La Borie: Ypresian, France
collected 2006

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Polypodiopsida - Polypodiaceae
Polypodiaceae indet. Presl and Presl 1822
APSO.SP3-03, 04, 05, 06
Angiospermae - Rosales - Rhamnaceae
Rhamnaceae indet. Jussieu 1789
APSO.SP3-01, 02
Angiospermae - Fabales - Fabaceae
Fabaceae indet. Lindley 1836
APSO.SP3-07
Angiospermae - Sapindales - Anacardiaceae
Anacardiaceae indet. Brown 1818
APSO.SP3-08, 09, 10, 11, 12
unclassified
Charophyta indet. Migula 1890
Chondrichthyes - Carcharhiniformes - Carcharhinidae
Physogaleus tertius
Actinopteri - Amiiformes
Amiiformes indet. Hay 1929
Actinopteri - Lepisosteiformes
Lepisosteiformes indet. Hay 1929
Actinopteri - Siluriformes
Siluriformes indet.
APSO.SP2-181
Amphibia - Temnospondyli - Urodela
Salamandridae indet. Gray 1825
APSO.SP2-217
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
APSO.SP2-218, 219
Reptilia - Squamata
Squamata indet. Oppel 1811
APSO.SP2-229
Reptilia - Anguidae
Anguini indet. Augé 2005
APSO.SP2-224 and 225
Glyptosaurinae indet. Marsh 1872
APSO.SP2-223
Reptilia - Necrosauridae
Necrosauridae indet. Hoffstetter 1943
APSO.SP2-235 and 226-228
Reptilia - Boidae
Boidae informal Forme A
APSO.SP2-230 and 231
Boidae informal Forme B
APSO.SP2-232
Reptilia - Iguanidae
cf. Geiseltaliellus sp. Kuhn 1944
APSO.SP2-220 and 221
Reptilia - Eolacertidae
Eolacerta sp. Nöth 1940
APSO.SP2-222 and 234
Reptilia
Neornithes indet. Gadow 1893
Bourdon et al. 2014
Reptilia - Struthioniformes - Palaeotididae
Galligeranoides boriensis n. gen., n. sp. Bourdon et al. 2014
Bourdon et al. 2014
Reptilia - Gastornithiformes - Gastornithidae
Mourer-Chauviré and Bourdon 2020
Holotype: MHNT.PAL.2018.20.1 (mandible)
Gastornis parisiensis Hébert 1855
Bourdon et al. 2014
Reptilia
Crocodylus depressifrons Blainville 1855
recombined as Asiatosuchus depressifrons
APSO.SP1-150, APSO.SP1-133, 140, APSO. SP1-120, APSO.SP2-214, 215
Kentisuchus spenceri (Buckland 1836)
APSO.SP5-01, APSO.SP1-121; APSO.SP1- 74; APSO.SP5-05, 16
Reptilia - Testudines - Trionychidae
APSO.SP1-88, 123, 138; APSO.SP1-141; APSO.SP2-49, 82, 86, 99, 104, 137, 146, 160, 204; APSO.SP2-174, 176, 180, and APSO.SP5-22, 33
Reptilia - Testudines - Testudinidae
Achilemys cassouleti Claude and Tong 2004
recombined as Fontainechelon cassouleti
APSO.SP2-100
Palaeoemys sp. Schleich 1994
APSO.SP1-109, 112, 115, 132, 140, 146, 147, 148, 151; APSO.SP2-60, 203, and APSO.SP2-89
Reptilia - Testudines - Podocnemididae
Neochelys informal n. sp.
APSO.SP0-03, APSO.SP1-152, APSO.SP2-101, APSO.SP2-25, 134, and APSO.SP2-94, 95, 148
Mammalia - Primates - Notharctidae
Cercamoniinae informal n. gen.
APSO.SP2-233
Mammalia - Acreodi - Mesonychidae
Dissacus sp. Cope 1881
APSO.SP2-193 and APSO.SP5-02
    = Dissacus raslanloubatieri n. sp. Solé et al. 2018
Solé et al. 2018
MHNT.PAL.2015.6.1 - type
Mammalia - Creodonta - Hyaenodontidae
Proviverrinae informal n. gen.
APSO.SP5-26
    = Boritia duffaudi n. gen., n. sp. Solé et al. 2014
Solé et al. 2014
MHNT.PAL.2010.19.1 - holotype
Mammalia - Diacodexeidae
Diacodexis sp. Cope 1882
Boivin et al. 2018
UM BRI 1-4; MHNT.PAL.2017.21.2
Mammalia - Phenacodontidae
Phenacodus sp. Cope 1873
APSO.SP0-04
Mammalia - Perissodactyla - Palaeotheriidae
Pachynolophus sp. Pomel 1847
APSO.SP0-01, APSO.SP4-02 and APSO.SP0-06
Mammalia - Perissodactyla - Lophiodontidae
Lophiaspis sp. Depéret 1910
APSO.SP5-06, APSO.SP5- 08, APSO.SP5-15, 28, APSO.SP5-30, APSO.SP5-31 and APSO.SP6-01
    = Eolophiodon laboriense n. gen., n. sp. Robinet et al. 2015
Robinet et al. 2015
MHNT.PAL.2010.19.3 - holotype
Mammalia - Esthonychidae
Plesiesthonyx sp. Lemoine 1891
APSO.SP0.02
Gastropoda
Gastropoda indet. Cuvier 1795
Bivalvia - Unionida - Unionidae
Unionidae indet. Rafinesque 1820
Branchiopoda
Conchostraca indet. Sars 1867
invalid subgroup of Diplostraca
see common names

Geography
Country:France State/province:Aude
Coordinates: 43.3° North, 2.0° East (view map)
Paleocoordinates:38.8° North, 0.5° West
Basis of coordinate:estimated from map
Geographic resolution:small collection
Time
Period:Paleogene Epoch:Eocene
Stage:Ypresian 10 m.y. bin:Cenozoic 2
Key time interval:Ypresian Mammal zone: MP8 + 9
Age range of interval:56.00000 - 47.80000 m.y. ago
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: reference-level MP 8–9, early Eocene, middle Ypre- sian (Laurent et al., 2010; Danilo et al., 2013), with a numerical age of ca. 52 Ma (Escarguel et al., 1997).
Lithology and environment
Primary lithology: siltstone
Includes fossils?Y
Lithology description: organic rich clay deposits covered by layers of sand and conglomerates
Environment:fluvial-lacustrine indet.
Geology comments: All these deposits correspond to standing water bodies, suddenly invaded by sand and gravel during successive flood episodes. Analysis of the flora indicates a humid tropical climate, and a landscape of marshy wooded savanna in an alluvial plain (Laurent et al., 2010).
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Collection dates:2006
Metadata
Also known as:Saint-Papoul
Database number:182641
Authorizer:P. Mannion, M. Uhen Enterer:P. Mannion, G. Varnham, M. Uhen
Modifier:G. Varnham Research group:vertebrate
Created:2016-11-16 06:36:23 Last modified:2022-02-22 10:22:25
Access level:the public Released:2016-11-16 06:36:23
Creative Commons license:CC BY
Reference information

Primary reference:
60846. Y. Laurent, S. Adnet, E. Bourdon, D. Corbalan, L. Danilo, S. Duffaud, G. Fleury, G. Garcia, M. Godinot, G. Le Roux, C. Maisonnave, G. Métais, C. Mourer-Chauviré, B. Presseq, B. Sigé and F. Solé. 2010. La Borie (Saint-Papoul, Aude) : un gisement exceptionnel dans l’Éocène basal du Sud de la France. Bulletin de la Société d'Histoire Naturelle de Toulouse 146:89-103 [P. Mannion/P. Mannion]

Secondary references:

66245 M. Boivin, M. J. Orliac, M. Antunes Telles, M. Godinot, Y. Laurent, B. Marandat, D. Vidalenc and R. Tabuce. 2018. New material of Diacodexis (Mammalia, Artiodactyla) from the early Eocene of Southern Europe. Geobios 51(4):285-306 [P. Mannion/P. Mannion/P. Mannion]
74379 E. Bourdon, C. Mourer-Chauviré, and Y. Laurent. 2014. Early Eocene birds from La Borie, southern France. Acta Palaeontologica Polonica 61(1):175-190 [M. Uhen/M. Uhen]
80121 C. Mourer-Chauviré and E. Bourdon. 2020. Description of a new species of Gastornis (Aves, Gastornithiformes) from the early Eocene of La Borie, southwestern France. Geobios 63:39-46 [P. Mannion/G. Varnham/P. Mannion]
60849 C. Robinet, J. A. Remy, Y. Laurent, L. Danilo, and F. Lihoreau. 2015. A new genus of Lophiodontidae (Perissodactyla, Mammalia) from the early Eocene of La Borie (Southern France) and the origin of the genus Lophiodon Cuvier, 1822. Geobios 48:25-38 [P. Mannion/P. Mannion]
60836 F. Solé, J. Falconnet, and L. Yves. 2014. New proviverrines (Hyaenodontida) from the early Eocene of Europe; phylogeny and ecological evolution of the Proviverrinae. Zoological Journal of the Linnean Society 171:878-917 [P. Mannion/P. Mannion]
61876 F. Solé, M. Godinot, Y. Laurent, A. Galoyer, and T. Smith. 2018. The European Mesonychid Mammals: Phylogeny, Ecology, Biogeography, and Biochronology. Journal of Mammalian Evolution 25:339-379 [M. Uhen/M. Uhen/P. Mannion]