Nălaţ-Vad: Late/Upper Maastrichtian, Romania
collected by V. Codrea 2001–2002, 2007–2013

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Multituberculata - Kogaionidae
Multituberculata indet. (Cope 1884)
Smith et al. 2002 10 specimens
    = Kogaionon sp. Râdulescu and Samson 1996
Csiki et al. 2010
    = Kogaionidae indet. Râdulescu and Samson 1996
Csiki-Sava et al. 2016
Kogaionon radulescui n. sp. Smith et al. 2022
Smith et al. 2022
Barbatodon sp. Radulesco and Samson 1987
Smith et al. 2002 2 specimens
"n. sp."
Reptilia
Kallokibotion sp. Nopcsa 1923
Csiki et al. 2010 3 specimens
UBB NVK-7 (partial anterior region of a carapace of a juvenile specimen); UBB NVK-31 (almost complete shell, lacking the most posterior regions of both the carapace and the plastron); LPB [FGGUB] R.2086
    = Kallokibotion bajazidi Nopcsa 1923
Pérez-García and Codrea 2017
Scincomorpha indet. Camp 1923
Smith et al. 2002
possibly Scincidae
Theropoda indet. (Marsh 1881)
Smith et al. 2002 1 specimen
Reptilia - Alvarezsauridae
Elopteryx nopcsai Andrews 1913
Csiki-Sava et al. 2016
LPB [FGGUB] R.2399
Reptilia
Euenantiornithes indet. Chiappe 2002
Wang et al. 2011 2 specimens
NVEN 1 (complete right humerus), NVEN 2 (proximal end of left humerus)
Reptilia - Gargantuaviidae
Gargantuaviidae indet. Buffetaut and Angst 2019
Mayr et al. 2020
UBB V649 (incomplete pelvis)
Reptilia - Troodontidae
cf. Troodontidae indet. Gilmore 1924
Smith et al. 2002
    = Troodontidae indet. Gilmore 1924
Csiki et al. 2010
Reptilia - Dromaeosauridae
Velociraptorinae indet. Barsbold 1983
Smith et al. 2002
Reptilia
cf. Paronychodon sp. Cope 1876
Smith et al. 2002 1 specimen
Reptilia - Nodosauridae
Nodosauridae indet. Marsh 1890
Smith et al. 2002 1 specimen
tooth
Struthiosaurus sp. Bunzel 1870
Csiki et al. 2010
Reptilia
Iguanodontia indet. (Baur 1891)
Smith et al. 2002 1 individual
similar to Mochlodon robustus
cf. Telmatosaurus sp. Nopcsa 1903
Dalla Vecchia 2014 1 specimen
"? cf. Telmatosaurus" (LPB [FGGUB] R.2005, 2406)
Reptilia - Rhabdodontidae
? Rhabdodontidae indet. Weishampel et al. 2003
Brusatte et al. 2017 1 specimen
LPB (FGGUB) R.2408
Zalmoxes sp. Weishampel et al. 2003
Brusatte et al. 2017 4 specimens
LPB (FGGUB) R.2409, LPB (FGGUB) R.2167, LPB (FGGUB) R.2331, LPB (FGGUB) R.2407
? Zalmoxes sp. Weishampel et al. 2003
Brusatte et al. 2017 1 specimen
LPB (FGGUB) R.2167
Zalmoxes shqiperorum Weishampel et al. 2003
Csiki-Sava et al. 2016 6 specimens
LPB (FGGUB) R.2349, LPB (FGGUB) R.2592, LPB (FGGUB) R.2411, LPB (FGGUB) R.2403, LPB (FGGUB) R.2164, LPB (FGGUB) R.2410
Reptilia
cf. Paludititan sp. Csiki et al. 2010
Csiki-Sava et al. 2016 1 specimen
"? cf. Paludititan" (LPB [FGGUB] R.2344)
Titanosauridae indet. Lydekker 1885
Smith et al. 2002 1 individual
UBB NVM1
    = Magyarosaurus sp. Huene 1932
Codrea et al. 2010
    = Paludititan nalatzensis n. gen., n. sp. Csiki et al. 2010
Csiki et al. 2010
Reptilia - Titanosauridae
Megaloolithus cf. siruguei Vianey-Liaud et al. 1994
Smith et al. 2002 10 specimens
Reptilia - Azhdarchidae
? Hatzegopteryx sp. Buffetaut et al. 2002
Csiki-Sava et al. 2016 1 specimen
LPB [FGGUB] R.2396
Reptilia - Crocodyliformes
Crocodyliformes indet. (Hay 1930)
Csiki-Sava et al. 2016
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpetontidae indet. Fox and Naylor 1982
Smith et al. 2002 2 specimens
see common names

Geography
Country:Romania State/province:Hunedoara
Coordinates: 45.6° North, 22.9° East (view map)
Paleocoordinates:32.7° North, 21.1° East
Basis of coordinate:based on nearby landmark
Altitude:230 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
*Period:Late/Upper Cretaceous *Epoch:Senonian
Key time interval:Late/Upper Maastrichtian
Age range of interval:72.10000 - 66.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Râul Mare Member:upper
Stratigraphic resolution:group of beds
Stratigraphy comments: lower to middle part of the upper Maastricthtian (Csiki-Sava et al. 2016)
"Râul Mare beds", possibly upper section of ? Sînpetru Fm. or the Densus-Ciula Fm.
"can be considered as slightly older than at Totesti-baraj"; at least 7 horizons with eggs at the site.
also listed as early Maastrichtian Sînpetru Formation
Lithology and environment
Primary lithology:massive,paleosol/pedogenic,black silty mudstone
Secondary lithology:shelly/skeletal "limestone"
Includes fossils?Y
Includes fossils?Y
Lithology description: black marl and coquina limestone with microvertebrate remains; "dark gray silty mudstones" and "massive, dark gray mudstones"
Environment:wet floodplain
Geology comments: "fluvial palaeoenvironment with sandy channel infills and reddish brown overbank fines"; "Preserved in poorly drained floodplain deposits"
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Articulated whole bodies:some
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:occasional
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:bulk,surface (in situ),sieve,field collection
Rock censused:600 kg
Reason for describing collection:general faunal/floral analysis
Collectors:V. Codrea Collection dates:2001–2002, 2007–2013
Metadata
Also known as:Nalat-Vad; Nalatvad; Râul Mare River
Database number:26369
Authorizer:M. Carrano, M. Uhen, E. Vlachos, P. Mannion Enterer:M. Carrano, P. Mannion, S. Brown, E. Vlachos
Modifier:M. Carrano Research group:vertebrate
Created:2002-10-14 20:23:10 Last modified:2022-03-08 15:12:10
Access level:the public Released:2002-10-14 20:23:10
Creative Commons license:CC BY
Reference information

Primary reference:
37235. V. Codrea, M. Vremir, C. Jipa, P. Godefroit, Z. Csiki, T. Smith, and C. Farcas. 2010. More than just Nopcsa's Transylvanian dinosaurs: A look outside the Hateg Basin. Palaeogeography, Palaeoclimatology, Palaeoecology 293:391-405 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

76966 S. L. Brusatte, M. Dumbravă, M. Vremir, Z. Csiki-Sava, R. Totoianu and M. A. Norell. 2017. A catalog of Zalmoxes (Dinosauria: Ornithopoda) specimens from the Upper Cretaceous Nălaţ-Vad locality, Haţeg Basin, Romania. American Museum Novitates 3884:1-36 [M. Carrano/M. Carrano]
37255 Z. Csiki, V. Codrea, C. Jipa-Murzea and P. Godefroit. 2010. A partial titanosaur (Sauropoda, Dinosauria) skeleton from the Maastrichtian of Nalat-Vad, Hateg Basin, Romania. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 258(3):297-324 [M. Carrano/M. Carrano]
77788 Z. Csiki, D. Grigorescu, V. Codrea and F. Therrien. 2010. Taphonomic modes in the Maastrichtian continental deposits of the Haţeg Basin, Romania—palaeoecological and palaeobiological inferences. Palaeogeography, Palaeoclimatology, Palaeoecology 293(3-4):375-390 [M. Carrano/M. Carrano]
76092 Z. Csiki-Sava, M. Vremir, S. Vasile, S. L. Brusatte, G. Dyke, D. Naish, M. A. Norell and R. Totoianu. 2016. The East Side Story – The Transylvanian latest Cretaceous continental vertebrate record and its implications for understanding Cretaceous–Paleogene boundary events. Cretaceous Research 57:662-698 [P. Mannion/P. Mannion]
56054 F. M. Dalla Vecchia. 2014. An overview of the latest Cretaceous hadrosauroid record in Europe. In D. A. Eberth & D. C. Evans (ed.), Hadrosaurs 268-297 [M. Carrano/M. Carrano]
76794 V. Díez Díaz and X. Pereda Suberbiola. 2010. Titanosaurian fossil record of Europe: a review. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 30 [M. Carrano/M. Carrano/M. Carrano]
36651ETE D. Grigorescu. 2004. Vertebrate paleontology in Romania. Noesis 29:97-116 [M. Carrano/M. Carrano]
78206 D. Grigorescu. 2005. Rediscovery of a “forgotten land”: the last three decades of research on the dinosaur-bearing deposits from the Haţeg Basin. Acta Palaeontologica Romaniae 5:191-204 [M. Carrano/M. Carrano]
78208 D. Grigorescu and Z. Csiki. 2008. A new site with megaloolithid egg remains in the Maastrichtian of the Haţeg Basin. Acta Palaeontologica Romaniae 6:115-121 [M. Carrano/M. Carrano]
44509 D. Grigorescu, G. Garcia, Z. Csiki, V. Codrea, and A.-V. Bojar. 2010. Uppermost Cretaceous megaloolithid eggs from the Haţeg Basin, Romania, associated with hadrosaur hatchlings: search for explanation. Palaeogeography, Palaeoclimatology, Palaeoecology 293:360-374 [M. Carrano/M. Carrano]
80599 E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
76674 G. Mayr, V. Codrea, A. Solomon, M. Bordeianu, and T. Smith. 2020. A well-preserved pelvis from the Maastrichtian of Romania suggests that the enigmatic Gargantuavis is neither an ornithurine bird nor an insular endemic. Cretaceous Research 106:104271:1-10 [P. Mannion/P. Mannion/P. Mannion]
63553 A. Pérez-García and V. Codrea. 2017. New insights on the anatomy and systematics of Kallokibotion Nopcsa, 1923, the enigmatic uppermost Cretaceous basal turtle (stem Testudines) from Transylvania. Zoological Journal of the Linnean Society 182(2):419-443 [E. Vlachos/E. Vlachos/P. Mannion]
37256 T. Smith, V. A. Codrea, A. Sasaran, J. Van Itterbeeck, P. Bultynck, Z. Csiki, P. Dica, C. Farcas, A. Folie, G. Garcia, and P. Godefroit. 2002. A new exceptional vertebrate site from the Late Cretaceous of the Hateg Basin (Romania). Studia Universitatis Babes-Bolyai, Geologia, Special Issue 1:321-330 [M. Carrano/M. Carrano]
78786 T. Smith, V. A. Codrea, G. Devillet and A. A. Solomon. 2022. A new mammal skull from the Late Cretaceous of Romania and phylogenetic affinities of kogaionid multituberculates. Journal of Mammalian Evolution 29:1-26 [M. Uhen/S. Brown/P. Mannion]
39698 X. Wang, G. J. Dyke, V. Codrea, P. Godefroit, and T. Smith. 2011. A euenantiornithine bird from the Late Cretaceous Haţeg Basin of Romania. Acta Palaeontologica Polonica 56(4):853-857 [P. Mannion/P. Mannion]