Tsagan Khushu Quarry 1 [SMPE]: Bumbanian, Mongolia
collected by V. Reshetov & South Gobi party 1980

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Phoenicopteriformes - Juncitarsidae
Juncitarsidae informal large species
Zelenkov 2021
PIN 3104/100, 107, 116, 158, and 681
Juncitarsidae informal small species
Zelenkov 2021
PIN 3104/73, 506, 645, and 693
Reptilia - Anseriformes - Presbyornithidae
Presbyornis sp. Wetmore 1926
Kurochkin and Dyke 2010
PIN specimens (quadrates, sterna, furculae, coracoids, scapulae, humeri, radii, ulnae, carpometacarpi, manual phalanges, femora and tibiotarsi)
    = Bumbalavis anatoides n. gen., n. sp. Zelenkov 2021
Zelenkov 2021
PIN 3104/149 - holotype; referred specimens: PIN 3104/118, 214 and 712
Presbyornis mongoliensis n. sp. Kurochkin and Dyke 2010
Kurochkin and Dyke 2010
PIN 3104-607 - type (distal end of left tarsometarsus) and paratypes: PIN 3104-69, 229, 601, 684 (distal ends of left tarsometatarsi) and PIN 3104-115, 716 (distal ends of right tarsometatarsi)
Reptilia - Galliformes
Galliformes indet. Temminck 1820
Hwang et al. 2010
ISMD-VP-10 (proximal portion of a left humerus)
Reptilia - Testudines - Lindholmemydidae
Paramongolemys khosatzkyi Danilov and Sukhanov 2013
Danilov and Sukhanov 2013
PIN, no. 3104
Reptilia - Testudines - Emydidae
Pseudochrysemys gobiensis n. gen., n. sp. Sukhanov and Narmandakh 1976
Dashzeveg and McKenna 1977
Mammalia - Condylarthra - Hyopsodontidae
Midiagnus gracilis n. sp. Kondrashov 2004
Kondrashov 2004
Hyopsodus orientalis
Mammalia - Primates
Altanius orlovi n. gen., n. sp. Dashzeveg and McKenna 1977
Dashzeveg and McKenna 1977
ASMPR GISPS coll. 7, no. 20-8 - holotype
Mammalia
Eomylus zhigdensis
Mammalia - Rodentia
Ivanantonia efremovi Shevyreva 1989
Mammalia - Rodentia - Yuomyidae
? Esesempomys centralasiae Shevyreva 1989
Mammalia - Rodentia - Alagomyidae
Alagomys inopinatus n. gen., n. sp. Dashzeveg 1990
Dashzeveg 1990
PSS 20-176 - holotype; referred material: PSS 20-177
Mammalia - Rodentia - Tamquammyidae
Orogomys obscurus n. gen., n. sp. Dashzeveg 1990
Dashzeveg 1990
PSS 20-174 - holotype; referred material: PSS 20-175
Adolomys ameristus n. gen., n. sp. Shevyreva 1989
Bumbanomys edestus Shevyreva 1989
Tsagamys subitus n. gen., n. sp. Dashzeveg 1990
Dashzeveg 1990
PSS 20-96 - holotype
Sharomys singularis Dashzeveg 1990
Sharomys parvus n. sp. Dashzeveg 1990
Dashzeveg 1990
PSS 20-24; referred material: PSS 20-23
Kharomys gracilis n. sp. Dashzeveg 1990
Dashzeveg 1990
PSS 20-92
Tsagankhushumys deriphatus Shevyreva 1989
Mammalia - Rodentia
Euboromys grandis (Dashzeveg 1990)
Averianov 1996
Mammalia - Mixodontia - Decipomyidae
Decipomys mongoliensis Dashzeveg et al. 1998
Mammalia - Mimotonidae
Gomphos elkema Shevyreva 1975
Mammalia - Eurymylidae
Rhombomylus cf. turpanensis Zhai 1978
Zagmys insolitus Dashzeveg et al. 1987
Mammalia - Lipotyphla
Lipotyphla indet. Haeckel 1866
Russell and Dashzeveg 1986
synonym of Eulipotyphla
PSS 20-62
Mammalia - Nyctitheriidae
Nyctitheriidae indet. Simpson 1928
Russell and Dashzeveg 1986
PSS 20-60
Bumbanius rarus n. gen., n. sp. Russell and Dashzeveg 1986
Russell and Dashzeveg 1986
Oedolius perexiguus n. gen., n. sp. Russell and Dashzeveg 1986
Russell and Dashzeveg 1986
Edzenius lus n. gen., n. sp. Lopatin 2006
Lopatin 2006
PIN 3104/949 - holotype
Eosoricodon terrigena n. gen., n. sp. Lopatin 2005
Lopatin 2006
PIN 3104/900 - holotype
Mammalia - Plesiosoricidae
cf. Hyracolestes sp. Matthew and Granger 1925
Russell and Dashzeveg 1986
PSS 20-124
    = Ordolestes ordinatus n. gen., n. sp. Lopatin 2006
Lopatin 2006
PIN 3104/952 - holotype
Mammalia - Acreodi - Mesonychidae
Dissacus zanabazari n. sp. Geisler and McKenna 2007
Geisler and McKenna 2007 1 individual
Mammalia - Creodonta - Hyaenodontidae
Arfia langebadreae n. sp. Lavrov and Lopatin 2004
Lavrov and Lopatin 2004
PIN, no. 3104/785 - type
? Prototomus sp. Cope 1874
Lavrov and Lopatin 2004
PIN, no. 3104/787
Mammalia - Homacodontidae
Tsaganohyus pecus n. gen., n. sp. Kondrashov et al. 2004
Kondrashov et al. 2004 1 specimen
Mammalia - Perissodactyla
Homogalax namandicus
Hyracotherium gabuniai Hooker and Dashzeveg 2004
recombined as Protomoropus gabuniai
Mammalia - Cimolestidae
Naranius infrequens n. gen., n. sp. Russell and Dashzeveg 1986
Russell and Dashzeveg 1986
Bagalestes trofimovi n. gen., n. sp. Lopatin 2006
Lopatin 2006
PIN 3104/960 - holotype; referred material: PIN 3104/959
Mammalia - Sarcodontidae
Prosarcodon maturus n. sp. Lopatin and Kondrashov 2004
Lopatin 2006
PIN 3104/778 - type
Mammalia - Proteutheria - Palaeoryctidae
Tsaganius ambiguus n. gen., n. sp. Russell and Dashzeveg 1986
Russell and Dashzeveg 1986
see common names

Geography
Country:Mongolia State/province:Southern Mongolia
Coordinates: 44.6° North, 106.2° East (view map)
Paleocoordinates:45.2° North, 98.0° East
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period:Paleogene Epoch:Eocene
Stage:Ypresian 10 m.y. bin:Cenozoic 2
Key time interval:Bumbanian
Age range of interval:56.00000 - 47.80000 m.y. ago
Stratigraphy
Formation:Naran Bulak Member:Bumban
Stratigraphic resolution:bed
Lithology and environment
Primary lithology:red gravel
Includes fossils?Y
Lithology description: Red gravelstone
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils,microfossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Museum repositories:PIN
Collectors:V. Reshetov & South Gobi party Collection dates:1980
Metadata
Also known as:Tsagan-Khusu
Database number:37206
Authorizer:X. Wang, P. Mannion, K. Beard, P. Holroyd, M. Uhen Enterer:X. Wang, P. Mannion, K. Beard, P. Holroyd, M. Uhen
Modifier:M. Carrano Research group:vertebrate
Created:2004-03-04 18:01:34 Last modified:2019-06-19 14:01:26
Access level:the public Released:2004-03-04 18:01:34
Creative Commons license:CC BY
Reference information

Primary reference:
9976.5% 1320D. Dashzeveg, J.-L. Hartenberger, T. Martin and S. Legendre. 1998. A peculiar minute Glires (Mammalia) from the early Eocene of Mongolia. Bulletin of Carnegie Museum of Natural History 34:194-209 [X. Wang/X. Wang/P. Mannion]

Secondary references:

59967 A. O. Averianov. 1996. Early Eocene Rodentia of Kyrgyzstan. Bulletin du Museum national d'Histoire naturelle, Paris, 4e serie 18:629-662 [P. Mannion/P. Mannion]
45403 I. G. Danilov and V. B. Sukhanov. 2013. A new basal testudinoid turtle (Testudinoidea: “Lindholmemydidae”) from the Upper Paleocene of Mongolia. Paleontological Journal 47(1):104-113 [P. Mannion/P. Mannion]
59992 D. Dashzeveg. 1990. The earliest rodents (Rodentia, Ctenodactyloidea) of Central Asia. Acta Zoologica Cracoviensia 33(2):11-35 [P. Mannion/P. Mannion]
59993 D. Dashzeveg. 1990. New trends in adaptive radiation of Early Tertiary rodents (Rodentia, Mammalia). Acta Zoologica Cracoviensia 33(3):37-44 [P. Mannion/P. Mannion]
59493 D. Dashzeveg and M. C. McKenna. 1977. Tarsioid primate from the Early Tertiary of the Mongolian People’s Republic. Acta Palaeontologica Polonica 22(2):119-137 [P. Mannion/P. Mannion]
31106 J. H. Geisler and M. C. McKenna. 2007. A new species of mesonychian mammal from the lower Eocene of Mongolia and its phylogenetic relationships. Acta Palaeontologica Polonica 52(1):189-212 [M. Uhen/M. Uhen]
37599 S. H. Hwang, G. Mayr, and M. Bolortsetseg. 2010. The earliest record of a galliform bird in Asia, from the late Paleocene–early Eocene of the Gobi Desert, Mongolia. Journal of Vertebrate Paleontology 30(5):1642-1644 [P. Mannion/P. Mannion]
19557 P. E. Kondrashov. 2004. A new hyopsodontid (Mammalia, Condylarthra) from the early Eocene of Mongolia. in Paleogene Mammals, S. G. Lucas, K. E. Zeigler, and P.E. Kondrashov (eds.), New Mexico Museum of Natural History and Science Bulletin (26)165-167 [P. Holroyd/P. Holroyd]
36622 P. E. Kondrashov, A. V. Lopatin, and S. G. Lucas. 2004. The oldest known Asian artiodactyl (Mammalia). New Mexico Museum of Natural History and Science Bulletin 26:205-208 [M. Uhen/M. Uhen]
37600 E. N. Kurochkin and G. J. Dyke. 2010. A large collection of Presbyornis (Aves, Anseriformes, Presbyornithidae) from the late Paleocene and early Eocene of Mongolia. Geological Journal 45:375-387 [P. Mannion/P. Mannion]
59965 A. V. Lavrov and A. V. Lopatin. 2004. A New Species of Arfia (Hyaenodontidae, Creodonta) from the Basal Eocene of Mongolia. Paleontological Journal 38(4):448-457 [P. Mannion/P. Mannion]
50837 A. V. Lopatin. 2006. Early Paleogene insectivore mammals of Asia and establishment of the major groups of Insectivora. Paleontological Journal 40(3):S205-S405 [P. Wagner/P. Wagner/P. Mannion]
19559 A. V. Lopatin and P. E. Kondrashov. 2004. Sarcodontinae, a new subfamily of micropternodontid insectivores from the early Paleocene-middle Eocene of Asia. in Paleogene Mammals, S. G. Lucas, K. E. Zeigler, and P.E. Kondrashov (eds.), New Mexico Museum of Natural History and Science Bulletin (26)177-184 [P. Holroyd/P. Holroyd]
29424 D. E. Russell and D. Dashzeveg. 1986. Early Eocene insectivores (Mammalia) from the People's Republic of Mongolia. Palaeontology 29:269-291 [K. Beard/K. Beard]
60843 F. Solé, T. Smith, R. Tabuce and B. Marandat. 2015. New dental elements of the oldest proviverrine mammal from the early Eocene of Southern France support possible African origin of the subfamily. Acta Palaeontologica Polonica 60(3):527-538 [P. Mannion/P. Mannion]
77696 N. V. Zelenkov. 2021. A Revision of the Palaeocene–Eocene Mongolian Presbyornithidae (Aves: Anseriformes). Paleontological Journal 55(3):323-330 [P. Mannion/P. Mannion]