Laguna Fría: Lutetian, Argentina

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
? Amilnedwardsiidae informal gen. et sp. indet. H
two left mandibular fragments from Laguna Fr ́ıa, one with p2 and p3 par- tially broken, and p4–m2 (LIEB-PV 1616), and a second specimen with p3–4 (LIEB-PV 1617)
Mammalia
Palangania cf. brandmayri Goin et al. 1998
lower molar (LIEB-PV 1051)
Mammalia - Microbiotheria
? Microbiotheria informal Gen. et sp. indet. F
Two lower molars
Mammalia - Microbiotheria - Microbiotheriidae
Microbiotheriidae informal Gen. et sp. indet. D
LIEB-PV 1039, a fragment of right dentary with the alveoli for i1-m2, and LIEB-PV 1041, a right Mx.
Eomicrobiotherium sp. Marshall 1982
LIEB-PV 1040, right m?3
Mammalia - Didelphimorphia
Didelphimorphia informal Gen. et sp. indet. A
Three upper molars including LIEB-PV 1042
Didelphimorphia informal Gen. et sp. nov. 3
Several upper and lower molars
Mammalia - Derorhynchidae
Derorhynchus sp. Couto 1952
two specimens (LIEB- PV 1102, right m4; LIEB-PV 1140, isolated left talonid)
Derorhynchus cf. minutus Goin et al. 1999
eight specimens
Pauladelphys sp. Goin et al. 1999
dentition, maxillae, and man- dibles mostly from Laguna Fría
Mammalia - Protodidelphidae
Protodidelphis informal cf. sp. nov.
About a dozen isolated upper and lower teeth
Mammalia - Peradectidae
Peradectidae informal Gen. et sp. nov. 1
One upper and three lower molars from Laguna Fr ́ıa are referred to this new taxon (LIEB-PV 1031, a left M?3).
Mammalia - Caroloameghiniidae
Caroloameghiniidae informal Gen. et sp. nov. 2
spelled with current rank as Caroloameghiniinae
Three lower molars collected at Laguna Fría
Mammalia - Sparassodonta - Hathliacynidae
Hathliacynidae informal Gen. et sp. nov. 6
corrected as Hathlyacininae
LIEB-PV 1036, a maxillary fragment with M?3 (fig. 3H), and LIEB-PV 1037, isolated metastylar portion of a right M?2,
Mammalia - Sparassodonta
cf. Nemolestes indet. Ameghino 1902
LIEB-PV 1038 is an isolated left m?4
    = Nemolestes lagunafriensis Rangel et al. 2023
Rangel et al. 2023
LIEB-PV 1038, a left m4, Holotype of Nemolestes lagunafriensis sp. nov. The LIEB-PV 1038 was originally considered as a possible m4 by Tejedor et al. (2009: p. 12), a conclusion followed in this study, based on the presence of a very short talonid and the unicuspid condition of the talonid, only hypoconulid. The size and morphology, developed anterolabial cingulid and very short talonid, of this specimen support its referral to a third species of Nemolestes. Nevertheless, the more reduced condition of the talonid and the enlargement of the hypoconulid of N. lagunafriensis support its identification as a different species from N. brasiliensis.
Mammalia - Polydolopidae
Polydolops unicus Tejedor et al. 2009
LIEB-PV 1240*, left maxillary fragment with M2–3; LIEB-PV 1241*, left M1, LIEB-PV 1242*, right M1, LIEB-PV 1243*, right M1, LIEB-PV 1244*, right, broken M1; LIEB-PV 1212*, left dentary with broken m1, complete m2, and roots of m3; LIEB-PV 1217*, right dentary with m1–3; LIEB-PV 1259*, left mandible with m1–2 and roots of p3; LIEB- PV 1260*, left mandible with m2–3; LIEB- PV 1261*, left m2; LIEB-PV 1262*, right mandible with m1–2;
Polydolops rothi Simpson 1936
recombined as Pliodolops rothi
several upper and lower specimens
Polydolops informal sp. nov. 1 Ameghino 1897
Not clear which material comes from Laguna Fría and which from La Barda. "LIEB-PV 1153, a maxillary fragment with M1–2, as well as four lower molars are included in this new species recovered in La Barda and Laguna Fría"
    = Hypodolops sapoensis n. gen., n. sp. Chornogubsky 2020
Chornogubsky 2020
The holotype: LIEB-PV 1153, LIEB-PV 1154, LIEB-PV 1158, LIEB-PV 1160 and LIEB-PV 1163 were exhumed from levels of the Tufolitas Laguna del H
Polydolops informal sp. nov. 2 Ameghino 1897
Not clear which material comes from Laguna Fría and which from La Barda. "LIEB-PV 1173, a right maxillary fragment with M1–2, as well as several maxillary and mandibular remains and isolated teeth have been assigned to this new taxon collected in both localities of Paso del Sapo."
    = Hypodolops sapoensis Chornogubsky 2020
Chornogubsky 2020
The holotype LIEB-PV 1153, LIEB-PV 1154, LIEB-PV 1158, LIEB-PV 1160 and LIEB-PV 1163 were exhumed from levels of the Tufolitas Laguna del H
Amphidolops informal sp. nov. 2 Ameghino 1902
Not clear which material comes from Laguna Fría and which from La Barda. "LIEB-PV 1187, a right m2, and several isolated lower and upper molars from both localities of Paso del Sapo"
Amphidolops informal sp. nov. 1 Ameghino 1902
LIEB-PV 1173, a right maxillary fragment with M1–2, as well as several maxillary and mandibular remains and isolated teeth have been assigned to this new taxon collected in both localities of Paso del Sapo.
    = Amphidolops minimus n. sp. Chornogubsky 2020
Chornogubsky 2020
LIEB-PV 1170, an isolated right M1 (Fig.
Amphidolops intermedius Chornogubsky 2020
Chornogubsky 2020
LIEB-PV 1179, LIEB-PV 1180, LIEB-PV 1189 and LIEB-PV 1190
Mammalia - Pucadelphyidae
cf. Itaboraidelphys sp. Marshall and de Muizon 1984
illustrated is an isolated right m2 or m3 (LIEB-PV 1064), and addi- tional referred materials include an isolated left metastylar area (M1 or M2; LIEB-PV 1066); isolated lower left premolar (LIEB-PV 1067); fragment of left upper molar (Mx; LIEB-PV 1110); and right metastylar fragment (LIEB-PV 1075).
Mammalia - Notoungulata - Isotemnidae
Isotemnus sp. Ameghino 1897
LIEB-PV 1646, right maxilla with P4–M3, LIEB-PV 1659, left m2
Mammalia - Notoungulata - Oldfieldthomasiidae
Oldfieldthomasiidae informal gen. et sp. indet. I
mandibular fragments
Mammalia - Notoungulata - Henricosborniidae
Henricosbornia lophodonta Ameghino 1901
Maxillary and mandibular fragments re- ferred to the Family Henricosborniidae are abundant in both localities of Paso del Sapo
Othnielmarshia lacunifera Ameghino 1901
right maxillary fragment with complete M1–3 (LIEB-PV 1618, fig. 7F)
Mammalia - Notoungulata - Archaeopithecidae
Archaeopithecus cf. rogeri Ameghino 1897
Not clear which material comes from Laguna Fría and which from La Barda. "Several maxillary and mandibular fragments (e.g., LIEB-PV 1625 to 1628), as well as isolated teeth (e.g., LIEB-PV 1629) referred to this family have been found in La Barda and Laguna Fría."
Teratopithecus elpidophoros n. gen., n. sp. López et al. 2020
López et al. 2020
LIEB-PV 1627, partial right maxilla with P3- M2
Mammalia - Notoungulata - Notostylopidae
Edvardotrouessartia sola Ameghino 1901
Two maxillary fragments were collected at Laguna Fr ́ıa, one with broken P2–4 (LIEB-PV 1619; fig. 7G), and the other with incomplete P4 (LIEB-PV 1620)
Notostylops sp. Ameghino 1897
Mandibular fragments and isolated upper teeth that can be referred to the genus Notostylops are abundant in both localities of Paso del Sapo.
Mammalia - Notoungulata
Notopithecinae informal gen. et sp. indet. J
spelled with current rank as Notopithecidae
A few isolated teeth, including LIEB-PV 1655, left lower p3
Mammalia - Astrapotheria
Astrapotheria informal gen. et sp. indet. K
left m1 or m2 (LIEB-PV 1623)
? Astrapotheria informal gen. et sp. indet. L
right maxillary fragment with part of the hypertrophied and compressed canine that may be attributable to the Astrapotheria (LIEB-PV 1624),
Mammalia - Chiroptera
Chiroptera informal gen. et sp. nov. 9
Tejedor et al. 2005
well preserved left m?2 (LIEB-PV 999; Fig. 2a, b, c) and a right talonid (LIEB-PV 1000; Fig. 2d)
Mammalia - Cingulata - Dasypodidae
Prostegotherium cf. astrifer Ameghino 1902
osteoderms
Astegotherium sp. Ameghino 1902
isolated plates
Stegosimpsonia sp. Vizcaino 1994
isolated plates of the dorsal carapace
Riostegotherium sp. Oliveira and Bergqvist 1998
Several isolated plates of the dorsal carapace
Mammalia - Panameriungulata - Didolodontidae
Didolodontidae informal gen. et sp. nov. 7
well-preserved m3 (LIEB-PV 1611)
Didolodontidae informal gen. et sp. nov. 8
right partial mandible with part of the talonid of m2 and complete m3 (LIEB-PV 1612)
Mammalia - Panameriungulata - Protolipternidae
Asmithwoodwardia subtrigona Ameghino 1901
several isolated teeth
Mammalia - Panameriungulata - Macraucheniidae
? Victorlemoineia longidens
incomplete right lower molariform (p4 or m1 LIEB-PV 1614)
Mammalia - Panameriungulata - Sparnotheriodontidae
Sparnotheriodontidae informal gen. et sp. indet. G
right maxillary fragment (MLP 66-V-12-2; fig. 7D) with three molariforms (P4–M2?), right M3 (MLP 66-V- 12-1), and a broken left M1 or M2 (LIEB-PV 1615).
see common names

Geography
Country:Argentina State/province:Chubut
Coordinates: 42.7° South, 69.9° West (view map)
Paleocoordinates:45.2° South, 60.1° West
Basis of coordinate:stated in text
Altitude:844 meters
Geographic resolution:outcrop
Time
Period:Paleogene Epoch:Eocene
Stage:Lutetian 10 m.y. bin:Cenozoic 2
Key time interval:Lutetian
Age range of interval:47.80000 - 41.20000 m.y. ago
Age estimate:49.26 ± 0.3 to 43.5 ± 1.14 Ma (Ar/Ar)
Stratigraphy
Stratigraphic resolution:bed
Stratigraphy comments: Proposed "Sapoan" South American Land Mammal Age (Tejedor et al. 2009)

Gosses et al. (2020) Conclusions: The Laguna Fría fauna is younger, constrained between the 49.26 ± 0.30 Ma Laguna Fría orange ignimbrite and the 43.50 ± 1.14 Ma Laguna Fría capping basalt. The age of the nearby La Barda fauna is more difficult to confidently determine. If previous stratigraphic relations inferred by Tejedor et al. (2009) are correct, then the La Barda fauna is at least 2 m.y. younger than its previously assumed age range of 47–45 Ma. The detailed stratigraphy of basaltic eruptions is poorly known however, and it is therefore possible that the La Barda and Laguna Fría faunas are similar in age. Finally, the Laguna Fría age range overlaps with the age ranges proposed by Krause et al. (2017) for the Riochican and Vacan faunas and therefore does not directly support the existence of a temporally distinct “Sapoan” SALMA.
Lithology and environment
Primary lithology: tuff
Includes fossils?Y
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:general faunal/floral analysis
Metadata
Database number:176059
Authorizer:J. Zijlstra, J. Carrillo, P. Mannion, E. Vlachos Enterer:J. Zijlstra, M. Kouvari, K. Pino, E. Vlachos
Modifier:K. Pino Research group:vertebrate
Subset of collection #:58765
Created:2016-01-23 22:05:33 Last modified:2023-07-13 05:53:49
Access level:the public Released:2016-01-23 22:05:33
Creative Commons license:CC BY
Reference information

Primary reference:
57758. M. F. Tejedor, F. J. Goin, J. N. Gelfo, G. López, M. Bond, A. A. Carlini, G. J. Scillato-Yané, M. O. Woodburne, L. Chornogubsky, E. Aragón, M. A. Reguero, N. J. Czaplewski, S. Vincon, G. M. Martin, and M. R. Ciancio. 2009. New Early Eocene mammalian fauna from western Patagonia, Argentina. American Museum Novitates 3638:1-43 [J. Zijlstra/J. Zijlstra]

Secondary references:

74919 L. Chornogubsky. 2020. Interrelationships of Polydolopidae (Mammalia: Marsupialia) from South America and Antarctica. Zoological Journal of the Linnean Society 1-42 [P. Mannion/M. Kouvari]
85265 J. Gosses, A. R. Carroll, B. T. Bruck, B. S. Singer, B. R. Jicha, E. Aragón, A. P. Walters and P. Wilf. 2020. Facies interpretation and geochronology of diverse Eocene floras and faunas, northwest Chubut Province, Patagonia, Argentina. Geological Society America Bulletin 133(3-4):740-752 [J. Carrillo/K. Pino]
72744 G. M. López, J. N. Gelfo, N. Bauzá, M. Bond, and M. F. Tejedor. 2020. Biochron and diversity of Archaeopithecidae (Mammalia, Notoungulata) and a new genus and species from the Eocene of Patagonia, Argentina. Ameghiniana 57(2):103-116 [E. Vlachos/E. Vlachos]
85086 C. C. Rangel, L. M. Carneiro, M. F. Tejedor, L. P. Bergqvist, and E. V. Oliveira. 2023. A reassessment of Nemolestes (Mammalia, Metatheria): Systematics and evolutionary implications for Sparassodonta. Journal of Mammalian Evolution [J. Carrillo/K. Pino/K. Pino]
16676 M. F. Tejedor, N. J. Czaplewski, F. J. Goin and E. Aragón. 2005. The oldest record of South American bats. Journal of Vertebrate Paleontology 25(4):990-993 [D. Croft/D. Croft/J. Alroy]