Bangtoupo F40, Qingyan: Pelsonian - Illyrian, China

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Corals indet.
1 individual
solitary corals ge. et sp. indet.
Crinoids indet.
1 individual
Crinoidenstilglieder gen. et sp. indet.
Anthozoa - Scleractinia - Pamiroseriidae
Pamiroseris cf. silesiaca (Beyrich 1852)
1 individual
Anthozoa - Scleractinia - Margarophylliidae
Neoconophyllia leidapoensis Deng and Kong 1984
2 individuals
Actinopteri - Lepisosteiformes
? Lepidotes sp. Agassiz 1832
1 element
Lepidotes? sp. A (isolated teeth)
Crinoidea - Millericrinida - Bangtoupocrinidae
Silesiacrinus parvus Stiller 2000
1 individual
Bangtoupocrinus kokeni Stiller 2000
1 individual
Crinoidea - Qingyanocrinidae
Qingyanocrinus kueichounensis (Dubatolova and Shao 1959)
1 individual
Crinoidea - Encrinida - Encrinidae
Encrinus cf. liliiformis Lamarck 1801
6 individuals
Crinoidea - Isocrinida - Isocrinidae
Tyrolecrinus tyrolensis (Laube 1865)
1 individual
    = Tyrolecrinus wugangi Stiller 2011
Stiller 2011
Balanocrinus sp. n. sp. Agassiz 1847
1 individual
Balanocrinus n. sp. A
Echinoidea - Cidaroida - Cidaridae
"Cidaris" bangtoupoensis
1 individual
"Cidaris" lineola
1 individual
"Cidaris" venustespinosa
1 individual
"Cidaris" wachangpoensis
1 individual
unclassified
Bryozoa indet. Ehrenberg 1831
2 individuals
Bryozoa gen. et sp. indet. A
Brachiopoda indet. Cuvier 1805
1 individual
Brachiopoda gen. et sp. indet.
Rhynchonellata - Athyridida - Neoretziidae
Schwagerispira fuchsi (Koken 1900)
1 individual
Rhynchonellata - Terebratulida - Angustothyrididae
Angustothyris angustaeformis (Boekh 1872)
85 individuals
Rhynchonellata - Terebratulida - Zeilleriidae
Aulacothyris angustaeformis Yang and Xu 1966
1 individual
recombined as Sacothyris angustaeformis
Rhynchonellata - Spiriferinida - Bittnerulidae
Thecocyrtelloidea tubulosa Yang and Xu 1966
10 individuals
Rhynchonellata - Spiriferinida - Laballidae
Paralepismatina semiconica Yang and Xu 1966
1 individual
Rhynchonellata - Spiriferinida - Cyrtinidae
"Cyrtina" indet. Davidson 1858
4 individuals
Rhynchonellata - Spiriferinida - Mentzeliidae
Mentzelia mentzeli (Dunker 1851)
8 individuals
Mentzelia subspherica Yang and Xu 1966
39 individuals
Hirsutella multicostata (Yang and Xu 1966)
5 individuals
recombined as Dagyssia multicostata
Rhynchonellata - Spiriferinida - Lepismatinidae
Lepismatina hsui Wang 1955
4 individuals
Rhynchonellata - Rhynchonellida - Norellidae
Diholkorhynchia sinensis (Koken 1910)
64 individuals
Nudirostralina subtrinodosi Yang and Xu 1966
2 individuals
Rhynchonellata - Rhynchonellida - Allorhynchidae
Septaliphorioidea paucicostata Yang and Xu 1966
11 individuals
Polychaeta - Sabellida - Serpulidae
"Spirorbis" qingyani
1 individual
Ruxingella lianjiangensis
33 individuals
Weixiserpula weixi
179 individuals
Scaphopoda - Dentaliida - Laevidentaliidae
Laevidentalium bangtoupoensis
15 individuals
original and current combination Dentalium (Laevidentalium)
Bivalvia
Bivalvia indet. Linnaeus 1758
14 individuals
Bivalvia gen. et sp. indet.
Bivalvia - Mytilida - Mytilidae
Modiolus (Modiolus) salzstettensis (Hohenstein 1913)
9 individuals
Bivalvia - Cyrtodontida - Pichleriidae
Elegantarca subareata Chen 1976
14 individuals
Bivalvia - Arcida - Parallelodontidae
Parallelodon beyrichi
1 individual
Bivalvia - Arcida - Philobryidae
Philobryidae indet. n. gen., n. sp. Bernard 1897
32 individuals
Philobryidae n. gen. A, n. sp. A
    = Eophilobryoidella sinoanisica Stiller and Chen 2004
Stiller and Chen 2004
Bivalvia - Ostreida
Ostreoidea indet. n. gen., n. sp. (Rafinesque 1815)
9 individuals
original and current combination Ostreacea
Ostreoidea n. gen. A, n. sp. A
Bivalvia - Ostreida - Ostreidae
Lopha sp. n. sp. Röding 1798
12 individuals
Lopha n. sp. A
Bivalvia - Ostreida - Pteriidae
Pteria caudata
11 individuals
Pteria cassiana (Bittner 1895)
2 individuals
Pteria rugosa Chen 1976
1 individual
Arcavicula (Arcavicula) guizhouensis
27 individuals
Bivalvia - Ostreida - Cassianellidae
Cassianella sulcata
20 individuals
Cassianella ecki sulcata
Cassianella qingyanensis
27 individuals
Cassianella subcislonensis
36 individuals
Cassianella simplex Chen 1976
38 individuals
Bivalvia - Ostreida - Bakevelliidae
Gervillia (Cultriopsis) angulata Münster 1841
2 individuals
Gervillaria subelegans (Chen 1976)
2 individuals
Bivalvia - Pectinida - Dimyidae
Protostrea sinensis (Hsu 1943)
11 individuals
Bivalvia - Pectinida - Pectinidae
Praechlamys schroeteri
3 individuals
Praechlamys stenodictya
1 individual
Bivalvia - Pectinida - Terquemiidae
Terquemiidae indet. n. gen., n. sp. Cox 1964
1 individual
Terquemiidae n. gen. A, n. sp. A
Terquemiidae indet. n. gen., n. sp. Cox 1964
19 individuals
Terquemiidae n. gen. B, n. sp. A
Newaagia multicostata
1 individual
Newaagia sessilis
9 individuals
Bivalvia - Pectinida - Plicatulidae
Plicatula sp. n. sp. Lamarck 1801
2 individuals
Plicatula n. sp. A
Pseudoplacunopsis sp. n. sp. Bittner 1895
1 individual
Pseudoplacunopsis n. sp. A
Bivalvia - Pectinida - Anomiidae
Placunopsis subplana
12 individuals
Bivalvia - Pectinida - Limidae
Palaeolima acutecostata
2 individuals
Palaeolima subcostata
1 individual
Mysidioptera (Mysidioptera) punctata Chen 1976
1 individual
Plagiostoma sp. n. sp. Sowerby 1814
2 individuals
Plagiostoma n. sp. A
Bivalvia - Pectinida - Hunanopectinidae
Leptochondria plana
2 individuals
Leptochondria albertii (Goldfuss 1838)
1 individual
Bivalvia - Carditida - Cardiniidae
Cardinia sp. n. sp. Agassiz 1841
1 individual
Cardinia n. sp. A
Bivalvia - Carditida - Astartidae
Astarte emacerata
1 individual
Coelopis (Coelopis) sp. n. sp. Munier-Chalmas 1887
1 individual
Coelopis (Coelopis) n. sp. A
Bivalvia - Lucinida - Lucinidae
Schafhaeutlia astartiformis (Münster 1841)
18 individuals
recombined as Fimbria astartiformis
Bivalvia - Cardiida - Kalenteridae
Pseudomyoconcha goldfussi
1 individual
Bivalvia - Trigoniida - Myophoriidae
Quadratia quadrata Yin 1974
1 individual
recombined as Guizhoumyophoria quadrata
Elegantinia kulunensis
1 individual
Elegantinia sp. n. sp. (Waagen 1907)
29 individuals
Elegantinia n. sp. A
Elegantinia elegans (Dunker 1849)
17 individuals
Costatoria multiformis
11 individuals
Costatoria proharpa multiformis
Bivalvia - Nuculanida - Malletiidae
Phaenodesmia sp. n. sp. Bittner 1895
2 individuals
Phaenodesmia n. sp. A
Bivalvia - Nuculida - Nuculidae
Palaeonucula qingyanensis Chen 1976
8 individuals
Cephalopoda - Pseudorthocerida - Trematoceratidae
Michelinoceras cf. nyalamense Chen 1981
1 individual
recombined as Pseudotemperoceras nyalamense
Cephalopoda - Ammonoidea
Ammonoidea indet. Zittel 1884
1 individual
Ammonoidea gen. et sp. indet.
Cephalopoda - Ceratitida - Ceratitidae
Bulogites multicostatus Zhao and Wang 1974
1 individual
Gastropoda
Gastropoda indet. Cuvier 1795
4 individuals
Gastropoda gen. et sp. indet.
Gastropoda - Eucyclidae
Eucycloscala sp. n. sp. Cossmann 1895
4 individuals
Eucycloscala n. sp. A
Eucycloscala sp. n. sp. Cossmann 1895
16 individuals
Eucycloscala n. sp. B
Eucycloscala sp. n. sp. Cossmann 1895
3 individuals
Eucycloscala n. sp. C
Eucycloscala obliquicostata Yin and Yochelson 1983
8 individuals
Riselloidea sp. n. sp. Cossmann 1909
2 individuals
spelled with current rank as Calliotropis (Riselloidea)
Riselloidea n. sp. B
Gastropoda - Schizogoniidae
Schizogonium pentagonum Yu et al. 1974
1 individual
Gastropoda - Turbinidae
Turbinidae indet. n. gen., n. sp. Rafinesque 1815
28 individuals
Turbinidae n. gen. A, n. sp. A
Turbinidae indet. n. gen., n. sp. Rafinesque 1815
2 individuals
Turbinidae n. gen. C, n. sp. A
Gastropoda - Discohelicidae
Discohelicidae indet. n. gen., n. sp. Schröder 1995
3 individuals
Discohelicidae n. gen. A, n. sp. A
Gastropoda - Cycloneritida - Tricolnaticopsidae
Pachyomphalus sp. n. sp. Böhm 1895
16 individuals
Pachyomphalus n. sp. A
Gastropoda - Cycloneritida - Delphinulopsidae
Delphinulopsis binodosa (Münster 1841)
2 individuals
Delphinulopsis cf. laubei Kittl 1892
1 individual
synonym of Delphinulopsis binodosa
Gastropoda - Heterostropha - Tofanellidae
Tofanellidae indet. n. gen., n. sp. Bandel 1995
2 individuals
Tofanellidae n. gen. A, n. sp. A
Gastropoda - Heterostropha - Gordenellidae
Promathilda sp. n. sp. (Andreae 1887)
2 individuals
original and current combination Promathildia
Promathilda n. sp. C
Gastropoda
Omphaloptycha gregaria (Schlotheim 1820)
1 individual
original and current combination Buccinites gregarius
Gastropoda - Euomphalina
Triassocirrus guizhouensis Yin and Yochelson 1983
2 individuals
Gastropoda - Murchisoniina - Spirostylidae
Spirostylus subcolumnaris (Münster 1841)
2 individuals
Gastropoda - Murchisoniina - Loxonematidae
Tyrsoecus multinodosa Yin and Yochelson 1983
8 individuals
Gastropoda - Murchisoniina - Zygopleuridae
Zygopleura arctecostata (Münster 1841)
1 individual
Gastropoda - Murchisoniina - Wortheniellidae
Wortheniella conica
11 individuals
Wortheniella nuda
1 individual
Wortheniella regulocostata
1 individual
Wortheniella tuberculifera
1 individual
Wortheniella xui
1 individual
Wortheniella sp. n. sp. Schwardt 1992
1 individual
Wortheniella n. sp. D
Wortheniella indet. Schwardt 1992
2 individuals
Wortheniella sp. indet.
Gastropoda - Murchisoniina - Gosseletinidae
Dictyotomaria triassica (Yin and Yochelson 1983)
1 individual
Gastropoda - Murchisoniina - Rhaphistomellidae
Sisenna sp. n. sp. Koken 1896
2 individuals
Sisenna n. sp. A
? Sisenna sp. n. sp. Koken 1896
2 individuals
Sisenna? n. sp. B
Gastropoda - Murchisoniina - Murchisoniidae
Murchisoniidae indet. n. gen., n. sp. Koken 1896
1 individual
Murchisoniidae n. gen. A, n. sp. A
Gastropoda - Neritariidae
Neritaria sp. n. sp. Koken 1892
6 individuals
Neritaria n. sp. A
Neritaria sp. n. sp. Koken 1892
32 individuals
Neritaria n. sp. B
Trachynerita distincta (Kittl 1894)
1 individual
Gastropoda - Naticopsidae
Hologyra guizhoui Yin and Yochelson 1983
18 individuals
Naticopsis (Naticopsis) ladina Kittl 1892
2 individuals
Naticopsis (Naticopsis) sp. n. sp. M'Coy 1844
2 individuals
Naticopsis (Naticopsis) n. sp. A
Gastropoda - Coelostylinidae
Coelostylina ahlburgi (Assmann 1924)
2 individuals
Coelostylina liscaviensis (Giebel 1856)
3 individuals
Trypanostylus cf. turritelliformis
3 individuals
Trypanostylus sp. n. sp. Cossmann 1895
2 individuals
Trypanostylus n. sp. B
Trypanostylus konincki (Münster 1841)
9 individuals
Gastropoda - Goniasmatidae
Cheilotomona pentagonum Yu et al. 1974
3 individuals
Gastropoda - Turritellidae
Provermicularia sp. n. sp. Kittl 1899
1 individual
spelled with current rank as Vermicularia (Provermicularia)
Provermicularia n. sp. A
Rhodophyceae - Solenoporaceae
Solenopora sp. Dybowski 1877
744 individuals
Solenopora sp. A and calcareous red algae gen. et sp. indet.
see common names

Geography
Country:China State/province:Guizhou
Coordinates: 26.3° North, 106.7° East (view map)
Paleocoordinates:13.3° North, 95.1° East
Basis of coordinate:estimated from map
Geographic resolution:small collection
Time
Period:Triassic Epoch:Middle Triassic
Stage:Anisian 10 m.y. bin:Triassic 2
*Period:Middle Triassic
*International age/stage:Late/Upper Anisian *Local age/stage:Late/Upper Pelsonian - Early/Lower Illyrian
Key time interval:Pelsonian - Illyrian Ammonoid zone: Schreyerites binodosus - Paraceratites trinodosus
Age range of interval:244.70000 - 242.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Qingyan Member:Leidapo
Local section:Bangtoupo Local bed:40 m
Local order:bottom to top
Stratigraphic resolution:group of beds
Stratigraphy comments: Basal upper Anisian. Schreyerites binodosus and Paraceratites trinodosus zones. Yuqing subformation
Lithology and environment
Primary lithology: lithified calcareous claystone
Secondary lithology: lithified marl
Includes fossils?Y
Includes fossils?Y
Environment:open shallow subtidal Tectonic setting:passive margin
Taphonomy
Modes of preservation:body,original calcite
Size of fossils:macrofossils,mesofossils
Articulated whole bodies:some
Bioerosion:occasional
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:all microfossils
Collection methods:bulk,mechanical,peel or thin section,field collection
Collection size:1128 individuals
Reason for describing collection:paleoecologic analysis
Collection method comments: All abundance data given as "statistical individuals"
Bivalves, brachiopods: Double-valved shells + higher number of single shells (right or left)
Echinoderms, vertebrates: one indivudual if number of parts more than parts in living animal. crinoids > 150 columnalia = 1 individual,
Others: all fragments as one
Metadata
Also known as:Bangtoupo F40, Qingyan, China
Database number:31623
Authorizer:W. Kiessling, M. Aberhan, M. Clapham Enterer:W. Kiessling, S. Nurnberg, M. Clapham, M. Apel
Modifier:P. Wagner Research group:marine invertebrate
Created:2003-05-20 02:08:13 Last modified:2021-09-23 21:28:24
Access level:the public Released:2003-05-20 02:08:13
Creative Commons license:CC BY
Reference information

Primary reference:
8524. F. Stiller. 2001. Fossilvergesellschaftungen, Paläoökologie und paläosynökologische Entwicklung im Oberen Anisium (Mittlere Trias) von Qingyan, insbesondere Bangtoupo, Provinz Guizhou, Südwestchina. Münstersche Forschungen zur Geologie und Paläontologie 92:1-523 [W. Kiessling/W. Kiessling/W. Kiessling]

Secondary references:

38551 F. Stiller. 2011. An early isocrinid sea lily from the middle to late Anisian boundary (Middle Triassic) of south-west China - evidence for a far-east Tethyan origin of the family Isocrinidae. Palaeontology 54(6):1415-1433 [M. Clapham/M. Clapham]
37155 F. Stiller and J. Chen. 2004. Eophilobryoidella sinoanisica new genus and species, an early Philobryid bivalve from the upper Anisian (middle Triassic) of Qingyan, southwestern China. Journal of Paleontology 78(2):414-419 [M. Aberhan/D. Wellmanns/M. Uhen]