PBDB Collection

UCMP V6168, Avenay quarry: Ypresian, France

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Mammalia - Leptictidae

	cf. Didelphodus sp. Cope 1882	Rich 1971
	MNHN Av 4917
Mammalia - Apatemyidae

	Heterohyus informal sp. I Gervais 1848		4 specimens

	Heterohyus informal sp. II Gervais 1848		1 specimen

	Heterohyus informal sp. III Gervais 1848		1 specimen

	Apatemys mutiniacus n. sp. Russell et al. 1979

	Apatemys sigogneaui n. sp. Russell et al. 1979		1 specimen
Mammalia - Esthonychidae

	Franchaius minimus Baudry 1992	Hooker 2010
	recombined as Plesiesthonyx minimus
Mammalia - Sarcodontidae

	cf. Hyracolestes sp. Matthew and Granger 1925	Rich 1971
	MNHN Av 4614
Mammalia - Condylarthra - Paroxyclaenidae

	cf. Paroxyclaenidae indet. Weitzel	Rich 1971

	cf. Paroxyclaenus sp. Teilhard de Chardin 1922	Rich 1971
	MNHN Av 5714, 5909
Mammalia - Macroscelidea - Amphilemuridae

	Macrocranion cf. nitens (Matthew 1918)	Hooker 2010
	Russell et al. 1975
Mammalia - Chiroptera - Icaronycteridae

	Icaronycteris menui Russell et al. 1973	Russell et al. 1973
Mammalia - Chiroptera - Natalidae

	cf. Ageina sp. Russell et al. 1973	Russell et al. 1973
Mammalia - Chiroptera - Archaeonycteridae

	Archaeonycteris brailloni n. sp. Russell et al. 1973	Russell et al. 1973
	type locality
Mammalia - Diacodexeidae

	Bunophorus cappettai	Sudre and Erfurt 1996

	Diacodexis varleti Sudre 1983	Sudre and Erfurt 1996
Mammalia - Choeropotamidae

	Cuisitherium sp. Sudre et al. 1983	Sudre and Erfurt 1996
Mammalia - Viverravidae

	cf. Viverravus sp. Marsh 1872	Rich 1971
	MNHN Av 4916
Mammalia - Carnivora - Miacidae

	Miacidae indet. Cope 1880	Rich 1971
	MNHN Av 5895
Mammalia

	Uintacyon hookeri Solé 2014	Solé 2014	10 specimens
	MNHN.F.Av5005, Av5006, Av5734, Av5817, Av4969, Av5820, Av4934, Av229BN, Av4756, Av15820
Mammalia - Creodonta

	Morlodon vellerei Solé 2013	Solé 2013
	MNHN Av 4785, 5917, (50)4711, 290, 5012, 467, 5070 and 5773
Mammalia - Creodonta - Hyaenodontidae

	cf. Preregidens langebadrae Solé et al. 2015	Solé et al. 2015
	MNHN.F.Av5773; Rich (1971) referred the specimen to ‘cf. Deltatheridia or Carnivora’
Mammalia - Nyctitheriidae

	Placentidens lotus Russell et al. 1973	Russell et al. 1973
Mammalia - Primates - Plesiadapidae

	Platychoerops daubrei (Lemoine 1880)	Gingerich 1976
	specimens are in the MNHN and/or the Louis private collection
Mammalia - Primates - Paromomyidae

	Phenacolemur lapparenti n. sp. Russell et al. 1967	López-Torres and Silcox 2018
	MNHN AV 5849 type

	Arcius fuscus (Russell et al. 1967)	López-Torres and Silcox 2018
Mammalia - Primates - Notharctidae

	Cantius savagei Gingerich 1977	Hooker 2010
Mammalia - Primates - Toliapinidae

	Avenius amatorum n. gen., n. sp. Russell et al. 1992	Russell et al. 1992
	MNHN AV-1093-Ph - holotype

	Toliapina vinealis n. gen., n. sp. Hooker et al. 1999	Hooker et al. 1999
	MNHN AV-700-Ph - holotype
Reptilia - Boidae

	Calamagras gallicus Rage 1977	Rage 1977
	Rage (1977) mentions a total of 30 referred vertebrae which come from Avenay and other localities in the Paris Basin, but does not give details on specimen and catalogue numbers for individual localities
Reptilia - Tropidophiidae

	Dunnophis matronensis Rage 1973	Rage 1973
	Rage (1973) mentions "nombreuses vertèbres" which come from Avenay and other localities in the Paris Basin, but does not give details on specimen and catalogue numbers for individual localities
Reptilia - Iguanidae

	Geiseltaliellus louisi Auge 1990	Augé 1990	1 specimen
	synonym of Geiseltaliellus longicaudus
	1 right dentary (n AV 16462)
Reptilia - Agamidae

	Tinosaurus sp. (Marsh 1872)	Augé 1990	1 specimen
	synonym of Saniwa
	1 left dentary (n CB 16437)

	Tinosaurus europeocaenus Auge and Smith 1997	Augé 2005	1 specimen
	MHNH AV 16464 ,anterior left dentary
Reptilia - Eolacertidae

	Eolacerta sp. Nöth 1940	Augé 2005	1 specimen
	dorsal vertebra
Reptilia - Lacertidae

	Dormaalisaurus girardoti Augé and Smith 2002	Augé 2005	2 specimens
	incomplete left and right dentaries
Reptilia - Gekkonidae

	Gekkonidae indet. Gray 1825	Augé 2005	1 specimen
	fragment of left dentary
Reptilia

	Amphisbaenidae indet. (Gray 1865)	Augé 2005	2 specimens
	Av 7927, a vertebra; Av 7923, fragment of dentary
Reptilia - Necrosauridae

	Necrosaurus sp. Filhol 1876	Augé 2005	3 specimens
	MNHN, three osteoderms
Reptilia - Testudines - Podocnemididae

	Neochelys eocaenica (de Stefano 1902)	Lapparent de Broin 2003
	original and current combination Cistudo eocaenica
Reptilia - Testudines

	Allaeochelys sp. Noulet 1867	Broin 1977
Reptilia - Testudines - Trionychidae

	Trionyx sp. Geoffroy Saint-Hilaire 1809	Broin 1977

	Palaeotrionyx vittatus (Pomel 1847)	Broin 1977
	recombined as Axestemys vittata
Reptilia - Testudines - Geoemydidae

	Palaeochelys sp. Meyer 1847	Broin 1977
see common names

Geography

Country:	France	State/province:	Marne
Coordinates:	49.1° North, 4.0° East (view map)
Paleocoordinates:	44.5° North, 1.3° East
Basis of coordinate:	unpublished field data
Altitude:	119 meters
Geographic resolution:	small collection

Time

Period:	Paleogene	Epoch:	Eocene
Stage:	Ypresian	10 m.y. bin:	Cenozoic 2
Key time interval:	Ypresian
Age range of interval:	56.00000 - 47.80000 m.y. ago

Stratigraphy

Formation:	Lignites de Soissonais
Stratigraphic resolution:	formation
Stratigraphy comments: late Sparnacian, NP12, MP8+9 (Smith & Smith, 2003)

Lithology and environment

Primary lithology:	not reported
Environment:	terrestrial indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	mesofossils
Preservation of anatomical detail:	good
Disassociated major elements:	many
Disassociated minor elements:	some
Fragmentation:	occasional

Collection methods and comments

Collection methods:	field collection
Reason for describing collection:	taxonomic analysis

Metadata

Also known as:	Avenay-Val-d’Or
Database number:	35535
Authorizer:	M. Carrano, K. Beard, J. Alroy, P. Mannion, P. Holroyd, J. Head, D. Nicholson, R. Butler, J. Mueller, P. Barrett	Enterer:	M. Carrano, K. Beard, J. Alroy, T. Liebrecht, P. Mannion, P. Holroyd, J. Head, D. Nicholson, R. Butler, T. Cleary
Modifier:	P. Mannion	Research group:	vertebrate
Created:	2003-12-19 12:54:36	Last modified:	2016-11-16 08:15:25
Access level:	the public	Released:	2003-12-19 12:54:36
Creative Commons license:	CC BY

Reference information

Primary reference:

9599.

5% 31240

D. E. Russell, M. Godinot, P. Louis and D. E. Savage. 1979. Apatotheria (Mammalia) de l'Eocène inférieur de France et de Belgique. Bulletin du Muséum National d'Histoire Naturelle, section C: Science de la Terre 1(3):203-243 [M. Carrano/M. Carrano/J. Alroy]

Secondary references:

43530		M. Augé. 1990. La faune de lézards et d'amphisbaenes ed l'Eocene inférieur de Condé-en-Brie (France). Bulletin du Muséum National d'Histoire Naturelle. Paris. 4 séries 12(2):111-141 [J. Head/J. Head]
59290		M. L. Augé. 2005. Evolution des lézards du Paléogène en Europe. Mémoires du Muséum national d'histoire naturelle 192:1-369 [P. Barrett/T. Cleary/T. Cleary]
60047		F. Broin. 1977. Contribution a l'etude des Cheloniens. Cheloniens continentaux due Cretace et du Tertiare de France. Memoires du Museum National d'Histoire Naturelle, Nouvelle Serie, Serie C, Sciences de la terre 38:1-366 [P. Holroyd/P. Holroyd/P. Holroyd]
1635		P. D. Gingerich. 1976. Cranial Anatomy and Evolution of Early Tertiary Plesiadapidae (Mammalia, Primates). University of Michigan Papers on Paleontology 15:1-141 [J. Alroy/J. Alroy/P. Mannion]
60859		J. J. Hooker. 2010. The mammal fauna of the early Eocene Blackheath Formation of Abbey Wood, London. Monograph of the Palaeontographical Society 165:1-162 [P. Mannion/P. Mannion]
10304		J. J. Hooker, D. E. Russell, and A. Phelzion. 1999. A new family of Plesiadapiformes (Mammalia) from the Old World Lower Paleogene. Palaeontology 42(3):377-407 [J. Alroy/E. Leckey/P. Mannion]
58692		F. Lapparent de Broin. 2003. Neochelys sp. (Chelonii, Erymnochelyinae), from Silveirinha, early Eocene, Portugal. Ciencias da Terra (UNL) 15:117-132 [P. Holroyd/P. Holroyd]
66265		S. López-Torres and M. T. Silcox. 2018. The European Paromomyidae (Primates, Mammalia): taxonomy, phylogeny, and biogeographic implications. Journal of Paleontology [P. Mannion/P. Mannion]
55138		J.-C. Rage. 1973. Présence de Dunnophis (Reptilia, Serpentes) dans l'Eocène et l'Oligocène européen. Compte Rendu sommaire des séances de la Société Géologique de France 1973(3):76-78 [J. Mueller/T. Liebrecht/T. Liebrecht]
7882		J.-C. Rage. 1977. An erycine snake (Boidae) of the genus Calamagras from the French lower Eocene, with comments on the phylogeny of the Erycinae. Herpetologica 33(4):459-463 [J. Head/J. Head/J. Alroy]
60942		T. H. V. Rich. 1971. Deltatheridia, Carnivora, and Condylarthra (Mammalia) of the Early Eocene, Paris Basin, France. University of California Publications in Geological Sciences 88:1-72 [P. Mannion/P. Mannion]
9822	5% 15520	D. Russell, A. Phelizon, and P. Louis. 1992. Avenius n. gen. (Mammalia, Primates?, Mycrosyopidae) de l'Eocene inferieur de France. Comptes Rendus de l'Academie des Sciences, Paris 314(2):243-250 [E. Fara/E. Fara/J. Alroy]
30281		D. E. Russell, P. Louis, and D. E. Savage. 1973. Chiroptera and Dermoptera of the French early Eocene. University of California Publications in Geological Sciences 95:1-57 [K. Beard/K. Beard]
19707	5% 57680	T. Smith and R. Smith. 2003. Terrestrial mammals as biostratigraphic indicators in upper Paleocene-lower Eocene marine deposits of the southern North Sea Basin. Geological Society of America Special Paper 369:513-520 [M. Uhen/M. Uhen]
60835		F. Solé. 2013. New proviverrine genus from the Early Eocene of Europe and the first phylogeny of Late Palaeocene–Middle Eocene hyaenodontidans (Mammalia). Journal of Systematic Palaeontology 11(4):375-398 [P. Mannion/P. Mannion]
60656		F. Solé. 2014. New carnivoraforms from the early Eocene of Europe and their bearing on the evolution of the Carnivoraformes. Palaeontology 57(5):963-978 [R. Butler/R. Butler]
60838		F. Solé, J. Falconett, and D. Vidalenc. 2015. New fossil Hyaenodonta (Mammalia, Placentalia) from the Ypresian and Lutetian of France and the evolution of the Proviverrinae in southern Europe. Palaeontology 58(6):1049-1072 [P. Mannion/P. Mannion/P. Mannion]
61257		J. Sudre and J. Erfurt. 1996. Les artiodactyles du gisement yprésien terminal de Premontre (Aisne, France). Palaeovertebrata 25(2-4):391-414 [P. Mannion/P. Mannion]