected as Hathlyacininae | ||||||||||
"MPEF-PV 4771, un premolar inferior aislado." "MPEF-PV 4767, un talónido derecho aislado." "MPEF-PV 4766 (figura 6.K), un molar superior derecho (M?1)." "MPEF-PV 4770, una hemimandíbula casi completa con los i3-m4 (figura 6.L)." | ||||||||||
Mammalia
- Sparassodonta
|
||||||||||
Cladosictis sp.
Ameghino 1887
|
Goin et al. 2007 | 2 specimens | ||||||||
"MLP 82-V-2-109, un fragmento mandibular derecho con los m1-2 completos." "MPEF-PV 4768, un molar inferior derecho aislado (m?3)." | ||||||||||
Cladosictis centralis n. sp.
Ameghino 1902
|
Marshall et al. 1984 | 5 specimens | ||||||||
From ref. 58306 (Marshall, 1981): MACN 11639, partial skull, type. From ref. 57989 (Marshall et al. 1984): MNHN COL 4, mandible with right c-m4 and left p2-m4; MNHN COL 122, m3. From ref. 58052 (Goin et al. 2007): MPEF-PV 4772, M2 or M3 fragment; MPEF-PV 5854, M1 | ||||||||||
Sipalocyon sp.
Ameghino 1887
|
Goin et al. 2007 | 2 specimens | ||||||||
"MPEF-PV 4769 (figuras 6.E-F), un molar inferior derecho aislado (m4)." "MPEF-PV 4545 (figura 6.G), un M3 izquierdo aislado." | ||||||||||
Sipalocyon externus n. sp.
(Ameghino 1902)
|
Goin et al. 2007 | 5 specimens | ||||||||
Reported by ref. 58306 (Marshall, 1981): MACN 52-383, partial skull, holotype. Reported by ref. 58052: "MPEF-PV 5840, un fragmento mandibular iz- quierdo con los m1-2 completos (figura 6.A); MPEF-PV 5832, un fragmento de maxilar izquierdo con el P2, un M3 sin el protocono y el M4 completo (figuras 6.B-C), todos probablemente asociados; MLP 77-VI-13-7, un fragmento de M2 aislado; MLP 82-V-2-85, fragmento de maxilar derecho con el M1 completo (figura 6.D)." | ||||||||||
Anatherium herrerae n. sp.
Marshall 1981
|
Marshall 1981 | 1 specimen | ||||||||
synonym of Acyon tricuspidatus | ||||||||||
FMNH P13521, left and right dentaries, holotype | ||||||||||
Borhyaeninae indet.
Ameghino 1894
|
Goin et al. 2007 | 6 specimens | ||||||||
"MLP 85-VII-3-46, un fragmento de maxilar izquierdo con parte del alvéolo del canino, el P1 completo y las raíces de los P2-3; MLP 82-V-2-4b, un incisivo inferior aislado; MPEF-PV 5637a, un incisivo inferior aislado; MPEF-PV 5637c, un fragmento man- dibular derecho con los alvéolos del p2; MPEF-PV 5637b, un frag- mento de m2 derecho; MPEF-PV 5846b, un P3 izquierdo aislado." | ||||||||||
Arctodictis sinclairi
Marshall 1978
|
Goin et al. 2007 | 9 specimens | ||||||||
"MLP 85-VII-3-1, un cráneo y mandíbulas casi completos con buena parte del esqueleto postcraneano asociado; MLP 93-XI-18-9, un canino superior derecho aislado; MPEF-PV 5846c, un M1 izquierdo aislado, fragmentario (figura 7.H); MPEF- PV 4752, un fragmento mandibular izquierdo con los m3-4 com- pletos (figuras 7.I, J); MLP 82-V-2-4c, un canino superior izquierdo aislado; MLP 77-VI-13-27, un m2 izquierdo aislado; MLP 82-V-2- 4a, un m2 derecho aislado; MPEF-PV 5846d, un P2 derecho aislado; MLP 82-V-2-116, un P3 derecho aislado." | ||||||||||
cf. Arctodictis sinclairi
Marshall 1978
|
Goin et al. 2007 | 3 specimens | ||||||||
"MLP 77-VI-13-1, una hemimandíbula izquierda casi completa con los i1-m4 completos, una hemimandíbula dere- cha con los i1 e i3, y los c-m1 completos, y un canino superior ais- lado, probablemente asociado; MLP 84-III-15-1, fragmento de ra- ma mandibular derecha con los m2-4 completos y un M1 derecho aislado asociado; MPEF-PV 5846a, un p2 izquierdo aislado." | ||||||||||
Pseudoborhyaena macrodonta n. sp.
Ameghino 1902
|
Goin et al. 2007 | 4 specimens | ||||||||
recombined as Borhyaena macrodonta | ||||||||||
"MACN 52-390 (tipo de Pseudoborhyaena macrodonta Ameghino 1902), un resto craneano con los P2-M4 izquierdos, aso- ciado a un fragmento mandibular derecho con la raíz del p1, los p2-m3 completos y un fragmento del m4." "MLP 82-V-4-3, fragmento de maxi- lar derecho con los P2-3 completos (figuras 7.D-E); MLP 92-X-10-8, un m2 izquierdo aislado (figura 7.F); MPEF-PV 1467, fragmento de maxilar izquierdo con los P3-M3 (figura 7.G, detalle del M1)." | ||||||||||
Pseudoborhyaena longaeva n. sp.
Ameghino 1902
|
Goin et al. 2007 | 1 specimen | ||||||||
synonym of Borhyaena macrodonta | ||||||||||
"MACN 52-366 (tipo de Pseudoborhyaena longeva Ameghino, 1902), un fragmento mandibular derecho con los c-p1 completos, parte del p2, raíces del p3, m1 completo, raíces del m2 y los m3-4 parcialmente rotos" | ||||||||||
cf. Acrocyon riggsi
(Sinclair 1930)
|
Marshall et al. 1984 | 1 specimen | ||||||||
MNHN COL 125, P3 | ||||||||||
Pseudothylacynus rectus n. gen., n. sp.
Ameghino 1902
|
Goin et al. 2007 | 6 specimens | ||||||||
MNHN COL 5, dentary p3-m4, isolated m2, isolated m4; MPEF-PV 5838, incisor, p1, p?3, premolar; MLP 92-X-10-7, maxillary M1-2; MPEF-PV 4765, dentary m3-4; MLP 92-X-10-5, dentary p2-3. Described by ref. 58324 (Marshall, 1979): MACN 52-369, dentary p1-m4, holotype | ||||||||||
Mammalia
- Argyrolagidae
|
||||||||||
Proargyrolagus argentinus n. sp.
Goin and Abello 2013
|
Goin and Abello 2013 | 24 specimens | ||||||||
"Holotipo. M2 derecho, MPEF-PV 4711 (Fig. 4.9–10). Material referido. M1 derecho, MPEF-PV 4715; M1 dere- cho, MPEF-PV 4714 (Fig. 4.7–8); M1 derecho, MPEF-PV 4786; M2 derecho MPEF-PV 4781; M2 izquierdo MPEF- PV 4547; M3 izquierdo, MPEF-PV 4713 (Fig. 4.11–12); M3 derecho, MPEF 4783-PV; M?4 derecho, MPEF-PV 4782; Mx izquierdo, MPEF-PV 4784; Mx izquierdo, MPEF-PV 4785; m1 izquierdo, MPEF-PV 4712 (Fig. 4.1– 2); m2 izquierdo, MPEF-PV 4778 (Fig. 4.3–4); m2 izquier- do, MPEF-PV 4789; m2 derecho, MPEF-PV 4776; m3 izquierdo, MPEF-PV 4716; m3 derecho, MPEF-PV 4710 (Fig. 4.5–6); m3 derecho, MPEF-PV 4788; m3 derecho, MPEF-PV 4790; m4 derecho, MPEF-PV 4717; mx dere- cho, MPEF-PV 4780; mx izquierdo, MPEF-PV 4787; mx derecho, MPEF-PV 4775." | ||||||||||
Mammalia
- Meridiolestida
|
||||||||||
Necrolestes mirabilis n. sp.
Goin et al. 2007
|
Goin et al. 2007 | 6 specimens | ||||||||
MLP 92-X-10-14, mandibular fragment with p?4 and dp?4, holotype; MPEF-PV 4706, P?4; MPEF-PV 6030, m?1; MPEF-PV 4548, Mx; MPEF-PV 4549, upper premolariform; MPEF-PV 4546, upper premolariform | ||||||||||
see common names |
Geography
Country: | Argentina | State/province: | Chubut |
Coordinates: | 45.7° South, 68.7° West (view map) | ||
Paleocoordinates: | 46.4° South, 62.8° West | ||
Basis of coordinate: | estimated from map | ||
Altitude: | 257 meters | ||
Geographic resolution: | local area |
Time
Period: | Neogene | Epoch: | Miocene |
10 m.y. bin: | Cenozoic 5 | ||
Key time interval: | Colhuehuapian | Mammal zone: | Colhuehuapian |
Age range of interval: | 21.00000 - 17.50000 m.y. ago | ||
Age estimate: | 20.2 to 20 Ma (Ar/Ar) |
Stratigraphy
Formation: | Sarmiento | Member: | Colhué Huapí | ||
Stratigraphic resolution: | member | ||||
Stratigraphy comments: Busker, et al. 2019: Colhuehuapian SALMA (early Miocene) (Flynn and Swisher, 1995; Simpson, 1940; Vucetich et al., 2010b).
Novo et al 2018: The Colhue-Huapi Member of the Sarmiento Formation is, early Miocene (Burdigalian Age) (see Simpson 1964; Madden & Scarano 2010 and references therein), approximately 20.0-20.2 Ma (Ré et al. 2010). |
Lithology and environment
Primary lithology: | not reported |
Environment: | terrestrial indet. |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils,mesofossils |
Collection methods and comments
Collection methods: | field collection |
Reason for describing collection: | paleoecologic analysis |
Museum repositories: | MLP,MNHN |
Collectors: | Tournouër and others |
Collection method comments: Busker, et al. 2019: Only MPEF-PV 5421 comes from Gran Barranca locality, Chubut Province |
Metadata
Database number: | 141368 | ||
Authorizer: | C. Jaramillo, J. Zijlstra, J. Carrillo, P. Mannion | Enterer: | J. Carrillo, P. Mannion, K. Pino, M. Kouvari, G. Varnham, J. Zijlstra |
Modifier: | M. Kouvari | Research group: | vertebrate |
Collections that are a subset of this one: | |||
Created: | 2013-03-26 08:17:32 | Last modified: | 2020-11-30 05:46:45 |
Access level: | database members | Released: | 2014-03-26 08:17:32 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
45974. | M. G. Vucetich and A. G. Kramarz. 2003. New Miocene rodents from Patagonia (Argentina) and their bearing on the early radiation of the Octodontoids (Hystricognathi). Journal of Vertebrate Paleontology 23(2):435-444 [C. Jaramillo/J. Carrillo] |
Secondary references:
75307 | M. A. Abello, G. M. Martin, and Y. Cardoso. 2021. Review of the extinct ‘shrew-opossums’ (Marsupialia: Caenolestidae), with descriptions of two new genera and three new species from the Early Miocene of southern South America. Zoological Journal of the Linnean Society 1-35 [P. Mannion/M. Kouvari] | |
69887 | F. Busker, M. E. Pérez, and M. T. Dozo. 2019. A new chinchilloid (Rodentia, Hystricognathi) from the early Miocene of the localities of Bryn Gwyn and Gran Barranca (Patagonia, Argentina). Comptes Rendus Palevol 18(5):525-540 [P. Mannion/M. Kouvari/E. Vlachos] | |
58217 | A. M. Candela. 1999. The evolution of the molar pattern of the Erethizontidae (Rodentia, Hystricognathi) and the validity of Parasteiromys Ameghino, 1904. Palaeovertebrata 28(1):53-73 [J. Zijlstra/J. Zijlstra] | |
71055 | A. M. Candela. 2003. A new porcupine (Rodentia, Hystricognathi, Erethizontidae) from the Early and Middle Miocene of Patagonia. Ameghiniana 40(3):483-493 [P. Mannion/M. Kouvari] | |
26483 | M. A. Candela and M.G. Vucetich. 2002. Hypsosteiromys (Rodentia, Hystricognathi) from the Early Miocene of Patagonia (Argentina), the only Erethizontidae with a tendency to hypsodonty. Geobios 35:153-161 [L. van den Hoek Ostende/L. van den Hoek Ostende] | |
73071 | A. A. Carlini, M. Cancio, and G. J. Scillato-Yané. 2005. Los Xenarthra de Gran Barranca: Más de 20 Ma de historia. Actas del XVI Congreso Geológico Argentino. La Plata 2005 [P. Mannion/G. Varnham] | |
57869 | A. A. Carlini, M. R. Ciancio, and G. J. Scillato-Yane. 2010. Middle Eocene-Early Miocene Dasypodidae (Xenarthra) of southern South America: biostratigraphy and palaeoecology. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 106-129 [J. Zijlstra/J. Zijlstra/P. Mannion] | |
58245 | R. L. Cifelli and M. F. Soria. 1983. Systematics of the Adianthidae (Litopterna, Mammalia). American Museum Novitates 2771:1-25 [J. Zijlstra/J. Zijlstra/J. Zijlstra] | |
57940 | N. J. Czaplewski. 2010. Bats (Mammalia: Chiroptera) from Gran Barranca (early Miocene, Colhuehuapian), Chubut Province, Argentina. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 240-252 [J. Zijlstra/J. Zijlstra] | |
62548 | M. T. Dozo and B. Vera. 2010. First skull and associated postcranial bones of Macraucheniidae (Mammalia, Litopterna) from the Deseadan SALMA (late Oligocene) of Cabeza Blanca (Chubut, Argentina). Journal of Vertebrate Paleontology 30(6):1818-1826 [P. Mannion/P. Mannion] | |
58216 | M. T. Dozo, M. G. Vucetich, and A. M. Candela. 2004. Skull anatomy and neuromorphology of Hypsosteiromys, a Colhuehuapian erethizontid rodent from Argentina. Journal of Vertebrate Paleontology 24(1):228-234 [J. Zijlstra/J. Zijlstra] | |
70891 | J. C. Fernicola and S. F. Vizcaino. 2008. Revision del genero Stegotherium Ameghino, 1887 (Mammalia, Xenarthra, Dasypodidae). Ameghiniana 45(2):321-332 [P. Mannion/M. Kouvari] | |
58052 | F. Goin, A. Abello, E. Bellosi, R. Kay, R. Madden and A. Carlini. 2007. Los Metatheria sudamericanos de comienzos del Neógeno (Mioceno Temprano, Edad-mamífero Colhuehuapense). Parte I: Introducción, Didelphimorphia y Sparassodonta. Ameghiniana 44(1):29-71 [J. Zijlstra/J. Zijlstra/J. Zijlstra] | |
58272 | F. J. Goin and M. A. Abello. 2013. Los Metatheria sudamericanos de comienzos del Neógeno (Mioceno temprano, Edad Mamífero Colhuehuapense): Microbiotheria y Polydolopimorphia. Ameghiniana 50(1):51-78 [J. Zijlstra/J. Zijlstra/J. Zijlstra] | |
69517 | S. Hernández Del Pino, M. G. Fernández, E. Cerdeño and J. C. Fernicola. 2019. Anatomy and systematics of Notohippus toxodontoides Ameghino, 1891 (Mammalia, Notoungulata), from the Miocene of Santa Cruz Province, Argentina. Journal of Vertebrate Paleontology e1577870 [P. Mannion/G. Varnham] | |
73094 | P. Hershkovitz. 1981. Comparative Anatomy of Platyrrhine Mandibular Cheek Teeth dpm4, pm4, m1 with Particular Reference to Those of Homunculus (Cebidae), and Comments on Platyrrhine Origins. Folia Primatologica 35:179-217 [P. Mannion/G. Varnham] | |
57939 | R. F. Kay. 2010. A new primate from the early Miocene of Gran Barranca, Chubut Province, Argentina: paleoecological implications. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 220-239 [J. Zijlstra/J. Zijlstra] | |
57912 | A. G. Kramarz and M. Bond. 2010. Colhuehuapian Astrapotheriidae (Mammalia) from Gran Barranca south of Lake Colhue Huapi. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 182-192 [J. Zijlstra/J. Zijlstra] | |
57896 | G. M. López, A. M. Ribeiro, and M. Bond. 2010. The Notohippidae (Mammalia, Notoungulata) from Gran Barranca: preliminary considerations. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 143-151 [J. Zijlstra/J. Zijlstra/J. Zijlstra] | |
57760 | L. Marshall. 1982. Systematics of the South American marsupial family Microbiotheriidae. Fieldiana Geology 10:1-75 [J. Zijlstra/J. Zijlstra] | |
58306 | L. G. Marshall. 1981. Review of the Hathlyacyninae, an extinct subfamily of South American "dog-like" marsupials. Fieldiana Geology 7:1-120 [J. Zijlstra/J. Zijlstra] | |
57989 | L. G. Marshall, C. de Muizon, and R. Hoffstetter. 1984. Fossil Marsupialia in the Muséum national d'Histoire naturelle collected by André Tournouër from Patagonia, southern Argentina. Bulletin du Muséum national d'histoire naturelle 6(1):33-58 [J. Zijlstra/J. Zijlstra/J. Zijlstra] | |
72535 | N. M. Novo, M. F. Tejedor, and L. R. González-Ruiz. 2018. Previously unknown fossil platyrrhines (Primates) of Patagonia from the Tournouër collection at the Muséum national d'Histoire naturelle, Paris. Geodiversitas 40(22):529-535 [P. Mannion/M. Kouvari] | |
58215 | B. Patterson. 1958. A new genus of erethizontid rodents from the Colhuehuapian of Patagonia. Breviora 92:1-4 [J. Zijlstra/J. Zijlstra] | |
58075 | B. Patterson and R. Pascual. 1968. New echimyid rodents from the Oligocene of Patagonia, and a synopsis of the family. Breviora 301:1-14 [J. Zijlstra/J. Zijlstra] | |
58202 | F. J. Prevosti, A. M. Forasiepi, M. D. Ercoli and G. F. Turazzini. 2012. Paleoecology of the mammalian carnivores (Metatheria, Sparassodonta) of the Santa Cruz Formation (late Early Miocene). In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 173-193 [J. Zijlstra/J. Zijlstra/J. Zijlstra] | |
57906 | A. M. Ribeiro, G. M. López, and M. Bond. 2010. The Leontiniidae (Mammalia, Notoungulata) from the Sarmiento Formation at Gran Barranca, Chubut Province, Argentina. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 170-181 [J. Zijlstra/J. Zijlstra/J. Zijlstra] | |
72438 | F. D. Seoane and E. Cerdeño. 2019. Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae. Journal of Systematic Palaeontology 17(19):1635-1663 [P. Mannion/M. Kouvari] | |
85085 | G. G. Simpson. 1932. Cochilius volvens from the Colpodon beds of Patagonia. American Museum Novitates 577 [J. Carrillo/K. Pino] | |
70877 | M. F. Soria and R. Hoffstetter. 1985. Pternoconius tournoueri, nueva especie de Macraucheniidae (Mammalia; Litopterna) de edad Colhuehuapense (Oligoceno tardio) ; Pcia. del Chubut, Republica Argentina. Ameghiniana 22(3-4):149-158 [P. Mannion/M. Kouvari] | |
58073 | B. Vera, M. Fornasiero, and L. Del Favero. 2015. The Egidio Feruglio's collection in the Museum of Geology and Palaeontology of the University of Padova: its importance to the knowledge of Cenozoic mammals from South America. Museologia Scientifica 9:35-44 [J. Zijlstra/J. Zijlstra] | |
57933 | M. G. Vucetich, A. G. Kramarz, and A. M. Candela. 2010. Colhuehuapian rodents from Gran Barranca and other Patagonian localities: the state of the art. In R. H. Madden, A. A. Carlini, M. G. Vucetich, R. F. Kay (eds.), The Paleontology of Gran Barranca: Evolution and Environmental Change through the Middle Cenozoic of Patagonia 206-219 [J. Zijlstra/J. Zijlstra] |