Csarnota locality 2: Ruscinian - Villanyian, Hungary
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
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Megalobatrachidae indet.
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2 specimens | |||||||||
Procamptoceros cf. brivatense
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Myriapoda
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Diplopoda indet.
Gervais 1844
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Gastropoda
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Gastropoda indet.
Cuvier 1795
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Reptilia
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Aves indet.
Linnaeus 1758
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Reptilia
- Phasianidae
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Francolinus capeki
Lambrecht 1933
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sporadic. Francolinus capeki wezensis | ||||||||||
Tetrao macropus
Jánossy 1967
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few | ||||||||||
Reptilia
- Testudines
- Testudinidae
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Testudo cf. lambrechti
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rare | ||||||||||
Reptilia
- Varanidae
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Varanus deserticolus
Bolkay 1913
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few | ||||||||||
= Varanus sp.
Merrem 1820
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Villa et al. 2018 | |||||||||
Reptilia
- Anguidae
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Anguis sp.
Linnaeus 1758
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several thousand | ||||||||||
Ophisaurus intermedius
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common | ||||||||||
Reptilia
- Ophidia
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Ophidia indet.
Latreille 1804
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abundant | ||||||||||
Reptilia
- Colubridae
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Elaphe quatuorlineata
(Lacépède 1789)
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Venczel and Vardai 2000 | 1 element | ||||||||
a single vertebra (GMH uncatalogued) | ||||||||||
Reptilia
- Lacertidae
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Lacerta ruscinensis
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several thousand | ||||||||||
Lacerta aff. agilis
Linneaus 1758
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few | ||||||||||
Mammalia
- Chiroptera
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Chiroptera indet.
Blumenbach 1779
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common | ||||||||||
Mammalia
- Carnivora
- Felidae
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Felis sp.
Linnaeus 1758
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1 specimen | |||||||||
Felis s. 1. indet. (size of catus) | ||||||||||
? Lynx sp.
Kerr 1792
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Lynx (?) sp. indet. | ||||||||||
Mammalia
- Carnivora
- Mustelidae
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Mustela aff. palerminea
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few. | ||||||||||
Baranogale beremendensis
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few. | ||||||||||
Xenictis pilgrimi
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few. | ||||||||||
Mammalia
- Carnivora
- Ursidae
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Ursus sp.
Linnaeus 1758
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few. Ursus s. 1. sp. indet. | ||||||||||
Mammalia
- Carnivora
- Canidae
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Canidae indet.
Fischer de Waldheim 1817
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few. Canidae I-III indet. | ||||||||||
Mammalia
- Cervidae
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? Cervus pyrenaicus
(Mennecart et al. 2017)
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recombined as Croizetoceros pyrenaicus | ||||||||||
Small Cervidae (Cervus pyrenaicus Deperet?) | ||||||||||
Cervus (? Rusa) sp.
(Smith 1827)
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original and current combination Rusa | ||||||||||
Cervus (?Rusa) sp. indet. | ||||||||||
Megaceros cf. dupuisi
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synonym of Megaloceros | ||||||||||
(unlikely according to Kretzoi) | ||||||||||
"Alces" sp.
Gray 1821
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Alces sp. indet. | ||||||||||
Mammalia
- Bovidae
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Gazellospira cf. torticornis
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Hemitragus cf. bonali
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Mammalia
- Soricidae
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Episoriculus gibberodon
Petenyi 1864
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100 specimens | |||||||||
c. 100 | ||||||||||
Blarinoides mariae
Sulimski 1959
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few | ||||||||||
Sorex sp.
Linnaeus 1758
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Sorex s. 1. sp. | ||||||||||
Petenyia hungarica
Kormos 1934
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100 specimens | |||||||||
c. 100 | ||||||||||
Petenyiella gracilis
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Beremendia fissidens
(Petényi 1864)
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400 specimens | |||||||||
c. 400 | ||||||||||
Mammalia
- Talpidae
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Talpa csarnotana
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100 specimens | |||||||||
c. 100 | ||||||||||
Talpa cf. fossilis
Petenyi 1864
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rare | ||||||||||
Desmana sp.
Guldenstaedt 1777
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Desmana sp. indet. I-II, rare | ||||||||||
Mammalia
- Erinaceidae
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Hemiechinus cf. samsonowiczi
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few | ||||||||||
Mammalia
- Rodentia
- Sciuridae
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Pliopetes hungaricus
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few | ||||||||||
? Eutamias sp.
Trouessart 1880
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Eutamias sp. indet. few | ||||||||||
Pliopetaurista pliocaenica
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2 specimens | |||||||||
Sciurus sp.
Linnaeus 1758
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1 specimen | |||||||||
Sciurus sp. indet. | ||||||||||
Mammalia
- Rodentia
- Muridae
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Apodemus dominans
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8000 specimens | |||||||||
c. 8000 | ||||||||||
Micromys praeminutus
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few | ||||||||||
Rhagapodemus frequens
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common | ||||||||||
Mammalia
- Rodentia
- Cricetidae
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Cricetinus europaeus
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few | ||||||||||
Cseria gracilis
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synonym of Mimomys | ||||||||||
common | ||||||||||
Propliomys hungaricus
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abundant | ||||||||||
Prospalax priscus
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600 specimens | |||||||||
c. 600 | ||||||||||
Baranomys loczyi
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few | ||||||||||
Dolomys nehringi
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common | ||||||||||
Promimomys cor
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1 specimen | |||||||||
Mammalia
- Rodentia
- Gliridae
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Dryomimus eliomyoides
Kretzoi 1959
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1000 specimens | |||||||||
c. 1000 | ||||||||||
Glirurus gemmula
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1 specimen | |||||||||
Muscardinus sp.
Kaup 1829
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few | ||||||||||
Glis minor
Kowalski 1963
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few | ||||||||||
Mammalia
- Lagomorpha
- Ochotonidae
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Ochotonoides csarnotanus
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1-2 specimens | ||||||||||
Mammalia
- Lagomorpha
- Leporidae
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Hypolagus cf. beremendensis
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few. | ||||||||||
Mammalia
- Primates
- Cercopithecidae
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Macaca sp.
Lacépède 1799
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Macaca sp. indet. sporadic | ||||||||||
cf. Dolichopithecus sp.
Depéret 1889
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cf. Dolichopithecus sp. indet. | ||||||||||
Amphibia
- Temnospondyli
- Albanerpetontidae
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Albanerpeton sp.
Estes and Hoffstetter 1976
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Venczel and Gardner 2003 | |||||||||
"new species" | ||||||||||
= Albanerpeton pannonicus n. sp.
Venczel and Gardner 2005
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Venczel and Gardner 2005 | |||||||||
corrected as Albanerpeton pannonicum | ||||||||||
MAFI V.22000 - holotype (fused pair of nearly complete premaxillae, articulated with anterior end of right maxilla); referred material: fragmentary skulls (n = 24), each consisting of either a maxillary arcade comprising a maxilla articulated with some combination of a jugal, lacrimal, prefrontal and palatal bones or an incomplete skull roof comprising fused frontals articulated with some combination of a prefrontal, lacrimal and nasal: MAFI V.22003 ⁄ 1, V.22011, V.22012 ⁄ 1–3, V.22015, V.22021 ⁄ 1–2, V.22033 ⁄ 2, V.22034, V.22039, V.22053, V.22055, V.22075, V.22076 ⁄ 1–2, V.22077, V.22093 ⁄ 1–3, V.22094 ⁄ 1, V.22100 ⁄ 1–2, V.22111; fused premaxillae (n = 6): MAFI V.22017 ⁄ 1–2, V.22023, V.22026 ⁄ 1, V.22041 ⁄ 1, V.22072 ⁄ 1; isolated premaxillae (n = 38): MAFI V.22006 ⁄ 1–5, V.22007, V.22018 ⁄ 1–3, V.22026 ⁄ 2, V.22030 ⁄ 1–2, V.22036 ⁄ 1–4, V.22041 ⁄ 2, V.22050, V.22057, V.22067, V.22072 ⁄ 2–3, V.22078 ⁄ 1–2, V.22083, V.22089 ⁄ 1–3, V.22095, V.22101 ⁄ 1–2, V.22105, V.22106 ⁄ 1–3, V.22113 ⁄ 1–3; isolated maxillae (n = 50): MAFI V.22002 ⁄ 1–6, V.22013 ⁄ 1–10, V.22014 ⁄ 1–2, V.22028 ⁄ 1, V.22038 ⁄ 1–3, V.22043 ⁄ 1–2, V.22048, V.22056 ⁄ 5, V.22070 ⁄ 1–4, V.22087 ⁄ 1–2, V.22092 ⁄ 1–4, V.22099 ⁄ 1–4, V.22110 ⁄ 1–10; fused frontals, not articulated with other bones (n = 39): MAFI V.22003 ⁄ 2–3; V.22004 ⁄ 1–3, V.22005 ⁄ 1–4, V.22016 ⁄ 1–7, V.22022, V.22025, V.22029, V.22035, V.22040, V.22045, V.22060, V.22063, V.22066 ⁄ 1–2, V.22071, V.22081, V.22082 ⁄ 1–2, V.22088, V.22094 ⁄ 2, V.22104 ⁄ 1–5, V.22112 ⁄ 1–2; mandibles (n = 20): MAFI V.22009, V.22010 ⁄ 1–4, V.22020 ⁄ 1–4, V.22032 ⁄ 1–2, V.22047, V.22052, V.22059, V.22062, V.22065, V.22080, V.22097, V.22108 ⁄ 1–2; dentaries (n = 302): MAFI V.22001 ⁄ 1– 18, V.22008 ⁄ 1–38, V.22019 ⁄ 1–37, V.22024 ⁄ 1–2, V.22027 ⁄ 1–8, V.22031 ⁄ 1–10, V.22037 ⁄ 1–24, V.22042 ⁄ 1–3, V.22044 ⁄ 1–6, V.22046 ⁄ 1–10, V.22049, V.22051 ⁄ 1–4, V.22056 ⁄ 1–4, V.22058 ⁄ 1– 3, V.22061 ⁄ 1–10, V.22064 ⁄ 1–3, V.22068 ⁄ 1–12, V.22073 ⁄ 1–4, V.22079, V.22084, V.22085 ⁄ 1–11, V.22090 ⁄ 1–17, V.22096 ⁄ 1–23, V.22102 ⁄ 1–11, V.22107 ⁄ 1–25, V.22114 ⁄ 1–16; presacral vertebrae (n = 10): MAFI V.22028 ⁄ 2, V.22054, V.22074 ⁄ 1, V.22086 ⁄ 1–2, V.22098 ⁄ 1, V.22103 ⁄ 1, V.22109 ⁄ 1–2, V.22114 ⁄ 17; humeri (n = 16): MAFI V.22033 ⁄ 1, V.22069, V.22074 ⁄ 2, V.22086 ⁄ 3, V.22091 ⁄ 1–5, V.22098 ⁄ 2–4, V.22103 ⁄ 2–3, V.22109 ⁄ 3–4. | ||||||||||
Amphibia
- Temnospondyli
- Urodela
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Salamandra sp.
Laurenti 1768
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rare | ||||||||||
Triturus sp.
Rafinesque 1815
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sporadic | ||||||||||
Amphibia
- Temnospondyli
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Anura indet.
(Fischer von Waldheim 1813)
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(minly Bufo (listed seperately) c. 20,000 specimens in total) | ||||||||||
Amphibia
- Temnospondyli
- Bufonidae
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Bufo sp.
Laurenti 1768
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Pisces
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Pisces indet.
Linnaeus 1758
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synonym of Osteichthyes | ||||||||||
see common names |
Geography
Country: | Hungary | State/province: | Barany |
Coordinates: | 45.9° North, 18.2° East (view map) | ||
Paleocoordinates: | 45.8° North, 18.1° East | ||
Basis of coordinate: | based on nearby landmark |
Time
Period: | Neogene | Epoch: | Pliocene |
10 m.y. bin: | Cenozoic 6 | ||
Key time interval: | Ruscinian - Villanyian | ||
Age range of interval: | 5.33300 - 2.58800 m.y. ago |
Stratigraphy
Stratigraphy comments: Csarnotian stage, Mid. Pliocene, MN15 (according to Fejfar et al., 1997). Janossy puts it slightly later (in the Villanyian). Villafranchian. |
Lithology and environment
Primary lithology: | not reported |
Lithology description: 'red clay.' | |
Environment: | karst indet. |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils,mesofossils |
Collection methods and comments
Collection methods: | bulk,field collection |
Reason for describing collection: | biostratigraphic analysis |
Collection method comments: This is a reconstructed faunal list, including several collections excavated from 1910 onwards. The original site lists (1910-1930s) do not give exact locality info., but the later excavations uncovered 25 strata in a trench 1m x 1.5m and 3m deep. | |
Taxonomic list comments:'Two Pliogallus species described by Galliard have been deleted from the list.' |
Metadata
Also known as: | Csarnóta 2 | ||
Database number: | 35354 | ||
Authorizer: | A. Turner, M. Carrano, J. Head, P. Mannion, J. Mueller | Enterer: | H. O'Regan, G. Varnham, M. Carrano, T. Liebrecht, P. Mannion |
Modifier: | M. Carrano | Research group: | PACED |
Created: | 2003-12-05 04:26:08 | Last modified: | 2021-07-14 11:11:08 |
Access level: | the public | Released: | 2003-12-05 04:26:08 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
9534. | PACED | D. Janossy. 1986. Pleistocene vertebrate faunas of Hungary. Developments in Palaeontology and Stratigraphy, 8. Elsevier, Amsterdam 1-208 [A. Turner/H. O'Regan/H. O'Regan] |
Secondary references:
77386 | J. D. Gardner, A. Villa, S. Colombero, M. Venczel, and M. Delfino. 2021. A Messinian (latest Miocene) occurrence for Albanerpeton Estes & Hoffstetter, 1976 (Lissamphibia: Albanerpetontidae) at Moncucco Torinese, Piedmont Basin, northwestern Italy, and a review of the European Cenozoic record for albanerpetontids. Geodiversitas 43(14):391-404 [M. Carrano/M. Carrano] | |
23463 | ETE | M. Venczel and J. Gardner. 2003. The geologically youngest albanerpetontid amphibian. Journal of Vertebrate Paleontology 23(3, suppl.):106A [M. Carrano/M. Carrano] |
44192 | M. Venczel and J. D. Gardner. 2005. The geologically youngest albanerpetontid amphibian, from the Lower Pliocene of Hungary. Palaeontology 48(6):1273-1300 [P. Mannion/P. Mannion] | |
54746 | M. Venczel and G. Vardai. 2000. The Genus Elaphe in the Carpathian Basin: Fossil Record. Nymphaea – Folia Naturae Bihariae 28:65-82 [J. Mueller/T. Liebrecht] | |
69946 | A. Villa, J. Abella, D. M. Alba, S. Almecija, A. Bolet, G. D. Koufos, F. Knoll, A. H. Lujan, J. Morales, J. M. Robles, I. M. Sanchez and M. Delfino. 2018. Revision of Varanus marathonensis (Squamata, Varanidae) based on historical and new material: morphology, systematics, and paleobiogeography of the European monitor lizards. PLoS ONE 13(12):1-46 [E. Vlachos/E. Vlachos] |