Level 3, Benguérir: Late/Upper Maastrichtian, Morocco

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Plesiosauria - Elasmosauridae
Elasmosauridae indet. Cope 1869
Reptilia - Mosasauridae
Eremiasaurus heterodontus LeBlanc et al. 2012
Carinodens minalmamar n. sp. Schulp et al. 2009
Schulp et al. 2009 1 specimen
OCP DEK/GE 453
Prognathodon sp. Dollo 1889
Prognathodon currii Christiansen and Bonde 2002
Bardet et al. 2005 2 specimens
OCP.DEK/GE (two teeth)
Globidens phosphaticus n. sp. Bardet et al. 2005
Bardet et al. 2005 24 specimens
OCP.DEK/GE 337 - type (mid-posterior tooth); paratypes: OCP.DEK/GE 338 (an anterior tooth) and OCP.DEK/GE 339-343 (all median teeth); referred material: OCP.DEK/GE unnumbered (21 teeth)
Halisaurus arambourgi Bardet et al. 2005
"Platecarpus" ptychodon Arambourg 1952
nomen dubium belonging to Plioplatecarpinae
Reptilia - Pachyvaranidae
Pachyvaranus crassispondylus Arambourg 1952
Houssaye et al. 2011 1 specimen
OCP.DEK/GE (vertebrae)
Reptilia - Testudines - Bothremydidae
Bothremydidae indet. Baur 1891
Reptilia - Testudines
Chelonioidea indet. Baur 1893
Actinopteri - Aulopiformes - Dercetidae
Stratodus apicalis Cope 1872
Actinopteri - Aulopiformes - Enchodontidae
Enchodus libycus
Enchodus bursauxi
Chondrichthyes - Myliobatiformes
Coupatezia elevata Noubhani and Cappetta 1997
Dasyrhombodus bondoni (Arambourg 1952)
Ixobatis mucronata (Arambourg 1952)
Chondrichthyes - Myliobatiformes - Rhombodontidae
Rhombodus microdon Arambourg 1952
Rhombodus meridionalis Arambourg 1952
Chondrichthyes - Rajiformes - Sclerorhynchidae
Ganopristis leptodon Arambourg 1935
Dalpiazia stomeri
Ctenopristis nougareti Arambourg 1940
Chondrichthyes - Rajiformes - Rhinobatidae
"Rhinobatos" informal sp. 1 Link 1790
Chondrichthyes - Orectolobiformes - Ginglymostomatidae
Ginglymostoma sp. Müller and Henle 1838
Plicatoscyllium gharbii Noubhani and Cappetta 1997
Chondrichthyes - Lamniformes - Otodontidae
Cretolamna maroccana
Chondrichthyes - Lamniformes - Serratolamnidae
Serratolamna aff. khderii
Serratolamna serrata (Agassiz 1838)
Chondrichthyes - Lamniformes - Anacoracidae
Squalicorax bassani
Squalicorax pristodontus (Agassiz 1835)
Squalicorax informal nov. sp. 1
    = Squalicorax microserratus Cappetta et al. 2014
Cappetta et al. 2014
Chondrichthyes - Lamniformes - Aquilolamnidae
Cretomanta sp. Case et al. 1990
Chondrichthyes
Youssoubatis aff. ganntourensis
Hypsobatis sp. Cappetta 1992
see common names

Geography
Country:Morocco
Coordinates: 32.3° North, 7.8° West (view map)
Paleocoordinates:24.4° North, 8.4° West
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
Key time interval:Late/Upper Maastrichtian
Age range of interval:72.10000 - 66.00000 m.y. ago
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: The use of dissimilarity indices and agglomerative method underscores two distinct associations: a lower one including the levels L6 and L5, and an upper one comprising the levels L4 to L2. These two associations allow to separate a lower and an upper Maastrichtian level. Moreover, the special morphology of the teeth of S. pristodontus occurring in L2, compared to what is observed in lower levels, permits us to distinguish a lower Upper Maastrichtian and an upper Upper Maastrichtian faunal assemblage.
Lithology and environment
Primary lithology: poorly lithified phosphorite
Environment:marine indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:surface (in situ),field collection
Reason for describing collection:general faunal/floral analysis
Museum repositories:MNHN
Collection method comments: Repository: University of Montpellier 2; Office Chérifien des Phosphates, Khouribga, Morocco (OCP)
Metadata
Database number:213665
Authorizer:M. Clapham Enterer:M. Clapham
Modifier:M. Clapham Research group:vertebrate
Created:2020-09-07 22:22:31 Last modified:2020-09-07 23:04:48
Access level:the public Released:2020-09-07 22:22:31
Creative Commons license:CC BY
Reference information

Primary reference:
73776. H. Cappetta, N. Bardet, X. Pereda Suberbiola, S. Adnet, D. Akkrim, M. Amalik, and A. Benabdallah. 2014. Marine vertebrate faunas from the Maastrichtian phosphates of Benguérir (Ganntour Basin, Morocco): Biostratigraphy, palaeobiogeography and palaeoecology. Palaeogeography, Palaeoclimatology, Palaeoecology 409:217-238 [M. Clapham/M. Clapham]

Secondary references:

37608 N. Bardet, X. Pereda Suberbiola, M. Iarochène, M. Amalik, and B. Bouya. 2005. Durophagous Mosasauridae (Squamata) from the Upper Cretaceous phosphates of Morocco, with description of a new species of Globidens. Netherlands Journal of Geosciences — Geologie en Mijnbouw 84(3):167-175 [P. Mannion/P. Mannion]
73775 H. Cappetta, S. Adnet, D. Akkrim and M. Amalik. 2014. New Squalicorax species (Neoselachii: Lamniformes) from the Lower Maastrichtian of Ganntour phosphate deposit, Morocco. Palaeovertebrata 38(3):e3 [M. Clapham/M. Clapham]
38134 A. Houssaye, N. Bardet, J.-C. Rage, X. Pereda-Suberbiola, B. Bouya, M. Amaghzaz, and M. Amalik. 2011. A review of Pachyvaranus crassispondylus Arambourg, 1952, a pachyostotic marine squamate from the latest Cretaceous phosphates of Morocco and Syria. Geological Magazine 148(2):237-249 [P. Mannion/P. Mannion]
46172 A. S. Schulp, N. Bardet, and B. Bouya. 2009. A new species of the durophagous mosasaur Carinodens (Squamata, Mosasauridae) and additional material of Carinodens belgicus from the Maastrichtian phosphates of Morocco. Netherlands Journal of Geosciences 88(3):161-167 [P. Mannion/J. Tennant]