Woodring Loc. 161a - Gatun Dam [middle Gatun Fm]: Tortonian, Panama
collected by Olsson 1918

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Bivalvia - Nuculanida - Nuculanidae
Saccella balboae (Brown and Pilsbry 1911)
Woodring 1973 R category
Bivalvia - Pectinida - Pectinidae
Lepidopecten scissuratus (Dall 1898)
Woodring 1982 R category
Bivalvia - Arcida - Arcidae
Anadara (Rasia) dariensis (Brown and Pilsbry 1911)
Woodring 1973 A category
SUBSPECIES: Anadara (Rasia) dariensis dariensis
Bivalvia - Cardiida - Cardiidae
Trachycardium (Dallocardia) dominicense (Gabb 1873)
Woodring 1982 R category
SUBSPECIES: Trachycardium (Dallocardia) dominicense dominicense
Bivalvia - Cardiida - Tellinidae
Tellina (Eurytellina) angulosa (Gmelin 1791)
Woodring 1982 R category
SUBSPECIES: Tellina (Eurytellina) angulosa costaricana
Bivalvia - Cardiida - Veneridae
Macrocallista (Paradione) maculata (Linnaeus 1758)
Woodring 1982 R category
recombined as Macrocallista maculata
Agiopoma (Pitarella) gatunensis (Dall 1903)
Woodring 1982 R category
recombined as Callocardia (Agriopoma) gatunensis
Bivalvia - Pholadida - Corbulidae
Caryocorbula oropendula (Olsson 1922)
Woodring 1982 F category
SUBSPECIES: Caryocorbula oropendula stena
Scaphopoda - Dentaliida - Dentaliidae
Dentalium (Dentalium) armillatum
Woodring 1973 R category
SUBSPECIES: Dentalium (Dentalium) armillatum armillatum
Gastropoda - Neogastropoda - Turbinellidae
Xancus validus (Sowerby 1850)
Woodring 1964 R category
recombined as Turbinella valida
Xancus validus falconensis in Woodring (1964); Xancus dodonaius praelavigatus
Gastropoda - Neogastropoda - Marginellidae
Volvarina leander (Brown and Pilsbry 1911)
Woodring 1970 R category
Gastropoda - Neogastropoda - Muricidae
Paziella (Panamurex) gatunensis (Brown and Pilsbry 1911)
Woodring 1959 R category
recombined as Poirieria (Panamurex) gatunensis
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria (Cancellaria) epistomifera (Guppy 1876)
Woodring 1970 C category
recombined as Bivetiella epistomifera
SUBSPECIES: Cancellaria (Cancellaria) epistomifera dariena
Gastropoda - Neogastropoda - Olividae
Oliva (Oliva) gatunensis (Toula 1908)
Woodring 1964 F category
original and current combination Oliva gatunensis
Gastropoda - Neogastropoda - Drilliidae
Clathrodrillia gatunensis (Toula 1909)
Woodring 1970 R category
Crassispira (Hindsiclava) consors (Sowerby 1850)
Woodring 1970 R category
recombined as Hindsiclava consors
SUBSPECIES: Crassispira (Hindsiclava) consors consors
Gastropoda - Neogastropoda - Conidae
Conus imitator Brown and Pilsbry 1911
Woodring 1970 F category
SUBSPECIES: Conus imitator imitator
Gastropoda - Neogastropoda - Terebridae
Strioterebrum spiriferum (Dall 1895)
Woodring 1970 C category
Gastropoda - Neogastropoda - Turridae
Pleurolira (Polystira) tenagos Gardner 1947
Woodring 1970 A category
    = Polystira sp. Woodring 1928
Todd and Johnson 2013
Gastropoda - Neogastropoda - Columbellidae
Strombina (Strombina) lessepsiana (Brown and Pilsbry 1911)
Woodring 1964 A category
Anachis (Costoanachis) mira (Dall 1896)
Woodring 1964 R category
SUBSPECIES: Anachis (Costoanachis) mira fugax
Gastropoda - Neogastropoda - Buccinidae
Metula gabbi Brown and Pilsbry 1911
Woodring 1964 R category
Hanetia dalli (Brown and Pilsbry 1911)
Woodring 1964 C category
SUBSPECIES: Hanetia dalli dalli
Antillophos (Antillophos) candei (d'Orbigny 1842)
Woodring 1964 A category
recombined as Antillophos candeanus
SUBSPECIES: Antillophos (Antillophos) candei gatunensis
Cymatophos veatchi (Olsson 1922)
Woodring 1964 R category
Gastropoda - Cassidae
Sconsia laevigata (Guppy 1850)
Woodring 1959 R category
SUBSPECIES Sconsia laevigata sublaevigata
Gastropoda - Tonnidae
Malea camura Guppy 1866
Woodring 1959 R category
Gastropoda - Naticidae
Natica (Naticarius) stenopa
original and current combination Naticarius
Natica (Natica) bolus Brown and Pilsbry 1913
Polinices canalizonalis
Gastropoda - Turritellidae
Turritella gatunensis Conrad 1857
SUBSPECIES: Turritella gatunensis gatunensis
see common names

Geography
Country:Panama State/province:Colon
Coordinates: 9.3° North, 79.9° West (view map)
Paleocoordinates:9.0° North, 78.6° West
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Tortonian 10 m.y. bin:Cenozoic 6
Key time interval:Tortonian
Age range of interval:11.62000 - 7.24600 m.y. ago
Age estimate:10.1 Ma (other)
Stratigraphy
Formation:Gatun Member:middle Gatun
Stratigraphic resolution:bed
Stratigraphy comments: LITHOSTRATIGRAPHY: From middle part of the Gatun Formation. The Gatun Formation is sibdivided into three faunal zones (proposed by Thomson and Keen, 1946): lower, middle, and upper. AGE: Late Miocene, on the basis of microfossil biostratigraphy (see Collins and Coates, 1999), refined by enterer to Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 10.1 Ma (Hendy, unpublished data).
Lithology and environment
Primary lithology: poorly lithified siltstone
Secondary lithology: poorly lithified silty sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: LITHOLOGY: Siltstone and silty sandstone.
Environment:marine indet.
Geology comments: No specific environmental data presented.
Taphonomy
Modes of preservation:body,original aragonite
Size of fossils:macrofossils
Collection methods and comments
Collection methods:surface (float),surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Museum repositories:USNM
Collectors:Olsson Collection dates:1918
Collection method comments: COLLECTOR: A.A. Olsson. REPOSITORY: USNM.
Taxonomic list comments:COVERAGE: Exhaustive for gastropods and bivalves. NOMENCLATURE: Authoritative publications, with modern nomenclature, and species-resolution identifications.
Metadata
Also known as:USGS 8395; WG161a
Database number:145335
Authorizer:A. Hendy Enterer:A. Hendy
Research group:marine invertebrate
Created:2013-06-01 00:16:57 Last modified:2013-11-25 13:37:23
Access level:the public Released:2013-06-01 00:16:57
Creative Commons license:CC BY
Reference information

Primary reference:

11306. W. P. Woodring. 1957. Geology and Paleontology of Canal Zone and adjoining parts of Panama. United States Geological Survey Professional Paper 306(A) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

48647 J. A. Todd and K. G. Johnson. 2013. Dissecting a marine snail species radiation (Conoidea: Turridae: Polystira) over 12 million years in the southwestern Caribbean. Bulletin of Marine Science 89(4) [A. Hendy/A. Hendy]
11372 W. P. Woodring. 1959. Geology and paleontology of Canal Zone and adjoining parts of Panama: Description of Tertiary mollusks (gastropods: Vermetidae to Thaididae). United States Geological Survey Professional Paper 306(B):147-239 [A. Miller/A. Hendy/A. Hendy]
13232 W. P. Woodring. 1964. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Columbellidae to Volutidae). United States Geological Survey Professional Paper 306(C):241-297 [A. Miller/A. Hendy/A. Hendy]
13236 W. P. Woodring. 1970. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). United States Geological Survey Professional Paper 306(D):299-452 [A. Miller/A. Hendy/A. Hendy]
13235 W. P. Woodring. 1973. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae). United States Geological Survey Professional Paper 306(E):453-539 [A. Miller/A. Hendy/A. Hendy]
13359 W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy]