Bangtoupo F45, Qingyan, China: Pelsonian - Illyrian, China

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Anthozoa - Scleractinia - Margarophylliidae
Margarophyllia vortica Deng and Kong 1984
2 individuals
Neoconophyllia leidapoensis Deng and Kong 1984
1 individual
Semidistichophyllum guizhouensis Deng and Kong 1984
1 individual
Crinoidea - Millericrinida - Bangtoupocrinidae
Bangtoupocrinus kokeni Stiller 2000
1 individual
Crinoidea - Qingyanocrinidae
Qingyanocrinus kueichounensis (Dubatolova and Shao 1959)
1 individual
Crinoidea - Encrinida - Encrinidae
Encrinus cf. liliiformis Lamarck 1801
3 individuals
Crinoidea - Isocrinida - Isocrinidae
Tyrolecrinus tyrolensis (Laube 1865)
2 individuals
    = Tyrolecrinus wugangi Stiller 2011
Stiller 2011
Balanocrinus sp. n. sp. Agassiz 1847
1 individual
Balanocrinus n. sp. A
Echinoidea - Cidaroida - Cidaridae
"Cidaris" bangtoupoensis
1 individual
"Cidaris" lineola
1 individual
"Cidaris" venustespinosa
1 individual
Echinoidea
"Cidaris" wissmanni Desor 1847
1 individual
nomen dubium belonging to Echinoidea
unclassified
Bryozoa indet. Ehrenberg 1831
3 individuals
Bryozoa gen. et sp. indet. A
Brachiopoda indet. Cuvier 1805
2 individuals
Brachiopoda gen. et sp. indet.
Rhynchonellata - Terebratulida - Angustothyrididae
Angustothyris angustaeformis (Boekh 1872)
55 individuals
Rhynchonellata - Terebratulida - Dielasmatidae
Adygella elongata Yang and Xu 1966
1 individual
recombined as Coenothyris elongata
Rhynchonellata - Terebratulida - Zeilleriidae
Aulacothyris angustaeformis Yang and Xu 1966
2 individuals
recombined as Sacothyris angustaeformis
Rhynchonellata - Spiriferinida - Bittnerulidae
Thecocyrtelloidea tubulosa Yang and Xu 1966
23 individuals
Neocyrtina mixodeltidiumosa Yang and Xu 1966
1 individual
Rhynchonellata - Spiriferinida - Cyrtinidae
"Cyrtina" indet. Davidson 1858
9 individuals
Rhynchonellata - Spiriferinida - Mentzeliidae
Mentzelia mentzeli (Dunker 1851)
23 individuals
Mentzelia subspherica Yang and Xu 1966
101 individuals
Hirsutella multicostata (Yang and Xu 1966)
1 individual
recombined as Dagyssia multicostata
Rhynchonellata - Spiriferinida - Lepismatinidae
Lepismatina hsui Wang 1955
15 individuals
Rhynchonellata - Rhynchonellida - Cyclothyrididae
Costirhynchopsis sinensis (Yang and Xu 1966)
1 individual
Rhynchonellata - Rhynchonellida - Norellidae
Diholkorhynchia sinensis (Koken 1910)
47 individuals
Nudirostralina subtrinodosi Yang and Xu 1966
14 individuals
Rhynchonellata - Rhynchonellida - Allorhynchidae
Septaliphorioidea paucicostata Yang and Xu 1966
23 individuals
Polychaeta - Sabellida - Serpulidae
"Spirorbis" qingyani
1 individual
Ruxingella lianjiangensis
19 individuals
Weixiserpula weixi
42 individuals
Scaphopoda - Dentaliida - Laevidentaliidae
Laevidentalium bangtoupoensis
3 individuals
original and current combination Dentalium (Laevidentalium)
Bivalvia
Bivalvia indet. Linnaeus 1758
11 individuals
Bivalvia gen. et sp. indet.
Bivalvia - Mytilida - Mytilidae
Modiolus (Modiolus) salzstettensis (Hohenstein 1913)
5 individuals
Botulopsis cf. cassiana Reis 1926
2 individuals
replaced by Botulopsis reisi
Bivalvia - Cyrtodontida - Pichleriidae
Elegantarca subareata Chen 1976
2 individuals
Bivalvia - Arcida - Parallelodontidae
Parallelodon beyrichi
1 individual
Bivalvia - Arcida - Philobryidae
Philobryidae indet. n. gen., n. sp. Bernard 1897
10 individuals
Philobryidae n. gen. A, n. sp. A
    = Eophilobryoidella sinoanisica Stiller and Chen 2004
Stiller and Chen 2004
Bivalvia - Ostreida
Ostreoidea indet. n. gen., n. sp. (Rafinesque 1815)
17 individuals
original and current combination Ostreacea
Ostreoidea n. gen. A, n. sp. A
Bivalvia - Ostreida - Ostreidae
Lopha sp. n. sp. Röding 1798
1 individual
Lopha n. sp. A
Bivalvia - Ostreida - Pinnidae
Atrina sp. n. sp. Gray 1842
1 individual
Atrina n. sp. A
Bivalvia - Ostreida - Pteriidae
Pteria caudata
3 individuals
Pteria cassiana (Bittner 1895)
3 individuals
Pteria rugosa Chen 1976
10 individuals
Arcavicula sp. n. sp. Cox 1964
2 individuals
Arcavicula n. subgen. A, n. sp. A
Arcavicula (Arcavicula) guizhouensis
9 individuals
Bivalvia - Ostreida - Cassianellidae
Cassianella sulcata
34 individuals
Cassianella ecki sulcata
Cassianella qingyanensis
11 individuals
Cassianella subcislonensis
14 individuals
Cassianella simplex Chen 1976
26 individuals
Bivalvia - Ostreida - Bakevelliidae
Gervillaria subelegans (Chen 1976)
2 individuals
Bivalvia - Ostreida - Posidoniidae
Posidonia cf. wengensis (Wissmann 1841)
1 individual
recombined as Bositra wengensis
Bivalvia - Pectinida - Dimyidae
Protostrea sinensis (Hsu 1943)
10 individuals
Bivalvia - Pectinida - Pectinidae
Pleuronectites difformis Chen 1976
1 individual
Praechlamys schroeteri
2 individuals
Bivalvia - Pectinida - Terquemiidae
Terquemiidae indet. n. gen., n. sp. Cox 1964
15 individuals
Terquemiidae n. gen. B, n. sp. A
Newaagia multicostata
2 individuals
Newaagia sessilis
7 individuals
Newaagia sp. n. sp. Hertlein 1952
1 individual
Newaagia n. sp. A
Bivalvia - Pectinida - Plicatulidae
Pseudoplacunopsis sp. n. sp. Bittner 1895
3 individuals
Pseudoplacunopsis n. sp. A
Bivalvia - Pectinida - Anomiidae
Placunopsis subplana
3 individuals
Bivalvia - Pectinida - Limidae
Plagiostoma sp. n. sp. Sowerby 1814
1 individual
Plagiostoma n. sp. A
Bivalvia - Pectinida - Hunanopectinidae
Leptochondria plana
3 individuals
Leptochondria albertii (Goldfuss 1838)
3 individuals
Bivalvia - Carditida - Astartidae
Opinae indet. n. gen., n. sp. Chavan 1952
1 individual
Opinae n. gen. A, n. sp. A
Bivalvia - Lucinida - Lucinidae
Schafhaeutlia astartiformis (Münster 1841)
6 individuals
recombined as Fimbria astartiformis
Bivalvia - Trigoniida - Myophoriidae
Quadratia quadrata Yin 1974
1 individual
recombined as Guizhoumyophoria quadrata
Elegantinia kulunensis
1 individual
Elegantinia sp. n. sp. (Waagen 1907)
16 individuals
Elegantinia n. sp. A
Elegantinia elegans (Dunker 1849)
14 individuals
Costatoria multiformis
10 individuals
Costatoria proharpa multiformis
Bivalvia - Nuculanida - Malletiidae
Palaeoneilo distincta
1 individual
Bivalvia - Nuculida - Nuculidae
Palaeonucula qingyanensis Chen 1976
2 individuals
Cephalopoda - Pseudorthocerida - Trematoceratidae
Michelinoceras cf. nyalamense Chen 1981
1 individual
recombined as Pseudotemperoceras nyalamense
Gastropoda
Gastropoda indet. Cuvier 1795
2 individuals
Gastropoda gen. et sp. indet.
Gastropoda - Eucyclidae
Eucycloscala sp. n. sp. Cossmann 1895
3 individuals
Eucycloscala n. sp. A
Eucycloscala sp. n. sp. Cossmann 1895
2 individuals
Eucycloscala n. sp. B
Eucycloscala binodosa (Münster 1841)
3 individuals
Eucycloscala obliquicostata Yin and Yochelson 1983
5 individuals
Riselloidea sp. n. sp. Cossmann 1909
1 individual
spelled with current rank as Calliotropis (Riselloidea)
Riselloidea n. sp. B
Riselloidea guizhouensis Yu et al. 1974
1 individual
Gastropoda - Discohelicidae
Discohelicidae indet. n. gen., n. sp. Schröder 1995
1 individual
Discohelicidae n. gen. A, n. sp. A
Gastropoda - Cycloneritida - Tricolnaticopsidae
Pachyomphalus sp. n. sp. Böhm 1895
6 individuals
Pachyomphalus n. sp. A
Gastropoda - Cycloneritida - Delphinulopsidae
Delphinulopsis binodosa (Münster 1841)
1 individual
Gastropoda - Cycloneritida - Scalaneritinidae
Natiria striatocostata (Braun 1841)
1 individual
recombined as Ladinaticella striatocostata
Gastropoda
Omphaloptycha gregaria (Schlotheim 1820)
2 individuals
original and current combination Buccinites gregarius
Gastropoda - Euomphalina
Triassocirrus applanatus (Kutassy 1937)
2 individuals
Gastropoda - Murchisoniina - Zygopleuridae
Allocosmia cf. turricula (Assmann 1937)
1 individual
Gastropoda - Murchisoniina - Wortheniellidae
Wortheniella conica
3 individuals
Wortheniella hausmanni
4 individuals
Wortheniella johannisaustriae
2 individuals
Wortheniella nuda
2 individuals
Wortheniella sp. n. sp. Schwardt 1992
1 individual
Wortheniella n. sp. C
Wortheniella indet. Schwardt 1992
2 individuals
Wortheniella sp. indet.
Gastropoda - Murchisoniina - Gosseletinidae
Dictyotomaria triassica (Yin and Yochelson 1983)
1 individual
Gastropoda - Murchisoniina - Rhaphistomellidae
Sisenna sp. n. sp. Koken 1896
3 individuals
Sisenna n. sp. A
? Sisenna sp. n. sp. Koken 1896
1 individual
Sisenna? n. sp. B
Gastropoda - Murchisoniina - Murchisoniidae
Murchisoniidae indet. n. gen., n. sp. Koken 1896
3 individuals
Murchisoniidae n. gen. B, n. sp. A
Gastropoda - Neritariidae
Neritaria sp. n. sp. Koken 1892
1 individual
Neritaria n. sp. A
Neritaria sp. n. sp. Koken 1892
4 individuals
Neritaria n. sp. B
Gastropoda - Naticopsidae
Hologyra guizhoui Yin and Yochelson 1983
3 individuals
Naticopsis (Naticopsis) ladina Kittl 1892
1 individual
Naticopsis (Naticopsis) signata (Koken 1900)
7 individuals
Gastropoda - Coelostylinidae
Coelostylina ahlburgi (Assmann 1924)
1 individual
? Siphonophyla sp. Kittl 1894
1 individual
spelled with current rank as Coelostylina (Siphonophyla)
Siphonophyla sp. A
Coelostylina liscaviensis (Giebel 1856)
4 individuals
Gastropoda - Goniasmatidae
Cheilotomona pentagonum Yu et al. 1974
2 individuals
Rhodophyceae - Solenoporaceae
Solenopora sp. Dybowski 1877
57 individuals
Solenopora sp. A and calcareous red algae gen. et sp. indet.
see common names

Geography
Country:China State/province:Guizhou
Coordinates: 26.3° North, 106.7° East (view map)
Paleocoordinates:13.3° North, 95.1° East
Basis of coordinate:estimated from map
Geographic resolution:small collection
Time
Period:Triassic Epoch:Middle Triassic
Stage:Anisian 10 m.y. bin:Triassic 2
*Period:Middle Triassic
*International age/stage:Late/Upper Anisian *Local age/stage:Late/Upper Pelsonian - Early/Lower Illyrian
Key time interval:Pelsonian - Illyrian
Age range of interval:244.70000 - 242.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Qingyan Member:Leidapo
Local section:Bangtoupo Local bed:45
Local order:bottom to top
Stratigraphic resolution:group of beds
Stratigraphy comments: Basal upper Anisian. Schreyerites binodosus and Paraceratites trinodosus zones. Yuqing subformation
Lithology and environment
Primary lithology: lithified calcareous claystone
Secondary lithology: lithified marl
Includes fossils?Y
Includes fossils?Y
Environment:open shallow subtidal Tectonic setting:passive margin
Taphonomy
Modes of preservation:body,original calcite
Size of fossils:macrofossils,mesofossils
Articulated whole bodies:some
Bioerosion:occasional
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:all microfossils
Collection methods:bulk,mechanical,peel or thin section,field collection
Collection size:748 individuals
Reason for describing collection:paleoecologic analysis
Collection method comments: All abundance data given as "statistical individuals"
Bivalves, brachiopods: Double-valved shells + higher number of single shells (right or left)
Echinoderms, vertebrates: one indivudual if number of parts more than parts in living animal. crinoids > 150 columnalia = 1 individual,
Others: all fragments as one
Metadata
Database number:31625
Authorizer:W. Kiessling, M. Aberhan, M. Clapham Enterer:W. Kiessling, S. Nurnberg, M. Clapham, M. Apel
Modifier:W. Kiessling Research group:marine invertebrate
Created:2003-05-20 02:09:29 Last modified:2003-05-20 05:09:31
Access level:the public Released:2003-05-20 02:09:29
Creative Commons license:CC BY
Reference information

Primary reference:
8524. F. Stiller. 2001. Fossilvergesellschaftungen, Paläoökologie und paläosynökologische Entwicklung im Oberen Anisium (Mittlere Trias) von Qingyan, insbesondere Bangtoupo, Provinz Guizhou, Südwestchina. Münstersche Forschungen zur Geologie und Paläontologie 92:1-523 [W. Kiessling/W. Kiessling/W. Kiessling]

Secondary references:

38551 F. Stiller. 2011. An early isocrinid sea lily from the middle to late Anisian boundary (Middle Triassic) of south-west China - evidence for a far-east Tethyan origin of the family Isocrinidae. Palaeontology 54(6):1415-1433 [M. Clapham/M. Clapham]
37155 F. Stiller and J. Chen. 2004. Eophilobryoidella sinoanisica new genus and species, an early Philobryid bivalve from the upper Anisian (middle Triassic) of Qingyan, southwestern China. Journal of Paleontology 78(2):414-419 [M. Aberhan/D. Wellmanns/M. Uhen]