Proetus concinnus-Warburgella stokesii Assemblage, Much Wenlock Limestone: Homerian, United Kingdom

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Semicoscinum sp.
Aulacopora sp.
Dokophyllym turbinatus
Lophostroma sp.
Chlorophyceae - Dasycladales - Diploporaceae
Diplopora sp. Schafhaeutl 1863
Anthozoa - Auloporida - Syringoporidae
Syringopora sp. Goldfuss 1826
Anthozoa - Favositida - Alveolitidae
Subalveolites sp. Sokolov 1955
Anthozoa - Favositida - Coenitidae
Coenites sp. Eichwald 1829
Planocoenites sp. Sokolov 1952
Anthozoa - Favositida - Theciidae
Thecia expatiata
Angopora sp. (Jones 1936)
original and current combination Thecia (Angopora)
Anthozoa - Favositida - Pachyporidae
Cladopora sp. Hall 1851
Thamnopora sp. Steininger 1831
Anthozoa - Favositida - Favositidae
Favosites multipora
Paleofavosites asper
Paleofavosites rugosus
Paleofavosites sp. Twenhofel 1914
Anthozoa - Heliolitida - Proporidae
Propora tubulata
Anthozoa - Heliolitida - Plasmoporidae
Plasmopora sp. Milne-Edwards and Haime 1849
Anthozoa - Heliolitida - Heliolitidae
Heliolites interstinctus
Heliolites megastoma
Heliolites sp. Dana 1846
Anthozoa - Heliolitida - Stelliporellidae
Stelliporella parvistella
Anthozoa - Heliolitida - Halysitidae
Cystihalysites sp. Chernyshev 1941
Halysites sp. von Waldheim 1813
Halysites catenularius (Linnaeus 1767)
Anthozoa - Stauriida - Kodonophyllidae
Kodonophyllum sp. Wedekind 1927
Anthozoa - Stauriida - Laccophyllidae
Syringaxon sp. Lindström 1882
Anthozoa - Stauriida - Lykophyllidae
Pycnactis sp. Ryder 1926
Plasmophyllum sp. Dybowski 1873
Phaulactis sp. Ryder 1926
Anthozoa - Stauriida - Calostylidae
Calostylis sp. Lindström 1868
Helminthidium sp. Lindström 1882
Anthozoa - Stauriida - Ptenophyllidae
Spongophylloides sp. Meyer 1881
Cymatelasma sp. Hill and Butler 1936
Anthozoa - Stauriida - Acervulariidae
Acervularia sp. Schweigger 1819
Anthozoa - Stauriida - Entelophyllidae
Entelophyllum articulatus
Anthozoa - Stauriida - Arachnophyllidae
Arachnophyllum murchisoni (Edwards and Haime 1851)
Anthozoa - Cystiphyllida - Fletcheriidae
Fletcheria sp. Milne-Edwards and Haime 1851
Anthozoa - Cystiphyllida - Goniophyllidae
Goniophyllum sp. Milne-Edwards and Haime 1850
Anthozoa - Cystiphyllida - Palaeocyclidae
Rhabdocyclus sp. Lang and Smith 1939
Anthozoa - Cystiphyllida - Tryplasmatidae
Tryplasma sp. Lonsdale 1845
Wenlockia sp. Kato 1966
Anthozoa - Cystiphyllida - Cystiphyllidae
Cystiphyllum siluriense
Microplasma sp. Dybowski 1873
Stromatoporoidea - Stromatoporida - Stromatoporidae
Stromatopora antiqua
Stromatopora carteri
Stromatopora sp. Goldfuss 1826
Stromatoporoidea - Stromatoporellida - Stromatoporellidae
Diplostroma sp. Bogoyavlenskaya 1965
synonym of Simplexodictyon
Stromatoporoidea - Actinostromatida - Actinostromatidae
Plectostroma sp. Nestor 1964
Stromatoporoidea - Actinostromatida - Densastromatidae
Densastroma sp. Flügel 1959
Stromatoporoidea - Labechiida - Labechiidae
Labechia conferta (Lonsdale 1839)
Stromatoporoidea - Clathrodictyida - Actinodictyidae
Ecclimadictyon sp. Nestor 1964
Stromatoporoidea - Clathrodictyida - Clathrodictyidae
Clathrodictyon striatellum
Clathrodictyon sp. Nicholson and Murie 1878
Clathrodictyon vesiculosum Nicholson and Murie 1878
unclassified
Echinodermata indet. Klein 1754
"Pelmatozoan elements"
Conodonta - Ozarkodinida
Ozarkodina sagitta (Walliser 1964)
Ozarkodina confluens (Rhodes 1953)
original and current combination Trichonodella symmetrica
Conodonta - Ozarkodinida - Spathognathodontodae
Ozarkodina excavata (Branson and Mehl 1933)
recombined as Hindeodella excavata
Conodonta - Panderodontida - Panderodontidae
Panderodus spp.
Cephalopoda - Orthocerida - Dawsonoceratidae
Dawsonoceras sp. Hyatt 1884
Gastropoda
Gastropoda indet. Cuvier 1795
Gastropoda - Murchisoniina - Hormotomidae
Hormotoma articulata (Murchison 1839)
Donald 1902
Gastropoda - Murchisoniina - Murchisoniidae
? Murchisonia dudleyensis n. sp. Donald 1902
Donald 1902
Gastropoda - Murchisoniina - Phanerotrematidae
Bembexia lloydii (de Sowerby and Murchison 1839)
Donald 1905
recombined as Ulrichospira lloydii
Gastropoda - Murchisoniina - Lophospiridae
Lophospira cyclonema (Salter 1873)
Donald 1906
recombined as Longstaffia cyclonema
Gastropoda - Murchisoniina - Loxonematidae
Loxonema planatum Longstaff 1909
Longstaff 1909
Loxonema (Rhabdostropha) pseudofasciatum n. sp. Donald 1905
Donald 1905
recombined as Loxonema pseudofasciatum
Gastropoda - Euomphalina - Platyceratidae
Capulus haliotis n. sp. de Sowerby and Murchison 1839
de Sowerby and Murchison 1839
    = Platyceras haliotis n. sp. de Sowerby and Murchison 1839
Wagner 2023
original and current combination Capulus haliotis
Gastropoda - Euomphalina - Euomphalidae
Poleumita walmstedti (Lindström 1884)
Wagner 2023
Poleumita discors (Sowerby 1814)
Poleumita rugosa (Sowerby 1814)
Wagner 2023
Euomphalopterus alatus (Wahlenberg 1821)
Gastropoda - Bellerophontida - Pterothecidae
Bucaniopsis expansus (de Sowerby and Murchison 1839)
Reed 1921
recombined as Phragmosphaera expansus
Gastropoda - Bellerophontida - Bucaniidae
Tremanotus dilatatus (de Sowerby and Murchison 1839)
Reed 1921
Gastropoda - Bellerophontida - Bellerophontidae
Prosoptychus aff. sphaera (Lindström 1884)
Wagner 2023
ornate species
Bellerophon wenlockensis Murchison 1839
Reed 1921
recombined as Prosoptychus wenlockensis
Cymbularia merita n. sp. Reed 1921
Reed 1921
Bucaniopsis latevittatus (Lindström 1884)
Reed 1921
recombined as Bucanopsis latevittatus
Gastropoda - Bellerophontida - Plectonotidae
Bellerophon (Bucaniella) trilobatus (Conrad 1839)
Reed 1921
recombined as Plectonotoides trilobatus
Gastropoda - Bellerophontida - Tropidodiscidae
Temnodiscus fletcheri n. sp. Reed 1921
Reed 1921
Rostroconchia - Conocardiida - Bransoniidae
Pleurorhynchus aequicostatus n. sp. Phillips 1848
Phillips 1848
recombined as Mulceodens aequicostatus
Bivalvia
Pelecypoda indet. Goldfuss 1820
synonym of Bivalvia
Bivalvia - Ostreida - Pterineidae
Pteronitella retroflexa
Bivalvia - Grammysiidae
Grammysia cingulata
Tentaculitida - Tentaculitidae
Tentaculites ornatus
Strophomenata - Strophomenida - Strophochonetidae
Protochonetes sp. Muir-Wood 1962
Strophomenata - Strophomenida - Amphistrophiidae
Amphistrophia sp. Hall and Clarke 1892
Rhynchonellata - Atrypida
Atrypida indet. Rzhonsnitskaya 1960
"ribbed atrypoid"
Rhynchonellata - Atrypida - Atrypidae
Atrypa sp. Dalman 1828
Rhynchonellata - Atrypida - Glassiidae
Glassia sp. Davidson 1881
genus name in quotes
Rhynchonellata - Orthida - Dalmanellidae
Resserella sp. Bancroft 1928
Isorthis sp. Kozlowski 1929
Rhynchonellata - Pentamerida - Clorindidae
Antirhynchonella sp. Oehlert 1887
Rhynchonellata - Pentamerida - Gypidulidae
Gypidula sp. Hall 1867
Rhynchonellata - Spiriferida - Delthyrididae
Howellella sp. Kozlowski 1946
Rhynchonellata - Spiriferida - Cyrtiidae
Eospirifer sp. Schuchert 1913
Striispirifer sp. Cooper and Muir-Wood 1951
Cyrtia sp. Dalman 1828
Rhynchonellata - Rhynchonellida - Hebetoechiidae
Sphaerirhynchia sp. Cooper and Muir-Wood 1951
Rhynchonellata - Rhynchonellida - Trigonirhynchiidae
Rhynchotreta sp. Hall 1879
Microsphaeridiorhynchus sp. Sartenaer 1970
Stenolaemata - Trepostomida - Amplexoporidae
Monotrypa sp. Nicholson 1879
? Amplexopora sp.
Monotrypella sp.
Stenolaemata - Trepostomida
Nicholsonella sp. Ulrich 1890
Stenolaemata - Cystoporata - Fistuliporidae
Fistulipora sp. McCoy 1850
Stenolaemata - Cystoporata - Ceramoporidae
Favositella sp. Etheridge and Foord 1884
Ceramopora sp. Hall 1851
Stenolaemata - Cyclostomata - Diastoporidae
Sagenella sp. Hall 1851
Stenolaemata - Cyclostomata - Corynotrypidae
Corynotrypa sp. Bassler 1911
Stenolaemata - Cyclostomata - Crownoporidae
Mitoclema sp. Ulrich 1882
Stenolaemata - Phylloporinidae
Pseudohornera sp. Roemer 1876
Stenolaemata - Cryptostomata - Fenestellidae
Archaeofenestella sp. Miller 1962
Fenestella sp. Lonsdale 1839
Reteporina sp. d'Orbigny 1849
Stenolaemata - Cryptostomata - Rhinidictyidae
Pachydictya sp. Ulrich 1882
Stenolaemata - Cryptostomata - Rhinoporidae
Lichenalia sp. Hall 1851
Stenolaemata - Cryptostomata - Arthrostylidae
Glauconomella sp. Bassler 1952
Stenolaemata - Rhabdomesida - Rhomboporidae
Rhombopora sp. Meek 1872
Stenolaemata - Trepostomata - Trematoporidae
Trematopora sp. Hall 1852
Stenolaemata - Trepostomata - Halloporidae
Hallopora sp. Bassler 1911
Stenolaemata - Trepostomata - Batostomellidae
Batostomella sp. Bassler 1911
Stenolaemata - Trepostomata - Heterotrypidae
Asperopora sp.
Stenolaemata - Trepostomata - Aisenvergiidae
Eridotrypa sp. Ulrich 1893
Ostracoda
Ostracoda indet. Latreille 1802
Ostracoda - Palaeocopida
Osmotoxotis phalacra
Sarmatotoxotis sp.
Aetholicotoxotis nidicola
Tribotoxotis dorsistriata
Zorotoxotis medioculta
Charitoxotis gravida
Amphitoxotis sp.
Strepula concentrica
Ostracoda - Palaeocopida - Beyrichiidae
Beyrichia clausa
Ostracoda - Palaeocopida
Tinotoxotis velivola
Gongylostonyx sp.
Ostracoda - Podocopida
Sleia grumulata
Sleia troglodytophila
Trilobita - Proetida - Proetidae
Warburgella stokesii (Murchison 1839)
Proetus concinnus (Dalman 1827)
recombined as Proetus (Proetus) concinnus
Trilobita - Corynexochida - Styginidae
Styginidae indet. Vogdes 1890
"illaenimorph styginids"
Trilobita - Phacopida - Dalmanitidae
Dalmanites sp. Barrande 1852
Trilobita - Phacopida - Phacopidae
Ananaspis sp. Campbell 1967
Trilobita - Phacopida - Acastidae
Acaste sp. Goldfuss 1843
Trilobita - Phacopida - Encrinuridae
Encrinurus sp. Emmrich 1844
Trilobita - Phacopida - Calymenidae
Calymene sp. Brongniart 1822
see common names

Geography
Country:United Kingdom State/province:England
Coordinates: 52.5° North, 2.1° West (view map)
Paleocoordinates:28.4° South, 12.9° West
Basis of coordinate:based on political unit
Geographic resolution:local area
Time
Period:Silurian Epoch:Wenlock
Stage:Homerian 10 m.y. bin:Silurian 2
Key time interval:Homerian
Age range of interval:430.50000 - 427.40000 m.y. ago
Stratigraphy
Formation:Much Wenlock Limestone
Stratigraphic resolution:formation
Lithology and environment
Primary lithology: grainstone
Secondary lithology: lime mudstone
Lithology description: "..several lithofacies...Trilobites most common in interbedded lst and silty mudst. and nodular lst. lithofacies. The lst. beds and nodules variously consist of carbonate mudstone, grainstone, packstone, and wackestone. "
Environment:marine indet.
Geology comments: "Level-bottom carbonate shelf...This is a BA 3 unit....several lithofacies...Trilobites most common in interbedded lst and silty mudst. and nodular lst. lithofacies. The lst. beds and nodules variously consist of carbonate mudstone, grainstone, packstone, and wackestone. Microfacies analysis indicates that the nodular lst. was depos. below storm wave base, although algae indicate photic zone..Deposition of interbedded lst and silty mudst in somewhat shallower water, up to depths approaching normal wave base. In view of wide extent of local Wenlock shelf and consequent attenuation of wave activity, the actual depths would not have been great. A well-oxygenated, mid-shelf location for the assemblage was envisaged by Ratcliffe (1988)."
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Reason for describing collection:paleoecologic analysis
Collection method comments: The Assemblage type is broadly distributed, but it seems fairly clear from the text that the elements listed here are known from the Much Wenlock Limestone.
Metadata
Database number:23057
Authorizer:M. Foote, P. Wagner Enterer:M. Foote, P. Wagner
Modifier:M. Uhen Research group:marine invertebrate
Created:2002-07-08 15:07:55 Last modified:2018-11-26 18:00:06
Access level:the public Released:2002-07-08 15:07:55
Creative Commons license:CC BY
Reference information

Primary reference:
6632. A. T. Thomas and P. D. Lane. 1999. Trilobite assemblages of the North Atlantic Region. in A. J. Boucot and J. D. Lawson, eds. Paleocommunities--a case study from the Silurian and Lower Devonian. Cambridge University Press, Cambrdige 444-457. [M. Foote/M. Foote/M. Foote]

Secondary references:

16207 J. C. de Sowerby and R. I. Murchison. 1839. The Silurian System. Part 2. Organic Remains. The Silurian System 2:579-768 [P. Wagner/P. Wagner/P. Wagner]
15868 J. G. Donald. 1902. On some of the Proterozoic Gasteropoda which have been referred to Murchisonia and Pleurotomaria, with descriptions of new subgenera and species. Quarterly Journal of the Geological Society of London 58(4):313-339 [P. Wagner/P. Wagner]
15869 J. G. Donald. 1905. Observations on some of the Loxonematidae, with descriptions of two species. Quarterly Journal of the Geological Society of London 61(5):564-566 [P. Wagner/P. Wagner]
15870 J. G. Donald. 1905. On some Gastropoda from the Silurian rocks of Llangadock (Caemarthenshire). Quarterly Journal of the Geological Society of London 61(5):567-578 [P. Wagner/P. Wagner]
11637 J. G. Donald. 1906. Notes on the genera Omospira, Lophospira and Turritoma, with descriptions of new Proterozoic species. Quarterly Journal of the Geological Society of London 62:552-572 [P. Wagner/P. Wagner/P. Wagner]
16154 J. D. Longstaff. 1909. On the genus Loxonema, with descriptions of new Proterozoic species. Quarterly Journal of the Geological Society of London 65(2):210-228 [P. Wagner/P. Wagner]
50497 J. Phillips. 1848. The Malvern Hills, compared with the Palaeozoic districts of Abberley, Woolhope, May Hill, Tortworth, and Usk. Memoir of the Geological Survey of the United Kingdom 2(1):1-330 [P. Wagner/P. Wagner]
11640 F. R. C. Reed. 1921. A monograph of the British Ordovician and Silurian Bellerophontacea. The Palaeontographical Society 31:1-92 [P. Wagner/P. Wagner/P. Wagner]
9042 P. J. Wagner. 2023. Paleozoic Gastropod, Rostroconch, Helcionelloid and Tergomyan Database (2006 - 2023). [P. Wagner/P. Wagner/P. Wagner]