PBDB Collection

Kitadani Dinosaur Quarry (lower, BB I): Aptian, Japan
collected 1988–

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

unclassified

	Nagdongia sp.	Currie and Azuma 2006

	Plantae indet. Haeckel 1866	Currie and Azuma 2006
	"plants"
Actinopteri - Amiiformes

	Amiiformes indet. Hay 1929	Azuma 2003
	scales
Reptilia - Theropoda

	Theropoda indet. Marsh 1881	Azuma 2003	2 specimens
Reptilia - Spinosauridae

	Spinosauridae indet. Stromer 1915	Hattori and Azuma 2020	18 specimens
	FPDM-V-546, 9475, 9999, 10000, 10237–10249, 10251
Reptilia

	Coelurosauria indet. Huene 1914	Currie and Azuma 2006

	Aves indet. Linnaeus 1758	Currie and Azuma 2006	1 individual
	FPDM-V-9769
	= Fukuipteryx prima n. gen., n. sp. Imai et al. 2019	Imai et al. 2019
Reptilia - Deinocheiridae

	Tyrannomimus fukuiensis n. gen., n. sp. Hattori et al. 2023	Hattori et al. 2023
	Holotype: FPDM-V-11311; Paratype: FPDM-V-10295; Additional specimens: FPDM-V-11313; FPDM-V-11314–11316; FPDM-V-11317–11320; FPDM-V-11321–11323; FPDM-V-8579, 8580, 8945, 11324–11329, 11332; FPDM-V-9348, 11333–11337; FPDM-V-11338; FPDM-V-11339–11341; FPDM-V-11343–11345; FPDM-V-11343, 11346–11348;
Reptilia - Neovenatoridae

	Fukuiraptor kitadaniensis n. gen., n. sp. Azuma and Currie 2000	Azuma and Currie 2000	14 individuals
	MNI; multiple individuals of different ontogenetic stages (11 measurements)
Reptilia - Stegosauridae

	Stegosauridae indet. Marsh 1880	Isaji and Hasegawa 1990	1 specimen
	spine
Reptilia

	Ornithopoda indet. Marsh 1881	Azuma 2003	2 specimens

	Fukuisaurus tetoriensis n. gen., n. sp. Kobayashi and Azuma 2003	Kobayashi and Azuma 2003	1 individual
Reptilia - Brachiosauridae

	Brachiosauridae indet. Riggs 1904	Azuma 2003	1 individual
	"brachiosaurid"
Reptilia - Goniopholididae

	Goniopholididae indet. Cope 1875	Azuma 2003	1 individual
Reptilia - Testudines - Nanhsiungchelyidae

	Basilemys sp. Hay 1902	Azuma 2003
	= Nanhsiungchelyidae indet. Yeh 1966	Danilov and Syromyatnikova 2008
Reptilia - Testudines

	Testudinoidea indet. Gray 1825	Azuma 2003
Reptilia - Testudines - Trionychidae

	Trionychidae indet. Gray 1825	Azuma 2003	5 specimens
	FPDM-V9487, V0127, V2761, V6370 and V1836
Reptilia - Testudines - Adocidae

	Adocus sp. Cope 1868	Azuma 2003
Reptilia - Testudines - Sinemydidae

	? Sinemydidae indet. Yeh 1963	Azuma 2003
Mammalia - Multituberculata - Eobaataridae

	Eobaataridae indet. Kielan-Jaworowska et al. 1987	Miyata et al. 2016	1 specimen
	FPDM-V9171 (left p4 crown)
Mammalia - Eutriconodonta - Triconodontidae

	? Triconodontidae indet. Marsh 1887	Miyata et al. 2016	1 specimen
	FPDM-V9172 (fragment of possible right dentary with a damaged molariform crown and roots)
Gastropoda

	Gastropoda indet. Cuvier 1795	Shibata and Goto 2008
Bivalvia - Unionida - Unionidae

	Plicatounio sp. Kobayashi and Suzuki 1936	Azuma 2002
Bivalvia - Trigoniida - Pseudohyriidae

	Pseudohyria sp. MacNeil 1936	Azuma 2002
Bivalvia - Trigoniida - Trigonioididae

	Trigonioides sp. Kobayashi and Suzuki 1936	Azuma 2002

	Nippononaia ryosekiana	Isaji and Hasegawa 1990
see common names

Geography

Country:	Japan	State/province:	Fukui	County:	Katsuyama
Coordinates:	36.1° North, 136.5° East (view map)
Paleocoordinates:	47.8° North, 137.0° East
Basis of coordinate:	stated in text
Altitude:	134 meters
Geographic resolution:	outcrop

Time

Period:	Cretaceous	Epoch:	Early/Lower Cretaceous
Stage:	Aptian	10 m.y. bin:	Cretaceous 3
Key time interval:	Aptian
Age range of interval:	121.40000 - 113.00000 m.y. ago

Stratigraphy

Geological group:	Tetori	Formation:	Kitadani
Stratigraphic resolution:	bed
Stratigraphy comments: Akaiwa Subgroup, lower part of formation *"Unit C") Tsukiji et al. 2018: Kubota (2005) assigned the age of the Kitadani Formation to the Barremian based on the co-occurrence of charophyte gyrogonites: Clavator harrisii var. reyi, Mesochara harrisi, and Mesoshara stipitata. Recently, Sano (2015) suggested that the co-occurrence of these three species indicates the Barre- mianeAptian age of the Kitadani Formation because these species are also reported elsewhere from the Aptian. Recent interpretation of the stratigraphy of the Kitadani Formation is younger than the thick sandstone dominant unit (Unit B of Sano et al., 2008), which is assigned to the BarremianeAptian Akaiwa Formation (Sano and Yabe, 2016). Thus the Kitadani Formation is likely assigned to the Aptian in age.

Lithology and environment

Primary lithology:	sandstone
Secondary lithology:	"shale"
Includes fossils?	Y
Lithology description: "alternating beds of thick sandstone and shale interbedded with "tuff" layers."
Environment:	fluvial indet.
Geology comments: "ancient meandering rivers"

Taphonomy

Modes of preservation:	body
Degree of concentration:	concentrated
Size of fossils:	macrofossils,mesofossils
Preservation of anatomical detail:	good
Articulated whole bodies:	some
Associated major elements:	some
Disassociated major elements:	some
Disassociated minor elements:	many
Fragmentation:	occasional

Collection methods and comments

Collection methods:	selective quarrying,mechanical,field collection
Reason for describing collection:	taxonomic analysis
Collection dates:	1988–
Collection method comments: excavated by Fukui Prefectural Museum

Metadata

Also known as:	Bonebed I, Katsuyama
Database number:	38020
Authorizer:	M. Carrano, E. Dunne, P. Mannion	Enterer:	M. Carrano, P. Mannion, E. Tasimov
Modifier:	G. Varnham	Research group:	vertebrate
Created:	2004-03-31 11:41:39	Last modified:	2022-02-02 08:40:24
Access level:	the public	Released:	2004-03-31 11:41:39
Creative Commons license:	CC BY

Reference information

Primary reference:

54260.

M. Matsukawa and I. Obata. 1994. Dinosaurs and sedimentary environments in the Japanese Cretaceous: a contribution to dinosaur facies in Asia based on molluscan palaeontology and stratigraphy. Cretaceous Research 15(1):101-125 [M. Carrano/M. Carrano]

Secondary references:

43728		L. Alcalá, E. Espílez, L. Mampel, J. I. Kirkland, M. Ortiga, D. Rubio, A. González, D. Ayala, A. Cobos, R. Royo-Torres, F. Gascó and M. D. Pesquero. 2012. A new Lower Cretaceous vertebrate bonebed near Arino (Tereul, Aragon, Spain); found and managed in a joint collaboration between a mining company and a palaeontological park. Geoheritage 4:275-286 [P. Mannion/J. Tennant/M. Carrano]
34308	ETE	Y. Azuma. 2002. Dinosaur remains from Katsuyama City, Fukui Prefecture. 2002 Annual Meeting, The Palaeontological Society of Japan, June 21-23, 2002, Katsuyama. Abstracts with Programs 14 [M. Carrano/M. Carrano]
10133		Y. Azuma. 2003. Early Cretaceous vertebrate remains from Katsuyama city, Fukui prefecture, Japan. Memoir of the Fukui Prefectural Dinosaur Museum 2:17-21 [M. Carrano/M. Carrano/M. Carrano]
10135		Y. Azuma and P. J. Currie. 2000. A new carnosaur (Dinosauria: Theropoda) from the Lower Cretaceous of Japan. Canadian Journal of Earth Sciences 37:1735-1753 [M. Carrano/M. Carrano/M. Carrano]
29361	ETE	Y. Azuma and M. Shibata. 2008. New dinosaurs of the Tetori Group. International Dinosaur Symposium in Fukui 2008: Recent Progress of the Study on Asian Dinosaurs and Paleoenvironments. Fukui Prefectural Dinosaur Museum, Fukui 29-31 [M. Carrano/M. Carrano]
37922		Y. Azuma and M. Shibata. 2010. Fukuititan nipponensis, a new titanosauriform sauropod from the Early Cretaceous Tetori Group of Fukui Prefecture, Japan. Acta Geologica Sinica 84(3):454-462 [P. Mannion/P. Mannion]
54268		Y. Azuma and Y. Tomida. 1995. Early Cretaceous dinosaur fauna of the Tetori Group in Japan. In A. Sun, Y. Wang (eds.), Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota, Short Papers 125-131 [M. Carrano/M. Carrano/P. Wagner]
54277		Y. Azuma and Y. Tomida. 1997. Japanese dinosaurs. In P. J. Currie, K. Padian (eds.), Encyclopedia of Dinosaurs 375-379 [M. Carrano/M. Carrano/P. Wagner]
24532	ETE	P. J. Currie and Y. Azuma. 2006. New specimens, including a growth series, of Fukuiraptor (Dinosauria, Theropoda) from the Lower Cretaceous Kitadani Quarry of Japan. Journal of the Paleontological Society of Korea 22(1):173-193 [M. Carrano/M. Carrano]
41748		I. G. Danilov and E. V. Syromyatnikova. 2008. New materials on turtles of the family Nanhsiungchelyidae from the Cretaceous of Uzbekistan and Mongolia, with a review of the Nanhsiungchelyid record in Asia. Proceedings of the Zoological Institute RAS 312(1/2):3-25 [R. Benson/R. Benson/P. Mannion]
55479		Fukui Kenritsu Kyoryu Hakubutsukan. 2010. [Exhibit Guidebook of the Fukui Prefectural Dinosaur Museum]. 1-212 [M. Carrano/M. Carrano]
75997		S. Hattori and Y. Azuma. 2020. Spinosaurid teeth from the Lower Cretaceous Kitadani Formation of the Tetori Group, Fukui, Japan. Memoir of the Fukui Prefectural Dinosaur Museum 19:1-9 [M. Carrano/M. Carrano]
85757		S. Hattori, M. Shibata, S. Kawabe, T. Imai, H. Nishi and Y. Azuma. 2023. New theropod dinosaur from the Lower Cretaceous of Japan provides critical implications for the early evolution of ornithomimosaurs. Scientific Reports 13(13842):1-16 [E. Dunne/E. Tasimov/M. Carrano]
70937		T. Imai, Y. Azuma, S. Kawabe, M. Shibata, K. Miyata, M. Wang, and Z. Zhou. 2019. An unusual bird (Theropoda, Avialae) from the Early Cretaceous of Japan suggests complex evolutionary history of basal birds. Communications Biology 2(1):399:1-11 [M. Carrano/M. Carrano]
53088		S. Isaji and Y.T. D. Hasegawa. 1990. [Discovery of Nippononaia ryosekiana and stegosaurid spine from the Kitadani Formation of the Tetori Group]. 139th Regular Meeting of the Palaeontological Society of Japan, June 30-July 1, 1990 35 [P. Wagner/P. Wagner/M. Carrano]
10136		Y. Kobayashi and Y. Azuma. 2003. A new iguanodontian (Dinosauria: Ornithopoda) from the Lower Cretaceous Kitadani Formation of Fukui Prefecture, Japan. Journal of Vertebrate Paleontology 23(1):166-175 [M. Carrano/M. Carrano/M. Carrano]
56885		K. Miyata, Y. Azuma, and M. Shibata. 2016. New mammalian specimens from the Lower Cretaceous Kitadani Formation, Tetori Group, Fukui, Japan. Historical Biology 28(1-2):139-150 [P. Mannion/P. Mannion]
65859		Y. Nakajima, I. G. Danilov, R. Hirayama, T. Sonoda, and T. M. Scheyer. 2017. Morphological and histological evidence for the oldest known softshell turtles from Japan. Journal of Vertebrate Paleontology 37(2):e1278606:1-11 [P. Mannion/P. Mannion]
41006		H. Saegusa and Y. Tomida. 2011. Titanosauriform teeth from the Cretaceous of Japan. Anais da Academia Brasileira de Ciências 83(1):247-265 [M. Carrano/M. Carrano]
69508		S.-i. Sano and A. Yabe. 2017. Fauna and flora of Early Cretaceous Tetori Group in central Japan: the clues to revealing the evolution of Cretaceous terrestrial ecosystem in Asia. Palaeoworld 26:253-267 [M. Carrano/M. Carrano]
54271		M. Shibata and Y. Azuma. 2015. New basal hadrosauroid (Dinosauria: Ornithopoda) from the Lower Cretaceous Kitadani Formation, Fukui, central Japan. Zootaxa 3914(4):421-440 [P. Mannion/J. Tennant/M. Carrano]
29441	ETE	M. Shibata and M. Goto. 2008. Report of the 3rd Dinosaur Excavation Project in Katsuyama, Fukui, 2007. Memoir of the Fukui Prefectural Dinosaur Museum 7:109-116 [M. Carrano/M. Carrano/M. Carrano]
60586		S. Suzuki, M. Shibata, Y. Azuma, H. Yukawa, T. Sekiya and Y. Masaoka. 2015. Sedimentary environment of dinosaur fossil bearing successions of the Lower Cretaceous Kitadani Formation, Tetori Group, Katsuyama City, Fukui, Japan. Memoir of the Fukui Prefectural Dinosaur Museum 14:1-9 [M. Carrano/M. Carrano]