Vathylakkos-3 (=VAT): Late/Upper Miocene, Greece

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Testudines - Testudinidae
Testudo sp. Linnaeus 1758
They mention "of large size"; Vlachos et al. (2014) were unable to locate these specimens and postulate a possible mixture with material from Megalon Emvolon
    = Titanochelon sp. Pérez-García and Vlachos 2014
Vlachos 2021
plate I in Arambourg and Piveteau (1929) depicts a block of bones from Vathylakkos. The caption states that a Testudo is present among these bones, referring to a leg osteoderm and a pelvis of giant tortoise; this specimen forms part of the exhibition of the MNHN.
Mammalia - Proboscidea - Mammutidae
Mastodon pentelici
synonym of Mammut
Mammalia - Carnivora - Mustelidae
Plesiogulo crassa Teilhard de Chardin 1945
de Bonis et al. 1988
Mammalia - Carnivora - Hyaenidae
Ictitherium robustum
Ictitherium hipparionum
Plioviverrops orbignyi (Gaudry and Lartet 1856)
de Bonis et al. 1988
Hyaena eximia
Mammalia - Carnivora - Felidae
Machairodus aphanistus (Kaup 1833)
Felis cf. attica Wagner 1857
recombined as Pristifelis attica
Mammalia - Perissodactyla - Equidae
Hipparion dietrichi (Wehrli 1941)
de Bonis et al. 1988
Hipparion gracile Guerin and Mein 1971
synonym of Hipparion depereti
Hipparion macedonicum Koufos 1984
de Bonis et al. 1988
Mammalia - Perissodactyla - Chalicotheriidae
Chalicotherium cf. goldfussi Kaup 1833
    = Anisodon macedonicus de Bonis et al. 1995
Koufos 2012
Mammalia - Perissodactyla - Rhinocerotidae
Rhinocerus pachygnathus
original and current combination Rhinoceros
Aceratherium sp. Kaup 1834
Mammalia - Suidae
Sus erymanthius
Microstonyx major (Gervais 1848)
de Bonis et al. 1988
Mammalia - Tragulidae
Dorcatherium puyhauberti Arambourg and Piveteau 1929
Mammalia - Giraffidae
Giraffa attica
Orasius eximius
synonym of Giraffa
Helladotherium duvernoyi Gaudry and Lartet 1856
Samotherium boissieri Forsyth Major 1888
de Bonis et al. 1988
Mammalia - Bovidae
Tragoportax rugosifrons (Schlosser 1904)
de Bonis et al. 1988
Helicotragus rotundicornis
Gazella deperdita
Gazella sp. Blaineville 1816
de Bonis et al. 1988
Gazella gaudryi Schlosser 1904
Palaeoreas lindermayeri (Wagner 1848)
original and current combination Antilope lindermayeri
Tragocerus amaltheus
Protoryx carolinae Major 1891
Prostrepsiceros zitteli
de Bonis et al. 1988
Tragelaphus sp. de Blainville 1816
Nisidorcas planicornis (Pilgrim 1939)
de Bonis et al. 1988
Mammalia - Rodentia - Muridae
Parapodemus schaubi Papp 1947
de Bonis and Koufos 1999
synonym of Parapodemus gaudryi
see common names

Geography
Country:Greece State/province:Chalkidona
Coordinates: 40.8° North, 22.7° East (view map)
Paleocoordinates:40.5° North, 22.4° East
Time
Period:Neogene Epoch:Miocene
10 m.y. bin:Cenozoic 6
Key time interval:Late/Upper Miocene Mammal zone: MN12
Age range of interval:11.63000 - 5.33300 m.y. ago
Stratigraphy
Lithology and environment
Primary lithology: conglomeratic sandstone
Secondary lithology: claystone
Includes fossils?Y
Environment:fluvial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Museum repositories:MNHN
Metadata
Also known as:Ravin de Vatiluk; Ravin G
Database number:182750
Authorizer:E. Vlachos Enterer:E. Vlachos
Created:2016-11-23 08:19:21 Last modified:2016-11-23 08:19:21
Access level:the public Released:2016-11-23 08:19:21
Creative Commons license:CC BY
Reference information

Primary reference:
60908. C. Arambourg and J. Piveteau. 1929. Les vertébrés du Pontien de Salonique. Annales de Paleontologie 18:59-83 [E. Vlachos/E. Vlachos]

Secondary references:

69452 L. de Bonis, G. Bouvrain, and G. D. Koufos. 1988. Late Miocene mammal localities of the lower Axios valley (Macedonia, Greece) and their stratigraphic significance. Modern Geology 13:141-147 [E. Vlachos/E. Vlachos]
70463 L. de Bonis and G. D. Koufos. 1999. The Miocene large mammal succession in Greece. In J. Agusti, L. Rook, P. Andrews (eds.), Hominoid Evolution and Climatic Change in Europe, Volume 1, The Evolution of Neogene Terrestrial Ecosystems in Europe 1:205-223 [E. Vlachos/E. Vlachos]
69308 G. D. Koufos. 2012. New material of Chalicotheriidae (Perissodactyla, Mammalia) from the Late Miocene of Axios Valley, Macedonia (Greece) with the description of a new species. Annales de Paléontologie 98(3):203-224 [E. Vlachos/E. Vlachos]
78401 E. Vlachos. 2021. The Fossil Record of Turtles and Tortoises (Reptilia: Testudines) in Greece. In E. Vlachos (ed.), Fossil Vertebrates of Greece Vol. 1, 1:245-281 [E. Vlachos/E. Vlachos]