Amöneburg: MN 2, Germany
collected by Thomas Keller et al.

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Squamata
Squamata indet. Oppel 1811
5 specimens
PAL EV 2013/272 (atlas fragemnt); PAL EV 2013/273 and 274 (fragments of humeri: two proximal and one distal part); PAL EV 2013/275 (1 left pelvis)
Reptilia
"Scincomorpha" indet. (Camp 1923)
1 element
PAL EV 2013/164 (a right dentary)
Gekkota indet. Cuvier 1817
1 element
PAL EV 2013/147 (left maxilla)
Merkurosaurus ornatus Klembara 2008
1 element
PAL EV 2013/268 (a jugal)
cf. Merkurosaurus ornatus Klembara 2008
1 element
PAL EV 2013/277 (a single osteoderm)
Platynota indet. Baur 1890
1 element
PAL EV 2013/270 (a single osteoderm)
Shinisauria indet. Conrad 2008
1 fragment
PAL EV 2013/269 (fragment of a parietal)
Reptilia - Anguidae
Anguinae indet. Gray 1825
92 elements
PAL EV 2013/176 (a right quadrate); PAL EV 2013/177 to 179 (3 dorsal vertebrae); PAL EV 2013/180 and 181 (2 caudal vertebrae); PAL EV 2013/182 to 267 (86 osteoderms)
Ophisaurus sp. Daudin 1803
5 elements
PAL EV 2013/172 to 174 (3 frontals: 1 left, 2 right); PAL EV 2013/175 (a right pterygoid); PAL EV 2013/276 (a right dentary)
Ophisaurus spinari Klembara 1979
1 element
PAL EV 2013/171 (a parietal)
Pseudopus sp. Merrem 1820
5 elements
PAL EV 2013/166 to 170 (5 osteoderms)
Pseudopus cf. ahnikoviensis Klembara 2012
1 element
PAL EV 2013/165 (a right dentary)
Reptilia
Blanus thomaskelleri n. sp. Cernansky et al. 2015
1 element
PAL EV 2013/271 (holotype: incomplete right dentary)
Reptilia - Lacertidae
Lacertidae indet. Bonaparte 1831
9 elements
PAL EV 2013/155 (fragment of a right frontal); PAL EV 2013/156 to 159 (4 jugals: 2 left, 2 right); PAL EV 2013/160 (a right pterygoid); PAL EV 2013/161 (a basisphenoid); PAL EV 2013/162 to 163 (2 dorsal vertebrae)
Lacerta poncenatensis Müller 1996
7 elements
PAL EV 2013/148 to 151 (4 dentary fragments: 2 of a left, 2 of a right dentary); PAL EV 2013/152 and 153 (2 maxilla fragments: 1 of a left, 1 of a right maxilla); PAL EV 2013/154 (a premaxilla)
Reptilia
Serpentes indet. (Linnaeus 1758)
36 elements
PAL EV 2013/111 (an incomplete compound bone); PAL EV 2013/112 to 146 (35 trunk and caudal vertebrae)
Reptilia - Tropidophiidae
? Falseryx sp. Szyndlar and Rage 2003
4 elements
PAL EV 2013/2 to 5 (4 trunk vertebrae)
Reptilia
Booidea indet. Gray 1825
2 elements
PAL EV 2013/6 and 7 (2 trunk vertebrae)
Colubroidea indet. Oppel 1811
1 element
PAL EV 2013/106 (a single trunk vertebra)
Reptilia - Crotalidae
Vipera sp. Laurenti 1768
3 elements
"aspis complex"; PAL EV 2013/107 to 109 (3 trunk vertebrae)
Reptilia - Colubridae
cf. Neonatrix sp. Holman 1973
2 elements
PAL EV 2013/11 and 12 (2 trunk vertebrae: 1 anterior, 1 posterior)
? Neonatrix sp. Holman 1973
1 element
PAL EV 2013/110 (a single cloacal vertebra)
Coluber informal sp. 1 Linnaeus 1758
72 elements
"Coluber sensu lato"; PAL EV 2013/13 to 84 (72 trunk vertebrae)
Coluber informal sp. 2 Linnaeus 1758
20 elements
"Coluber sensu lato"; PAL EV 2013/85 to 104 (20 trunk vertebrae)
Coluber indet. Linnaeus 1758
1 element
"Coluber sensu lato"; PAL EV 2013/105 (incomplete parabasisphenoid)
cf. Natrix sp. Laurenti 1768
3 elements
PAL EV 2013/8–10 (3 trunk vertebrae)
see common names

Geography
Country:Germany State/province:Rhineland-Palatinate County:Wiesbaden
Coordinates: 50.0° North, 8.3° East (view map)
Paleocoordinates:50.1° North, 6.2° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
10 m.y. bin:Cenozoic 5
Key time interval:MN 2
Age range of interval:22.40000 - 20.00000 m.y. ago
Stratigraphy
Formation:Wiesbaden
Stratigraphy comments: "The Formation [i.e. the Wiesbaden Formation, formerly referred to as the Lower Hydrobia beds; TL] can be divided into three parts. The lower (brackish) and middle (lacustrine) parts are middle Aquitanian, containing the MN 2 mammal zone (Tobien 1988; Gers & Medden 1991; Reichenbacher 2000; Reichenbacher & Keller 2002). Reichenbacher (2000) assumed that the MN 2b and MN 3a mammal sub-zones of the Late Aquitanian–Early Burdigalian interval corresponded to the upper part of the Wiesbaden Formation, named the ‘emerging facies’. The squamate microvertebrate material described here was collected by Thomas Keller and colleagues from the lower brackish part of the Wiesbaden Formation where numerous freshwater and terrestrial gastropods, including Gyraulus, Stagnicola and Planorbarius, also occur." Age of the lower part of the Wiesbaden Fm is given as MN2a.
Lithology and environment
Primary lithology: "limestone"
Includes fossils?Y
Lithology description: No lithology description is given in the primary reference. The Wiesbaden Formation, however, belongs to the so-called "Kalktertiär" ("limestone Tertiary") of the Mainz Basin. Hence, the host rocks may well have been calcareous rocks.
Environment:lagoonal/restricted shallow subtidal Tectonic setting:rift
Taphonomy
Modes of preservation:body
Size of fossils:mesofossils
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:sieve,survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:SMF
Collectors:Thomas Keller et al.
Collection method comments: "The fossils were recovered by screen washing and are housed in the SMF, prefixed by collection numbers of the PAL EV."
Metadata
Also known as:Amoneburg
Database number:167464
Authorizer:J. Mueller Enterer:T. Liebrecht
Modifier:T. Liebrecht Research group:vertebrate
Created:2015-03-18 06:11:05 Last modified:2015-03-18 11:11:11
Access level:the public Released:2015-03-18 06:11:05
Creative Commons license:CC BY
Reference information

Primary reference:
54643. A. Cernansky, J. C. Rage, and J. Klembara. 2015. The Early Miocene squamates of Amöneburg (Germany): the first stages of modern squamates in Europe. Journal of Systematic Palaeontology 13(2):97-128 [J. Mueller/T. Liebrecht]