Bangtoupo F32, Qingyan, China: Pelsonian - Illyrian, China

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Cephalopoda - Ceratitida - Ptychitidae
? Ptychites indet. Mojsisovics 1875
1 individual
Ptychites? sp. indet.
Cephalopoda - Ceratitida - Ceratitidae
Paraceratites brembanus (Mojsisovics 1882)
1 individual
Cephalopoda - Pseudorthocerida - Trematoceratidae
Michelinoceras cf. nyalamense Chen 1981
1 individual
recombined as Pseudotemperoceras nyalamense
Scaphopoda - Dentaliida - Laevidentaliidae
Laevidentalium bangtoupoensis
7 individuals
original and current combination Dentalium (Laevidentalium)
Scaphopoda - Dentaliida - Prodentaliidae
Prodentalium qingyanicum
2 individuals
Gastropoda
Gastropoda indet. Cuvier 1795
3 individuals
Gastropoda gen. et sp. indet.
Gastropoda - Heterostropha - Gordenellidae
Promathilda sp. n. sp. (Andreae 1887)
1 individual
original and current combination Promathildia
Promathilda n. sp. C
Gastropoda - Naticopsidae
Naticopsis (Naticopsis) sp. n. sp. M'Coy 1844
1 individual
Naticopsis n. sp. A
Naticopsis (Naticopsis) sp. n. sp. M'Coy 1844
2 individuals
Naticopsis n. sp. B
Naticopsis (Naticopsis) signata (Koken 1900)
1 individual
Hologyra guizhoui Yin and Yochelson 1983
10 individuals
Gastropoda - Neritariidae
Neritaria sp. n. sp. Koken 1892
4 individuals
Neritaria n. sp. A
Neritaria sp. n. sp. Koken 1892
11 individuals
Neritaria n. sp. A
Gastropoda - Goniasmatidae
Cheilotomona pentagonum Yu et al. 1974
1 individual
Gastropoda - Coelostylinidae
Coelostylinidae indet. n. gen., n. sp. Cossmann 1909
1 individual
Coelostylinidae n. gen. A, n. sp. A
Coelostylina ahlburgi (Assmann 1924)
3 individuals
Coelostylina liscaviensis (Giebel 1856)
6 individuals
Trypanostylus cf. turritelliformis
2 individuals
Trypanostylus sp. n. sp. Cossmann 1895
1 individual
Trypanostylus n. sp. A
Trypanostylus sp. n. sp. Cossmann 1895
2 individuals
Trypanostylus n. sp. B
Trypanostylus konincki (Münster 1841)
6 individuals
Toxoconcha brocchii (Stoppani 1857)
3 individuals
recombined as Toxoconcha brocchi
Gastropoda - Euomphalina
Triassocirrus moellendorfii
1 individual
Triassocirrus moellendorfi (Koken 1900)
1 individual
Gastropoda - Turbinidae
Turbinidae indet. n. gen., n. sp. Rafinesque 1815
5 individuals
Turbinidae n. gen. A, n. sp. A
Gastropoda - Trochidae
Trochidae indet. n. gen., n. sp. Rafinesque 1815
11 individuals
Trochidae n. gen. A, n. sp. A
Gastropoda - Eucyclidae
Riselloidea guizhouensis Yu et al. 1974
1 individual
Eucycloscala sp. n. sp. Cossmann 1895
2 individuals
Eucycloscala n. sp. A
Eucycloscala sp. n. sp. Cossmann 1895
10 individuals
Eucycloscala n. sp. B
Eucycloscala indet. Cossmann 1895
1 individual
Eucycloscala sp. indet.
Eucycloscala obliquicostata Yin and Yochelson 1983
3 individuals
Eucycloscala binodosa (Münster 1841)
1 individual
Gastropoda - Schizogoniidae
Schizogonium pentagonum Yu et al. 1974
2 individuals
Gastropoda - Discohelicidae
Discohelicidae indet. n. gen., n. sp. Schröder 1995
7 individuals
Discohelicidae n. gen. A, n. sp. A
Discogonius subvenustus
1 individual
original and current combination Discohelix (Discogonius)
Gastropoda - Cycloneritida - Delphinulopsidae
Delphinulopsis binodosa (Münster 1841)
5 individuals
Delphinulopsis cf. laubei Kittl 1892
2 individuals
synonym of Delphinulopsis binodosa
Gastropoda - Cycloneritida - Tricolnaticopsidae
Pachyomphalus sp. n. sp. Böhm 1895
26 individuals
Pachyomphalus n. sp. A
Gastropoda - Murchisoniina - Murchisoniidae
Murchisoniidae indet. n. gen., n. sp. Koken 1896
2 individuals
Murchisoniidae n. gen. B, n. sp. A
Gastropoda - Murchisoniina - Wortheniellidae
Wortheniella conica
7 individuals
Wortheniella nuda
3 individuals
Wortheniella tuberculifera
1 individual
Wortheniella xui
1 individual
Wortheniella sp. n. sp. Schwardt 1992
4 individuals
Wortheniella n. sp. D
Gastropoda - Murchisoniina - Rhaphistomellidae
Sisenna sp. Koken 1896
2 individuals
Sisenna sp. A
Gastropoda - Murchisoniina - Zygopleuridae
Zygopleura arctecostata (Münster 1841)
5 individuals
Zygopleura walmstedti (Klipstein 1843)
2 individuals
Gastropoda - Murchisoniina - Loxonematidae
Tyrsoecus multinodosa Yin and Yochelson 1983
10 individuals
Gastropoda - Murchisoniina - Spirostylidae
Spirostylus subcolumnaris (Münster 1841)
1 individual
Bivalvia
Bivalvia indet. Linnaeus 1758
10 individuals
Bivalvia gen. et sp. indet.
Bivalvia - Nuculanida - Malletiidae
Phaenodesmia sp. n. sp. Bittner 1895
2 individuals
Phaenodesmia n. sp. A
Bivalvia - Nuculanida - Polidevciidae
Phestia (Polidevcia) subperlonga
1 individual
spelled with current rank as Polidevcia
Bivalvia - Nuculida - Nuculidae
Palaeonucula qingyanensis Chen 1976
10 individuals
Bivalvia - Mytilida - Mytilidae
Modiolus (Modiolus) salzstettensis (Hohenstein 1913)
3 individuals
Bivalvia - Ostreida
Ostreoidea indet. n. gen., n. sp. (Rafinesque 1815)
15 individuals
original and current combination Ostreacea
Ostreoidea n. gen. A, n. sp. A
Bivalvia - Ostreida - Ostreidae
Lopha sp. n. sp. Röding 1798
1 individual
Lopha n. sp. A
Bivalvia - Ostreida - Halobiidae
Enteropleura guembeli (Mojsisovics 1874)
2 individuals
Bivalvia - Ostreida - Cassianellidae
Cassianella sulcata
12 individuals
Cassianella ecki sulcata
Cassianella qingyanensis
23 individuals
Cassianella subcislonensis
43 individuals
Cassianella simplex Chen 1976
48 individuals
Bivalvia - Ostreida - Bakevelliidae
Gervillia (Cultriopsis) angulata Münster 1841
2 individuals
Bivalvia - Ostreida - Pteriidae
Arcavicula (Arcavicula) guizhouensis
16 individuals
Arcavicula indet. n. sp. Cox 1964
2 individuals
Arcavicula (n. subgen. A) n. sp. A
Pteria caudata
3 individuals
Pteria rugosa Chen 1976
2 individuals
Pteria cassiana (Bittner 1895)
1 individual
Bivalvia - Pectinida - Limidae
Mysidioptera (Mysidioptera) punctata Chen 1976
1 individual
Palaeolima dunkeri
1 individual
Palaeolima subcostata
2 individuals
Palaeolima sp. n. sp. Hind 1903
2 individuals
Palaeolima n. sp. A
Bivalvia - Pectinida - Hunanopectinidae
Leptochondria plana
2 individuals
Leptochondria albertii (Goldfuss 1838)
1 individual
Bivalvia - Pectinida - Plicatulidae
Pseudoplacunopsis sp. n. sp. Bittner 1895
2 individuals
Pseudoplacunopsis n. sp. A
Bivalvia - Pectinida - Anomiidae
Placunopsis subplana
8 individuals
Bivalvia - Pectinida - Dimyidae
Protostrea sinensis (Hsu 1943)
7 individuals
Bivalvia - Pectinida - Entoliidae
Entolium discites (Schlotheim 1820)
2 individuals
Bivalvia - Pectinida - Terquemiidae
Terquemiidae indet. n. gen., n. sp. Cox 1964
6 individuals
Terquemiidae n. gen. B, n. sp. A
Newaagia multicostata
1 individual
Newaagia sessilis
4 individuals
Bivalvia - Pectinida - Pectinidae
Praechlamys schroeteri
1 individual
Praechlamys stenodictya
1 individual
Bivalvia - Cyrtodontida - Pichleriidae
Elegantarca subareata Chen 1976
9 individuals
Bivalvia - Arcida - Parallelodontidae
Parallelodon beyrichi
4 individuals
Bivalvia - Arcida - Philobryidae
Philobryidae indet. n. gen., n. sp. Bernard 1897
56 individuals
Philobryidae n. gen. A, n. sp. A
    = Eophilobryoidella sinoanisica Stiller and Chen 2004
Stiller and Chen 2004
Bivalvia - Trigoniida - Myophoriidae
Elegantinia kulunensis
5 individuals
Elegantinia sp. n. sp. (Waagen 1907)
13 individuals
Elegantinia n. sp. A
Elegantinia elegans (Dunker 1849)
13 individuals
Costatoria multiformis
5 individuals
Costatoria proharpa multiformis
Neoschizodus laevigatus (Ziethen 1830)
2 individuals
Quadratia quadrata Yin 1974
13 individuals
recombined as Guizhoumyophoria quadrata
Bivalvia - Lucinida - Lucinidae
Schafhaeutlia astartiformis (Münster 1841)
18 individuals
recombined as Fimbria astartiformis
Bivalvia - Cardiida - Kalenteridae
Pseudomyoconcha maximilianleuchtenbergensis
2 individuals
Bivalvia - Cardiida - Anthracosiidae
Unionites sp. n. sp. Wissman 1841
2 individuals
Unionites n. sp. A
Unionites sp. n. sp. Wissman 1841
1 individual
Unionites n. sp. B
Rhynchonellata - Spiriferinida - Bittnerulidae
Thecocyrtelloidea tubulosa Yang and Xu 1966
11 individuals
Rhynchonellata - Spiriferinida - Cyrtinidae
"Cyrtina" indet. Davidson 1858
1 individual
Rhynchonellata - Spiriferinida - Lepismatinidae
Lepismatina hsui Wang 1955
2 individuals
Rhynchonellata - Spiriferinida - Balatonospiridae
Nudispiriferina minima Yang and Xu 1966
4 individuals
Rhynchonellata - Spiriferinida - Mentzeliidae
Hirsutella multicostata (Yang and Xu 1966)
5 individuals
recombined as Dagyssia multicostata
Mentzelia subspherica Yang and Xu 1966
12 individuals
Mentzelia mentzeli (Dunker 1851)
1 individual
Rhynchonellata - Rhynchonellida - Cyclothyrididae
Costirhynchopsis sinensis (Yang and Xu 1966)
1 individual
Rhynchonellata - Rhynchonellida - Allorhynchidae
Septaliphorioidea paucicostata Yang and Xu 1966
6 individuals
Rhynchonellata - Rhynchonellida - Norellidae
Diholkorhynchia sinensis (Koken 1910)
24 individuals
Rhynchonellata - Terebratulida - Angustothyrididae
Angustothyris angustaeformis (Boekh 1872)
13 individuals
Polychaeta - Sabellida - Serpulidae
Ruxingella lianjiangensis
21 individuals
Weixiserpula weixi
43 individuals
Echinoidea - Triadocidaridae
"Triadocidaris" sp. Döderlein 1887
1 individual
"Triadocidaris" sp. A
Echinoidea - Cidaroida - Miocidaridae
"Miocidaris" sp. Döderlein 1887
1 individual
"Miocidaris" sp. B
Crinoidea - Qingyanocrinidae
Qingyanocrinus kueichounensis (Dubatolova and Shao 1959)
1 individual
Crinoidea - Millericrinida - Bangtoupocrinidae
Bangtoupocrinus kokeni Stiller 2000
1 individual
Silesiacrinus parvus Stiller 2000
1 individual
Crinoidea - Encrinida - Encrinidae
Encrinus cf. liliiformis Lamarck 1801
4 individuals
Crinoidea - Isocrinida - Isocrinidae
Balanocrinus sp. n. sp. Agassiz 1847
1 individual
Balanocrinus n. sp. A
Tyrolecrinus tyrolensis (Laube 1865)
1 individual
    = Tyrolecrinus wugangi Stiller 2011
Stiller 2011
Anthozoa - Scleractinia - Margarophylliidae
Neoconophyllia leidapoensis Deng and Kong 1984
1 individual
Promargarosmilia sp. Deng and Zhang 1985
3 individuals
Promargarosmilia sp. A
Demospongiae - Agelasida - Catenispongiidae
? Hartmanina bangtoupoensis
1 individual
Rhodophyceae - Solenoporaceae
Solenopora sp. Dybowski 1877
336 individuals
Solenopora sp. A and calcareous red algae gen. et sp. indet.
see common names

Geography
Country:China State/province:Guizhou
Coordinates: 26.3° North, 106.7° East (view map)
Paleocoordinates:13.3° North, 95.1° East
Basis of coordinate:estimated from map
Geographic resolution:small collection
Time
Period:Triassic Epoch:Middle Triassic
Stage:Anisian 10 m.y. bin:Triassic 2
*Period:Middle Triassic
*International age/stage:Late/Upper Anisian *Local age/stage:Late/Upper Pelsonian - Early/Lower Illyrian
Key time interval:Pelsonian - Illyrian
Age range of interval:244.70000 - 242.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Qingyan Member:Leidapo
Local section:Bangtoupo Local bed:32
Local order:bottom to top
Stratigraphic resolution:group of beds
Stratigraphy comments: Basal upper Anisian. Schreyerites binodosus and Paraceratites trinodosus zones. Yuqing subformation
Lithology and environment
Primary lithology: lithified calcareous claystone
Secondary lithology: lithified marl
Includes fossils?Y
Includes fossils?Y
Environment:open shallow subtidal Tectonic setting:passive margin
Taphonomy
Modes of preservation:body,original calcite
Size of fossils:macrofossils,mesofossils
Articulated whole bodies:some
Bioerosion:occasional
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:all microfossils
Collection methods:bulk,mechanical,peel or thin section,field collection
Collection size:732 individuals
Reason for describing collection:paleoecologic analysis
Collection method comments: All abundance data given as "statistical individuals"
Bivalves, brachiopods: Double-valved shells + higher number of single shells (right or left)
Echinoderms, vertebrates: one indivudual if number of parts more than parts in living animal. crinoids > 150 columnalia = 1 individual,
Others: all fragments as one
Metadata
Database number:31620
Authorizer:W. Kiessling, M. Aberhan, M. Clapham Enterer:W. Kiessling, M. Clapham, S. Nurnberg, M. Apel
Modifier:W. Kiessling Research group:marine invertebrate
Created:2003-05-20 02:03:57 Last modified:2003-05-20 05:03:59
Access level:the public Released:2003-05-20 02:03:57
Creative Commons license:CC BY
Reference information

Primary reference:
8524. F. Stiller. 2001. Fossilvergesellschaftungen, Paläoökologie und paläosynökologische Entwicklung im Oberen Anisium (Mittlere Trias) von Qingyan, insbesondere Bangtoupo, Provinz Guizhou, Südwestchina. Münstersche Forschungen zur Geologie und Paläontologie 92:1-523 [W. Kiessling/W. Kiessling/W. Kiessling]

Secondary references:

38551 F. Stiller. 2011. An early isocrinid sea lily from the middle to late Anisian boundary (Middle Triassic) of south-west China - evidence for a far-east Tethyan origin of the family Isocrinidae. Palaeontology 54(6):1415-1433 [M. Clapham/M. Clapham]
37155 F. Stiller and J. Chen. 2004. Eophilobryoidella sinoanisica new genus and species, an early Philobryid bivalve from the upper Anisian (middle Triassic) of Qingyan, southwestern China. Journal of Paleontology 78(2):414-419 [M. Aberhan/D. Wellmanns/M. Uhen]