Donguz I, PIN 952: Anisian, Russian Federation
collected by P. Klimov 1935–1936

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Chondrichthyes
Elasmobranchii indet. Bonaparte 1838
1 element
"Ichthyodorulit" is a spine, presumably from Elasmobranchii
Amphibia - Temnospondyli
Labyrinthodontia indet. Owen 1859
7 specimens
    = Eryosuchus tverdochlebovi n. gen., n. sp. Ochev 1966
Shishkin et al. 2000
PIN (SGU) 104/3090, skull and postcranial material (holotype). Additional postcranial material includes PIN (SGU) 104:194-196, 2965, 3684, 3693
Eryosuchus garjainovi Ochev 1966
Schoch and Milner 2000 5 specimens
synonym of Eryosuchus tverdochlebovi
PIN 104/1203, 1278, 2279, 2966, 2950, postcranial bones
Amphibia - Temnospondyli - Plagiosauridae
Plagioscutum ochevi Shishkin 1986
Tverdokhlebov et al. 2003
Bystrowianidae
Bystrowianidae indet. Vjushkov 1957
Tverdokhlebov et al. 2003
PIN 952/184, fragmentary scute
Reptilia - Erythrosuchidae
Erythrosuchus magnus Ochev 1980
Gower and Sennikov 2000 1 specimen
recombined as Uralosaurus magnus
PIN 952/95, four presacral vertebrae
Reptilia
Dorosuchus neoetus Sennikov 1989
Gower and Sennikov 2000 1 specimen
PIN 952/200, ilium
Dongusuchus efremovi n. gen., n. sp. Sennikov 1988
Gower and Sennikov 2000 2 specimens
PIN 952/15-1, femur (holotype). PIN 952/84-1, epipodials
Dongusia colorata n. gen., n. sp. Huene 1940
Huene 1940 1 specimen
nomen dubium belonging to Pseudosuchia
PIN 268/2, dorsal vertebra
Reptilia - Thecodontia
? Thecodontosaurus sp. Riley and Stutchbury 1836
1 specimen
    = Thecodontia indet. Owen 1859
Nessov 1995
Anomodontia - Dicynodontidae
Rabidosaurus cristatus n. gen., n. sp. Kalandadze 1970
Battail and Surkov 2000 1 specimen
PIN 952/100, almost complete skull
Anomodontia - Kannemeyeriidae
Lystrosaurus klimovi n. sp. Efremov 1940
4 specimens
recombined as Rhadiodromus klimovi
PIN 159/1, partial skull and incomplete skeleton (holotype). PIN 952/111, incomplete skull. PIN 952/13, 18, isolated bones
Therocephalia - Bauriidae
Dongusaurus schepetovi n. gen., n. sp. Vjuschkov 1964
Battail and Surkov 2000 1 specimen
nomen dubium belonging to Bauriidae
PIN 952/1, right dentary lacking teeth
Dipnoi - Ceratodontidae
Ceratodus gracilis
Tverdokhlebov et al. 2003
Dental plates
Ceratodus jechartiensis Minikh 1977
Tverdokhlebov et al. 2003
Dental plates
Ceratodus recticristatus Vorobyeva and Minikh 1968
Tverdokhlebov et al. 2003
Dental plates
see common names

Geography
Country:Russian Federation State/province:Orenburg County:Sol'-Iletsk
Coordinates: 51.4° North, 55.3° East (view map)
Paleocoordinates:33.6° North, 43.4° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Middle Triassic
Stage:Anisian 10 m.y. bin:Triassic 2
*Period:Early/Lower Triassic - Middle Triassic *Epoch:Scythian - Middle Triassic
Key time interval:Anisian Other zone: Perovkan
Age range of interval:247.20000 - 242.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Donguz
Stratigraphic resolution:group of beds
Stratigraphy comments: Outcrop is ~10 meters high. The author does not explicitly state which layer Lystrosuarus klimovi is from in the english text, but bed 3 is marked in Fig. 24. Detailed lithological descriptions are provided for each bed in the russian text.
Equivalent to the South African "Cynognathus Zone, corresponding to the Upper Buntsandstein of Germany, or to the Scythian." The Cynognathus Zone has recently been split into three time equivalent assemblages, therefore a Scythian age may not be correct.

Additional note: Tverdokhlebov et al. (2003) assign the locality to the Middle Triassic (Anisian) Donguz Svita. Original stratigraphic data are modified accordingly (TL).

The Donguz Svita is the "characteristic" tetrapod assemblage for the Perovkan Land Vertebrate Faunachron of Lucas, considered to be of Anisian age (see Lucas 2010)
Lithology and environment
Primary lithology:ferruginous,black,gray,green sandstone
Secondary lithology:lenticular conglomerate
Includes fossils?Y
Lithology description: There are at least three vertebrate-bearing layers at this outcrop, it is unclear whether all of the bones are found in concretions. General outcrop description includes variegated marls, clays, sands and sandstones. Lystrosaurus-bearing concretions = "rather mellow sandstone, which here and there is impregnated with ferrous and manganous compounds, dying the rock into an almost black colour." Layer 8 = clay sandstones of greenish tint. Layer 9 = grey, coarse-granulate sandstones and conglomerates with Ural pebbles. Tverdokhlebov et al. (2003) give the following description: "Grey, fine- to medium-grained sandstone with lenses of conglomerate and interlayers of siltstone."
Environment:fluvial-deltaic indet. Tectonic setting:foreland basin
Geology comments: "deltaic deposits" Tverdokhlebov et al. (2003).
Taphonomy
Modes of preservation:body,concretion
Degree of concentration:dispersed
Size of fossils:macrofossils
Articulated whole bodies:some
Disassociated minor elements:many
Collection methods and comments
Collection methods:surface (float),surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Collectors:P. Klimov Collection dates:1935–1936
Taxonomic list comments:Full taxa list as given in Tverdokhlebov et al. (2003) is not yet entered into the collection since indeterminate taxa already listed by Efremov may have been re-identified later. See references in Tverdokhlebov et al. (2003) for details.
Metadata
Also known as:Locality 36; Dongus river; Perovsky settlement;
Database number:28850
Authorizer:J. Alroy, A. Dunhill, R. Butler, M. Carrano Enterer:R. Whatley, M. Carrano, R. Butler, B. Allen
Modifier:M. Carrano Research group:vertebrate
Created:2003-02-25 13:00:03 Last modified:2021-10-06 12:57:36
Access level:the public Released:2003-02-25 13:00:03
Creative Commons license:CC BY
Reference information

Primary reference:

7854.5% 47000J. A. Efremov. 1940. Predvaritel'noye opisaniye novykh form permskoy i triasovoy fauny nazemnykh pozvonochnykh SSSR [Preliminary description of new forms from the Permian and Triassic terrestrial vertebrate fauna of the USSR]. Trudy Paleontologicheskogo Instituta Akademiy Nauk SSSR 10(2):1-140 [J. Alroy/R. Whatley/M. Carrano]

Secondary references:

28966 B. Battail and M. V. Surkov. 2000. Mammal-like reptiles from Russia. In M. J. Benton, M. A. Shishkin, D. M. Unwin & E. N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge 86-119 [J. Mueller/T. Liebrecht]
29101 D. J. Gower and A. G. Sennikov. 2000. Early Archosaurs from Russia. In M. J. Benton, M. A. Shishkin, D. M. Unwin & E. N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge 140-159 [J. Mueller/T. Liebrecht]
9920 F. v. Huene. 1940. Eine Reptilfauna aus der ältesten Trias Nordrußlands [A reptilian fauna from the earliest Triassic of northern Russia]. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, Beilagen-Band, Abteilung B 84:1-23 [J. Alroy/R. Whatley/T. Liebrecht]
46409 M. F. Ivakhnenko. 2011. Permian and Triassic therocephals (Eutherapsida) of Eastern Europe. Paleontological Journal 45(9):981-1144 [R. Butler/R. Butler]
16510ETE L. A. Nessov. 1995. Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii [Dinosaurs of northern Eurasia: new data about assemblages, ecology, and paleobiogeography]. Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg 1-156 [M. Carrano/K. Maguire/M. Carrano]
28828 R. Schoch and A. R. Milner. 2000. Stereospondyli. Handbuch der Paläoherpetologie - Encyclopedia of Paleoherpetology 3B:1-203 [J. Mueller/T. Liebrecht]
28553 M. A. Shishkin, I. V. Novikov, and Y. M. Gubin. 2000. Permian and Triassic temnospondyls from Russia. In M. J. Benton, M. A. Shishkin, D. M. Unwin & E. N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge 35-59 [J. Mueller/T. Liebrecht/T. Liebrecht]
28844 V. P. Tverdokhlebov, G. I. Tverdokhlebova, M. V. Surkov and M. J. Benton. 2003. Tetrapod localities from the Triassic of the SE of European Russia. Earth-Science Reviews 60(1-2):1-66 [J. Mueller/T. Liebrecht/B. Allen]
68327 K. B. Yuryev. 1954. Kratkiy obzor nakhodok dinozavrov na territorii SSSR [A brief reivew of dinosaur finds in the USSR]. Uchenyye Zapiski, Leningradskogo Ordena Lenina Gosudarstvennogo Universiteta. Seriya Biologicheskikh Nauk 181(38):183-197 [M. Carrano/M. Carrano]