Maramena: Turolian, Greece
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
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cf. Belgrandiella indet.
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Schmidt-Kittler et al. 1995 | |||||||||
Mastus sp.
|
Schmidt-Kittler et al. 1995 | |||||||||
Milax sp.
|
Schmidt-Kittler et al. 1995 | |||||||||
or Tandonia sp. | ||||||||||
Cadona sp.
|
Schmidt-Kittler et al. 1995 | |||||||||
Ouzoceros aff. gracilis
|
Schmidt-Kittler et al. 1995 | |||||||||
Norbertia hellenica n. gen., n. sp.
|
Schmidt-Kittler et al. 1995 | |||||||||
Ommatotriton sp.
|
Georgalis et al. 2019 | |||||||||
M1: a vertebra (UU MAA 7489) and two femurs (UU MAA 7248 and UU MAA 7509). | ||||||||||
Gastropoda
- Aciculidae
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cf. Platyla sp.
Moquin-Tandon 1855
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Schmidt-Kittler et al. 1995 | |||||||||
Gastropoda
- Bithyniidae
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Bithynia sp.
Leach 1818
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Schmidt-Kittler et al. 1995 | |||||||||
Gastropoda
- Heterostropha
- Planorbidae
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Gyraulus sp.
Charpentier 1837
|
Schmidt-Kittler et al. 1995 | |||||||||
Planorbarius sp.
Dumeril 1806
|
Schmidt-Kittler et al. 1995 | |||||||||
cf. conru-mantelli | ||||||||||
Planorbis sp.
Muller 1774
|
Schmidt-Kittler et al. 1995 | |||||||||
Gastropoda
- Heterostropha
- Carychiidae
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Carychium cf. nouleti
Bourguignat 1857
|
Schmidt-Kittler et al. 1995 | |||||||||
Gastropoda
- Heterostropha
- Cochlicopidae
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Cochlicopa sp.
Ferussac 1821
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Schmidt-Kittler et al. 1995 | |||||||||
Gastropoda
- Heterostropha
- Parmacellidae
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Parmacella sp.
Cuvier 1804
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Schmidt-Kittler et al. 1995 | |||||||||
nov | ||||||||||
Gastropoda
- Heterostropha
- Limacidae
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cf. Limax sp.
Linnaeus 1758
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Schmidt-Kittler et al. 1995 | |||||||||
Gastropoda
- Heterostropha
- Pristilomatidae
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Vitrea sp.
Fitzinger 1833
|
Schmidt-Kittler et al. 1995 | |||||||||
Gastropoda
- Heterostropha
- Clausiliidae
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Clausiliidae indet.
Gray 1855
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Schmidt-Kittler et al. 1995 | |||||||||
Gastropoda
- Heterostropha
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Helicoidea indet.
Rafinesque 1815
|
Schmidt-Kittler et al. 1995 | |||||||||
sp. nov. | ||||||||||
Gastropoda
- Heterostropha
- Elonidae
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Klikia goniostoma
(Sandberger 1875)
|
Schmidt-Kittler et al. 1995 | |||||||||
Bivalvia
- Cardiida
- Sphaeriidae
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Pisidium casertanum
(Poli 1791)
|
Schmidt-Kittler et al. 1995 | |||||||||
original and current combination Cardium casertanum | ||||||||||
Amphibia
- Temnospondyli
- Ranidae
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Ranidae indet.
Rafinesque 1814
|
Georgalis et al. 2019 | |||||||||
M2: three ilia (UU MAA 7195); M3: three humeri (UU MAA 7231). | ||||||||||
Pelophylax sp.
Fitzinger 1843
|
Georgalis et al. 2019 | |||||||||
M2: an angular (UU MAA 7194); M3: a coracoid (UU MAA 7233) and three ilia (UU MAA 7232). | ||||||||||
Amphibia
- Temnospondyli
- Hylidae
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Hyla sp.
Laurenti 1768
|
Georgalis et al. 2019 | |||||||||
M1: seven humeri (UU MAA 7119, UU MAA 7244, UU MAA 7251, and UU MAA 7474-UU MAA 7476) and an ilium (UU MAA 7118); M2: three humeri (UU MAA 7193). | ||||||||||
Amphibia
- Temnospondyli
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Latonia sp.
von Meyer 1843
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Georgalis et al. 2019 | |||||||||
M1: two fragmentary frontoparietals belonging to a single individual (UU MAA 7445 and UU MAA 7462), three premaxillae (UU MAA 7130), 66 maxillae (UU MAA 7120, UU MAA 7128, UU MAA 7131, UU MAA 7156, UU MAA 7249, UU MAA 7250, UU MAA 7458, and UU MAA 7464-UU MAA 7472), seven angulars (UU MAA 7154 and UU MAA 7529), an atlas (UU MAA 7459), 17 presacral vertebrae (UU MAA 7125, UU MAA 7158a, UU MAA 7442-UU MAA 7444, UU MAA 7452, UU MAA 7454, UU MAA 7457, UU MAA 7460, and UU MAA 7461), three sacral vertebrae (UU MAA 7455 and UU MAA 7522), four urostyles (UU MAA 7446, UU MAA 7450, and UU MAA 7521), a scapula (UU MAA 7520), four costae (UU MAA 7523), 14 humeri (UU MAA 7155, UU MAA 7158b, UU MAA 7453, and UU MAA 7463), and nine ilia (UU MAA 7447-UU MAA 7449, UU MAA 7456, and UU MAA 7528); M2: a fragmentary frontoparietal (UU MAA 7192); M3: seven maxillae (UU MAA 7223, UU MAA 7224, UU MAA 7227, UU MAA 7238, UU MAA 7239, UU MAA 7240, and UU MAA 7241), a presacral vertebra (UU MAA 7228), a humerus (UU MAA 7225), and an ilium (UU MAA 7229). | ||||||||||
Amphibia
- Temnospondyli
- Urodela
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Cryptobranchidae indet.
Cope 1889
|
Georgalis et al. 2019 | |||||||||
M1: an atlas (UU MAA 7441), probably also a second atlas (UU MAA 7530) | ||||||||||
Salamandra sp.
Laurenti 1768
|
Georgalis et al. 2019 | |||||||||
M1: 12 trunk vertebrae (UU MAA 7494- UU MAA 7505) and six caudal vertebrae (UU MAA 7117, UU MAA 7491-UU MAA 7493, UU MAA 7506, and UU MAA 7507). | ||||||||||
Salamandridae indet.
Gray 1825
|
Georgalis et al. 2019 | |||||||||
M1: an atlas (UU MAA 7490), four vertebrae (UU MAA 7153, UU MAA 7485, UU MAA 7488, and UU MAA 7243), and two ribs (UU MAA 7508 and UU MAA 7519). | ||||||||||
Salamandrina sp.
Fitzinger 1826
|
Georgalis et al. 2019 | |||||||||
M1: two vertebrae (UU MAA 7483 and UU MAA 7484) | ||||||||||
Lissotriton sp.
Bell 1839
|
Georgalis et al. 2019 | |||||||||
(Lissotriton vulgaris group) M1: two vertebrae (UU MAA 7517 and UU MAA 7518). | ||||||||||
Mammalia
- Rodentia
- Gliridae
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Myomimus maritsensis
de Bruijn et al. 1970
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70 specimens | |||||||||
approx. | ||||||||||
Myomimus dehmi
(de Bruijn 1966)
|
Schmidt-Kittler et al. 1995 | |||||||||
Glis cf. minor
Kowalski 1963
|
Schmidt-Kittler et al. 1995 | |||||||||
Muscardinus pliocaenicus
Kowalski 1963
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Rodentia
- Sciuridae
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Tamias informal sp. A
Illinger 1811
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Tamias + Protospermophilus=approx. 13 specimens | ||||||||||
Hylopetes macedoniensis
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Pliopetaurista + Hylopetes=approx. 50 specimens | ||||||||||
? Protospermophilus sp.
Gazin 1930
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Tamias + Protospermophilus=approx. 13 specimens | ||||||||||
Pliopetaurista dehneli
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Pliopetaurista + Hylopetes=approx. 50 specimens | ||||||||||
Miopetaurista thaleri
Mein 1970
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Rodentia
- Eomyidae
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Keramidomys aff. carpathicus
(Schaub and Zapfe 1953)
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probably less than 3 specimens | ||||||||||
Leptodontomys catalaunicus
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Rodentia
- Castoridae
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Castoridae indet.
Gray 1821
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probably less than 3 specimens | ||||||||||
Mammalia
- Rodentia
- Muridae
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Apodemus dominans
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A. gudrunae+A.dominans=approx. 390 specimens | ||||||||||
Apodemus gudrunae
van de Weerd 1976
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A. gudrunae+A.dominans=approx. 390 specimens | ||||||||||
Micromys informal sp. A
Dehne 1841
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18 specimens | |||||||||
approx. | ||||||||||
Occitanomys cf. neutrum
de Bruijn 1976
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40 specimens | |||||||||
approx. | ||||||||||
Mammalia
- Rodentia
- Cricetidae
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Kowalskia fahlbuschi
Bachmayer and Wilson 1970
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recombined as Neocricetodon fahlbuschi | ||||||||||
Kowalskia + Cricetus= approx. 83 specimens | ||||||||||
"Cricetus" informal sp. A
Leske 1779
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Kowalskia + Cricetus= approx. 83 specimens | ||||||||||
Mammalia
- Lagomorpha
- Leporidae
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Alilepus turolensis
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Alilepus + Prolagus = 41 specimens | ||||||||||
Mammalia
- Lagomorpha
- Ochotonidae
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Prolagus michauxi
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Alilepus + Prolagus = 41 specimens | ||||||||||
Mammalia
- Primates
- Cercopithecidae
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Mesopithecus pentelicus
Wagner 1839
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Erinaceidae
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Schizogalerix sp.
Engesser 1980
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Doukas 1989 | |||||||||
= Schizogalerix macedonica n. sp.
Doukas et al. 1995
|
Doukas et al. 1995 | |||||||||
Erinaceus sp.
Linnaeus 1758
|
Doukas 1989 | |||||||||
= Erinaceus samsonowiczi
Sulimski 1959
|
Doukas et al. 1995 | |||||||||
Erinaceus samsonowiczi
Sulimski 1959
|
Doukas 2005 | |||||||||
Mammalia
- Soricidae
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Soricidae informal sp. A
Fischer 1814
|
Doukas 1989 | |||||||||
= Asoriculus gibberodon
|
Doukas et al. 1995 | |||||||||
Petenyia sp.
Kormos 1934
|
Doukas 1989 | |||||||||
= Petenyia hungarica
Kormos 1934
|
Doukas et al. 1995 | |||||||||
Deinsdorfia kerkhoffi n. sp.
Doukas et al. 1995
|
Doukas et al. 1995 | |||||||||
Soricidae informal sp. B
Fischer 1814
|
Doukas 1989 | |||||||||
= Sorex sp.
Linnaeus 1758
|
Doukas et al. 1995 | |||||||||
Amblycoptus sp.
Kormos 1926
|
Doukas 1989 | |||||||||
= Amblycoptus jessiae n. sp.
Doukas et al. 1995
|
Doukas et al. 1995 | |||||||||
Mammalia
- Talpidae
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Desmanella sp.
Engesser 1972
|
Doukas 1989 | |||||||||
= Desmanella dubia
Rümke 1976
|
Doukas et al. 1995 | |||||||||
Talpidae informal sp. A
(Fischer de Waldheim 1817)
|
Doukas 1989 | |||||||||
= Urotrichus sp.
Temminck 1841
|
Doukas et al. 1995 | |||||||||
Dibolia dekkersi
Rümke 1985
|
Rümke 1985 | |||||||||
= Ruemkelia dekkersi
|
Doukas et al. 1995 | |||||||||
synonym of Archaeodesmana | ||||||||||
Talpidae informal sp. B
(Fischer de Waldheim 1817)
|
Doukas 1989 | |||||||||
= Talpa fossilis
Petenyi 1864
|
Doukas et al. 1995 | |||||||||
Mammalia
- Chiroptera
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Chiroptera indet.
Blumenbach 1779
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Perissodactyla
- Rhinocerotidae
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Rhinocerotidae indet.
Gray 1821
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Perissodactyla
- Equidae
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Hipparion sp.
de Christol 1832
|
Schmidt-Kittler et al. 1995 | |||||||||
2 species | ||||||||||
Mammalia
- Suidae
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Korynochoerus palaeochoerus
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Cervidae
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Pliocervus graecus n. sp.
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Bovidae
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Gazella sp.
Blaineville 1816
|
Schmidt-Kittler et al. 1995 | |||||||||
Boselaphini indet.
Knottnerus-Meyer 1997
|
Schmidt-Kittler et al. 1995 | |||||||||
Tragoportax gaudryi
Kretzoi 1941
|
Schmidt-Kittler et al. 1995 | |||||||||
Tragoportax cf. amalthea
(Roth and Wagner 1854)
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Giraffidae
|
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Samotherium cf. boissieri
Forsyth Major 1888
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Carnivora
- Hyaenidae
|
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Chasmaporthetes sp.
Hay 1921
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Carnivora
- Viverridae
|
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Viverridae indet.
Gray 1821
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Carnivora
- Mephitidae
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Promephitis sp.
Gaudry 1861
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Carnivora
- Mustelidae
|
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Martes lefkonensis n. sp.
|
Schmidt-Kittler et al. 1995 | |||||||||
Lutra affinis
Gervais 1859
|
Schmidt-Kittler et al. 1995 | |||||||||
Promeles macedonicus
|
Schmidt-Kittler et al. 1995 | |||||||||
Mammalia
- Proboscidea
- Choerolophodontidae
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Choerolophodon pentelicus
|
Schmidt-Kittler et al. 1995 | |||||||||
Reptilia
- Testudines
|
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Testudines indet.
Batsch 1788
|
Georgalis et al. 2019 | |||||||||
M1: an incomplete unidentified neural (UU MAA 7136), three unidentified peripherals (UU MAA 7137, UU MAA 7140, and UU MAA 7142), and several shell fragments (UU MAA 7138, UU MAA 7139, UU MAA 7147 [five fragments]); M3: an unidentified peripheral (UU MAA 7254), a shell fragment (UU MAA 7252), and several fragments of different elements (one caudal vertebra, one pelvic fragment, three neurals, four costals, one peripheral, and 24 shell fragments) (UU MAA 7255). | ||||||||||
Reptilia
- Testudines
- Testudinidae
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? Testudinidae indet.
Batsch 1788
|
Georgalis et al. 2019 | |||||||||
M1: an isolated ?limb osteoderm (UU MAA 7135). | ||||||||||
Testudinata indet.
(Oppel 1811)
|
Schmidt-Kittler et al. 1995 | |||||||||
= Testudo sp.
Linnaeus 1758
|
Vlachos 2015 | |||||||||
isolated plates | ||||||||||
Reptilia
- Testudines
- Geoemydidae
|
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Mauremys aristotelica
Vlachos et al. 2019
|
Georgalis et al. 2019 | |||||||||
M1: three incomplete unidentified neurals (UU MAA 7143, UU MAA 7144, and UU MAA 7146); M3: a nuchal (UU MAA 7283), a fragmentary unidentified neural (UU MAA 7253), a left costal VI (UU MAA 7284), unidentified costals (UU MAA 7286 and UU MAA 7288), a left peripheral III (UU MAA 7289), a right peripheral III (UU MAA 7290), unidentified peripherals (UU MAA 7285, UU MAA 7287, UU MAA 7291, UU MAA 7292, UU MAA 7293, UU MAA 7295, and UU MAA 7297), and a right hyoplastron (UU MAA 7296). | ||||||||||
Reptilia
- Squamata
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Squamata indet.
Oppel 1811
|
Georgalis et al. 2019 | |||||||||
M1: a premaxilla (UU MAA 7093), eight maxillae (UU MAA 7342, UU MAA 7355, UU MAA 7356, UU MAA 7360, and UU MAA 7428), six dentaries (UU MAA 7103, UU MAA 7105, and UU MAA 7427), 14 fragments of tooth bearing bones (UU MAA 7102, UU MAA 7311, UU MAA 7354, UU MAA 7393, UU MAA 7405, and UU MAA 7406), three axes (UU MAA 7346 and UU MAA 7378), three post-axial cervical vertebrae (UU MAA 7322, UU MAA 7348, and UU MAA 7789), 13 trunk vertebrae (UU MAA 7110, UU MAA 7345, UU MAA 7382-UU MAA 7384, UU MAA 7421), two cloacal vertebrae (UU MAA 7380 and UU MAA 7381), eight caudal vertebrae (UU MAA 7347, UU MAA 7385-UU MAA 7387, and UU MAA 7390), a fragment of caudal vertebra (UU MAA 7310), 11 ribs (UU MAA 7111 and UU MAA 7419), seven humeri (UU MAA 7309 and UU MAA 7352), two pelvic girdles (UU MAA 7349 and UU MAA 7376), 14 femurs (UU MAA 7109, UU MAA 7308, UU MAA 7351, UU MAA 7377, and UU MAA 7420), and a tibia (UU MAA 7350); M2: a fragment of a presacral vertebra (UU MAA 7299) and a caudal vertebra (UU MAA 7300); M3: a fragment of tooth bearing bone (UU MAA 7339) and a caudal vertebra (UU MAA 7221). | ||||||||||
Reptilia
|
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Serpentes indet.
(Linnaeus 1758)
|
Szyndlar 1995 | 6 elements | ||||||||
6 precaudal and caudal vertebrae (AMPG MA 190 to 195), "may have belonged either to Naja or Vipera" | ||||||||||
Scolecophidia indet.
(Duméril and Bibron 1844)
|
Georgalis et al. 2019 | |||||||||
M1: three trunk vertebrae (UU MAA 7373-UU MAA 7375). | ||||||||||
Reptilia
- Elapidae
|
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Naja sp.
Laurenti 1768
|
Szyndlar 1995 | 25 elements | ||||||||
25 fragmentary precaudal vertebrae (AMPG MA 163 to 187) | ||||||||||
Reptilia
- Crotalidae
|
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Vipera sp.
Laurenti 1768
|
Szyndlar 1995 | 2 elements | ||||||||
"oriental viper", 2 fragmentary precaudal vertebrae (AMPG MA 188, 189) | ||||||||||
Reptilia
- Colubridae
|
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Colubrinae indet.
Oppel 1811
|
Szyndlar 1995 | 36 elements | ||||||||
36 fragmentary precaudal vertebrae (AMPG MA 101 to 136) | ||||||||||
Paraxenophis spanios n. gen., n. sp.
Georgalis et al. 2019
|
Georgalis et al. 2019 | |||||||||
Holotype. Specimen UU MAA 7645 (a mid-trunk vertebra) (Figure 27). The holotype originates from M1 site. Paratypes. Specimen UU MAA 7536 (a posterior trunk vertebra) (Figure 29.1-5); UU MAA 7533 (a posteriormost trunk vertebra) (Figure 29.6-8). Both paratypes originate from M1 site. | ||||||||||
Natricinae indet.
Bonaparte 1838
|
Szyndlar 1995 | 26 elements | ||||||||
26 fragmentary precaudal vertebrae (AMPG MA 137 to 162) | ||||||||||
Periergophis micros n. gen., n. sp.
Georgalis et al. 2019
|
Georgalis et al. 2019 | |||||||||
Holotype. Specimen UU MAA 7615 (a posterior trunk vertebra) (Figures 22 and 24.1-2). The holotype originates from M1 site. Paratypes. Specimen UU MAA 7551 (a cervical vertebra) (Figure 23.1-2); UU MAA 7614 (a midtrunk vertebra) (Figure 23.3-8); UU MAA 7618 (a cloacal vertebra) (Figure 23.9-12); UU MAA 7636 (a caudal vertebra) (Figures 23.13-16, 24.3). All paratypes originate from M1 site. | ||||||||||
Natrix aff. rudabanyaensis
|
Georgalis et al. 2019 | |||||||||
M1: 34 precloacal vertebrae (UU MAA 7656-UU MAA 7689) and seven caudal vertebrae (UU MAA 7690-UU MAA 7694, UU MAA 7893, and UU MAA 7904); M3: four precloacal vertebrae (UU MAA 7741-UU MAA 7744). | ||||||||||
Natrix sp.
Laurenti 1768
|
Georgalis et al. 2019 | |||||||||
M1: six precloacal vertebrae (UU MAA 7695-UU MAA 7700). | ||||||||||
Reptilia
- Lacertidae
|
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Lacertidae indet.
Bonaparte 1831
|
Georgalis et al. 2019 | |||||||||
M1: three maxillae (UU MAA 7031, UU MAA 7032, and UU MAA 7438), a quadrate (UU MAA 7096), three pterygoids (UU MAA 7097-UU MAA 7099), 16 dentaries (UU MAA 7048-UU MAA 7061, UU MAA 7095, and UU MAA 7424), 10 fragments of tooth bearing bones (UU MAA 7087-UU MAA 7092, UU MAA 7094, UU MAA 7391, and UU MAA 7802), and an isolated tooth (UU MAA 7392); M3: a dentary (UU MAA 7220). | ||||||||||
Reptilia
|
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Laterata indet.
Vidal and Hedges 2005
|
Georgalis et al. 2019 | |||||||||
M1: 29 dentaries (UU MAA 7026, UU MAA 7106, UU MAA 7357, UU MAA 7361, UU MAA 7379, UU MAA 7402-UU MAA 7404, UU MAA 7435, and UU MAA 7436); M3: a dentary (UU MAA 7206). | ||||||||||
Reptilia
- Scincidae
|
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? Scincidae indet.
Gray 1825
|
Georgalis et al. 2019 | |||||||||
M1: two dentaries (UU MAA 7027 and UU MAA 7107). | ||||||||||
Reptilia
- Cordylidae
|
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aff. Palaeocordylus sp.
Roček 1984
|
Georgalis et al. 2019 | |||||||||
M1: 26 maxillae (UU MAA 7046, UU MAA 7047, UU MAA 7077-UU MAA 7086, UU MAA 7101, UU MAA 7273, UU MAA 7358, UU MAA 7362-UU MAA 7365, UU MAA 7370-UU MAA 7372, UU MAA 7395, UU MAA 7396, and UU MAA 7437), 57 dentaries (UU MAA 7001-UU MAA 7025, UU MAA 7034-UU MAA 7036, UU MAA 7039, UU MAA 7040, UU MAA 7104, UU MAA 7108, UU MAA 7266, UU MAA 7267, UU MAA 7270, UU MAA 7279, UU MAA 7359, UU MAA 7366-UU MAA 7369, UU MAA 7397-UU MAA 7401, UU MAA 7425, and UU MAA 7426), and six fragments of tooth bearing bones (UU MAA 7037 and UU MAA 7038); M3: a maxilla (UU MAA 7340), eight dentaries (UU MAA 7199-UU MAA 7205, and UU MAA 7209), and a fragment of tooth bearing bone (UU MAA 7210). | ||||||||||
Reptilia
- Agamidae
|
||||||||||
Agaminae indet.
|
Georgalis et al. 2019 | |||||||||
M1: a premaxilla (UU MAA 7076), three maxillae (UU MAA 7043-UU MAA 7045); five dentaries (UU MAA 7041, UU MAA 7042, UU MAA 7075, UU MAA 7276, and UU MAA 7305); 15 fragments of tooth bearing bones (UU MAA 7063, UU MAA 7065-UU MAA 7074, UU MAA 7264, UU MAA 7303, UU MAA 7304, and UU MAA 7394); five caudal vertebrae (UU MAA 7062, UU MAA 7064, and UU MAA 7302); M2: a frontal (UU MAA 7191); M3: a premaxilla (UU MAA 7341), a maxilla (UU MAA 7208), and a left dentary (UU MAA 7207). | ||||||||||
Reptilia
|
||||||||||
Anguimorpha indet.
Fürbringer 1900
|
Georgalis et al. 2019 | |||||||||
M1: a dentary (UU MAA 7353). | ||||||||||
Reptilia
- Varanidae
|
||||||||||
Varanus sp.
Merrem 1820
|
Georgalis et al. 2019 | |||||||||
M1: six teeth (UU MAA 7029, UU MAA 7030, and UU MAA 7148-UU MAA 7151) and 12 caudal vertebrae (UU MAA 7028, UU MAA 7278, UU MAA 7388, UU MAA 7389, UU MAA 7429-UU MAA 7434, UU MAA 7805, and UU MAA 7806); M3: a caudal vertebra (UU MAA 7198). | ||||||||||
Reptilia
- Anguidae
|
||||||||||
Anguinae indet.
Gray 1825
|
Georgalis et al. 2019 | |||||||||
M1: 188 osteoderms (UU MAA 7160-UU MAA 7170, UU MAA 7262, UU MAA 7265, UU MAA 7268, UU MAA 7271, UU MAA 7275, UU MAA 7306, and UU MAA 7314-UU MAA 7318); M2: eight osteoderms (UU MAA 7187, UU MAA 7188, and UU MAA 7298). | ||||||||||
Ophisaurus sp.
Daudin 1803
|
Georgalis et al. 2019 | |||||||||
M1: a maxilla (UU MAA 7324) and two dentaries (UU MAA 7176 and UU MAA 7177). | ||||||||||
Anguis sp.
Linnaeus 1758
|
Georgalis et al. 2019 | |||||||||
M1: 37 trunk vertebrae (UU MAA 7181, UU MAA 7263, UU MAA 7274, UU MAA 7280, UU MAA 7332-UU MAA 7334, UU MAA 7411, UU MAA 7418, and UU MAA 7804), a cloacal vertebra (UU MAA 7269); M3: two trunk vertebrae (UU MAA 7217 and UU MAA 7218). | ||||||||||
Pseudopus sp.
Merrem 1820
|
Georgalis et al. 2019 | |||||||||
M1: a maxilla (UU MAA 7171), a dentary (UU MAA 7407), a fragment of tooth bearing bone (UU MAA 7307), two trunk vertebrae (UU MAA 7172 and UU MAA 7328), and a cloacal vertebra (UU MAA 7329); M3: three trunk vertebrae (UU MAA 7212-UU MAA 7214). | ||||||||||
Pisces
|
||||||||||
Pisces indet.
Linnaeus 1758
|
Schmidt-Kittler et al. 1995 | |||||||||
synonym of Osteichthyes | ||||||||||
see common names |
Geography
Country: | Greece | State/province: | Central Macedonia | County: | Serres |
Coordinates: | 41.2° North, 23.5° East (view map) | ||||
Paleocoordinates: | 40.9° North, 23.2° East | ||||
Basis of coordinate: | estimated from map | ||||
Geographic resolution: | small collection |
Time
Period: | Neogene | Epoch: | Miocene |
10 m.y. bin: | Cenozoic 6 | ||
*Period: | Tertiary | *Local age/stage: | Middle Turolian - Late/Upper Turolian |
Key time interval: | Turolian | ||
Age range of interval: | 8.70000 - 5.33300 m.y. ago | ||
* legacy (obsolete) database fields |
Stratigraphy
Formation: | Lefkon | ||||
Stratigraphy comments: may be the same level as Ano Metochi 2-3 |
Lithology and environment
Primary lithology: | conglomerate |
Secondary lithology: | sandy marl |
Includes fossils? | Y |
Lithology description: A polymict poorly sorted conglomerate containing pebbles of schist, gneiss and granite. This coarse-grained series grades upwards into arkosic sands and finally into clays with occasional lignites. Within the top part of the formation, two "landslide" breccias occur. The lower of these consists of metamorphic rocks, the upper one of granite. | |
Environment: | alluvial fan |
Taphonomy
Modes of preservation: | body |
Size of fossils: | mesofossils |
Collection methods and comments
Collection methods: | field collection |
Reason for describing collection: | taxonomic analysis |
Museum repositories: | AMPG |
Metadata
Database number: | 32189 | ||
Authorizer: | J. Alroy, J. Mueller, E. Vlachos, L. van den Hoek Ostende | Enterer: | E. Leckey, E. Vlachos, T. Liebrecht, L. van den Hoek Ostende |
Modifier: | T. Liebrecht | Research group: | vertebrate |
Created: | 2003-06-09 16:30:14 | Last modified: | 2012-11-25 17:53:22 |
Access level: | the public | Released: | 2003-06-09 16:30:14 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
8408. | H. de Bruijn. 1989. Smaller mammals from the Upper Miocene and Lower Pliocene of the Strimon basin, Greece. Part 1. Rodentia and Lagomorpha. Bollettino della Societa Paleontologica Italiana 28(2-3):189-195 [J. Alroy/E. Leckey/E. Leckey] |
Secondary references:
8409 | 5% 15420 | C. S. Doukas. 1989. Smaller mammals from the Upper Miocene and Lower Pliocene of the Strimon basin, Greece. Part 2. Insectivora. Bollettino della Societa Palaeontologica Italiana 28(2-3):213-216 [J. Alroy/E. Leckey/E. Leckey] |
73905 | C. S. Doukas. 2005. Greece. In L. W. van den Hoek Ostende, C. S. Doukas, J. W. F. Reumer (eds.), The Fossil Record of the Eurasian Neogene Insectivores (Erinaceomorpha, Soricomorpha, Mammalia), Part I. Scripta Geologica Special Issue 5:99-112 [E. Vlachos/E. Vlachos] | |
19740 | C. S. Doukas, L. W. van den Hoek Ostende, C. D. Theocharopoulos and J. W. F. Reumer. 1995. The Vertebrate Locality Maramena (Macedonia, Greece) at the Turolian-Ruscinian Boundary (Neogene). 5. Insectivora (Erinaceidae, Talpidae, Soricidae, Mammalia). Münchner Geowissenschaftliche Abhandlungen (A) 28:43-64 [L. van den Hoek Ostende/L. van den Hoek Ostende/T. Liebrecht] | |
70786 | G. L. Georgalis, A. Villa, M. Ivanov, D. Vasilyan, and M. Delfino. 2019. Fossil amphibians and reptiles from the Neogene locality of Maramena (Greece), the most diverse European herpetofauna at the Miocene/Pliocene transition boundary. Palaeontologia Electronica 22.3.68:1-99 [E. Vlachos/E. Vlachos] | |
23416 | C. G. Rümke. 1985. A review of fossil and recent Desmaninae (Talpidae, Insectivora). Utrecht Micropaleontological Bulletins 4:1-241 [L. van den Hoek Ostende/L. van den Hoek Ostende] | |
61128 | N. Schmidt-Kittler, H. de Bruijn, and C. Doukas. 1995. The vertebrate locality Maramena (Macedonia, Greece) at the Turolian-Ruscinian Boundary (Neogene). 1. General Introduction. Munchner Goewiss. Abh 28:9-18 [E. Vlachos/E. Vlachos] | |
43602 | Z. Szyndlar. 1995. The Vertebrate Locality Maramena (Macedonia, Greece) at the Turolian-Ruscinian Boundary (Neogene). 4. Serpentes (Squamata, Reptilia). Münchner Geowissenschaftliche Abhandlungen (A) 28:39-42 [J. Mueller/T. Liebrecht] | |
65163 | E. Vlachos. 2015. The Fossil Chelonians of Greece. Systematics - Evolution - Stratigraphy - Palaeoecology. Scientific Annals of the School of Geology, Aristotle University of Thessaloniki, Greece 173:1-479 [E. Vlachos/E. Vlachos/E. Vlachos] |