MAD93-35 (SUNY): Maastrichtian, Madagascar

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Dinosauria
Dinosauria indet. Owen 1842
Roberts et al. 2007
Reptilia - Noasauridae
Masiakasaurus knopfleri Sampson et al. 2001
Carrano et al. 2011 8 specimens
FMNH PR 2220, 2199; UA 9183, 9734, 9773, 9199, 9185, 9137
Reptilia
Enantiornithes indet. Walker 1981
O'Connor and Forster 2010 1 specimen
FMNH PA 751
Mahajangasuchus insignis n. gen., n. sp. Buckley and Brochu 1999
Turner and Buckley 2008 3 specimens
UA 8654; FMNH PR 2389, 2449
Simosuchus clarki Buckley et al. 2000
Krause et al. 2010
FMNH PR 2600 (isolated tooth)
Reptilia - Serpentes
Serpentes indet. Linnaeus 1758
LaDuke et al. 2010 1 specimen
FMNH PR 2579
Reptilia - Nigerophiidae
Kelyophis hechti LaDuke et al. 2010
LaDuke et al. 2010
Reptilia - Madtsoiidae
Menarana nosymena LaDuke et al. 2010
LaDuke et al. 2010
Madtsoiidae indet. (Hoffstetter 1961)
LaDuke et al. 2010 1 specimen
FMNH PR 2572
    = Adinophis fisaka Pritchard et al. 2014
Pritchard et al. 2014
FMNH PR 2572 (mid-trunk vertebra)
Reptilia - Testudines
Sokatra antitra n. gen., n. sp. Gaffney and Krause 2011
Gaffney and Krause 2011
UA 9769 - holotype (partial skull, lacking the premaxillae, basioccipital, and with damaged basisphenoid, quadrates, prootics, and opisthotics). FMNH nos. PR 2647 (supraoccipital), PR 2648 (left frontal), PR 2649 (partial skull with articulated or associated quadrates, both opisthotics, left exoccipital, right prootic, left pterygoid, and right palatine)and UA nos. 9864 (partial right quadrate) and 9865 (partial left maxilla) were recovered from a small (2 m2) quarried surface at this locality
Reptilia - Testudines - Podocnemididae
cf. Erymnochelys sp. Baur 1888
Gaffney and Forster 2003 1 specimen
FMNH PR 2231, mandible
Mammalia - Multituberculata
? Multituberculata indet. Cope 1884
Krause 2013
FMNH PM 61152 (fragmentary molar)
Multituberculata indet. (Cope 1884)
Krause et al. 2017 1 specimen
DMNH EPV.100010 (fragmentary femur)
Mammalia - Sudamericidae
Sudamericidae indet. Scillato-Yané and Pascual 1984
Krause 2013
UA 9937 (fragmentary molariform cheek tooth)
Lavanify miolaka Krause et al. 1997
1 specimen
Mammalia - Zhelestidae
Marsupialia indet. (Illiger 1811)
Krause 2001 1 specimen
    = Zhelestidae indet. Nessov 1990
Averianov et al. 2003
UA 8699 (lower molar)
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Asher and Krause 1998
FMNH-PR 1960 (right premaxilla)
Beelzebufo ampinga Evans et al. 2008
Evans et al. 2014 19 specimens
UA 9623 – fragment of otic and ventral processes of squamosal; UA 9624 – posterior process of left quadratojugal; UA 9625 – posteroventral process of right squamosal; UA 9626 – cranial fragment; UA 9627 – partial vertebral spine table; UA 9635 – anterior fragment of right maxilla; UA 9676 – fragment of right maxilla bearing maxillary nerve canal; UA 9677 – fragment of frontoparietal or squamosal; UA 9679 – cranial fragment; UA 9947 – presacral vertebra, interpreted as PS3; UA 9948 – posterior presacral vertebra; FMNH PR 1960 – partial right premaxilla; FMNH PR 2504 – vertebral centrum and partial neural arch; FMNH PR 2505 – facial process of right maxilla; FMNH PR 2506 – partial right maxilla; FMNH PR 2507 – facial process and pars dentalis of right maxilla; FMNH PR 2508 – cranial fragment from antorbital margin, frontoparietal or nasal; FMNH PR 2509 – conjoined midline frontoparietal fragment; FMNH PR 2510 – partial left maxilla. Locality MAD93-36: UA 9678 – vertebral spine table; UA 9949 – fragment of otic plate of right squamosal
unclassified
Cubiculum ornatus n. gen., n. sp. Roberts et al. 2007
Roberts et al. 2007
see common names

Geography
Country:Madagascar State/province:Mahajanga
Coordinates: 15.9° South, 46.6° East (view map)
Paleocoordinates:30.1° South, 38.3° East
Basis of coordinate:stated in text
Altitude:151 meters
Geographic resolution:small collection
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
Key time interval:Maastrichtian
Age range of interval:72.10000 - 66.00000 m.y. ago
Stratigraphy
Formation:Maevarano Member:Anembalemba
Stratigraphic resolution:bed
Stratigraphy comments: "uppermost white sandstone unit" = Anembalemba Member
Lithology and environment
Primary lithology:white sandstone
Includes fossils?Y
Environment:"channel"
Geology comments: "shallow and very broad channel-belt system characterized by an ephemeral and flood-prone discharge regime."
Taphonomy
Modes of preservation:body,trace
Degree of concentration:dispersed
Size of fossils:mesofossils
Spatial orientation:random
Preservation of anatomical detail:medium
Disassociated major elements:all
Fragmentation:frequent
Feeding/predation traces:arthropod boring
Temporal resolution:snapshot
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:DMNH,FMNH
Metadata
Database number:39266
Authorizer:M. Carrano, P. Holroyd, P. Mannion, R. Benson Enterer:M. Carrano, P. Mannion, J. Tennant, R. Benson, P. Holroyd
Modifier:P. Holroyd Research group:vertebrate
Created:2004-05-25 12:14:38 Last modified:2017-09-22 12:31:49
Access level:the public Released:2004-05-25 12:14:38
Creative Commons license:CC BY
Reference information

Primary reference:
10611.ETE D. W. Krause, G. V. R. Prasad, W. Koenigswald, A. Sahni, and F. E. Grine. 1997. Cosmopolitanism among Gondwanan Late Cretaceous mammals. Nature 390:504-507 [M. Carrano/M. Carrano/P. Holroyd]

Secondary references:

37865 R. J. Asher and D. W. Krause. 1998. The first pre-Holocene (Cretaceous) record of Anura from Madagascar. Journal of Vertebrate Paleontology 18(4):696-699 [P. Mannion/P. Mannion]
42861 A. O. Averianov, J. D. Archibald, and T. Martin. 2003. Placental nature of the alleged marsupial from the Cretaceous of Madagascar. Acta Palaeontologica Polonica 48(1):149-151 [P. Mannion/P. Mannion]
44450 M. T. Carrano, M. A. Loewen, and J. J. W. Sertich. 2011. New materials of Masiakasaurus knopfleri Sampson, Carrano, and Forster, 2001, and implications for the morphology of the Noasauridae (Theropoda: Ceratosauria). Smithsonian Contributions to Paleobiology 95:1-53 [M. Carrano/M. Carrano/P. Mannion]
49974 S. E. Evans, J. R. Groenke, M. E. H. Jones, A. H. Turner, and D. W. Krause. 2014. New material of Beelzebufo, a hyperossified frog (Amphibia: Anura) from the Late Cretaceous of Madagascar. PLoS One 9(1):e87236 [P. Mannion/J. Tennant]
42114 E. S. Gaffney and C. A. Forster. 2003. Side-necked turtle lower jaws (Podocnemididae, Bothremydidae) from the Late Cretaceous Maevarano Formation of Madagascar. American Museum Novitates 3397:1-13 [R. Benson/R. Benson]
39063 E. S. Gaffney and D. W. Krause. 2011. Sokatra, a new side-necked turtle (Late Cretaceous, Madagascar) and the diversification of the main groups of Pelomedusoides. American Museum Novitates 3728:1-28 [P. Mannion/P. Mannion]
12457ETE D. W. Krause. 2001. Fossil molar from a Madagascan marsupial. Nature 412:497-498 [M. Carrano/M. Carrano/M. Carrano]
46395 D. W. Krause. 2013. Gondwanatheria and ? Multituberculata (Mammalia) from the Late Cretaceous of Madagascar. Canadian Journal of Earth Sciences 50(3):324-340 [P. Mannion/P. Mannion/M. Carrano]
63451 D. W. Krause, S. Hoffman, and S. Wernining. 2017. First postcranial remains of Multituberculata (Allotheria, Mammalia) from Gondwana. Cretaceous Research 80:91-100 [P. Holroyd/P. Holroyd]
38067 D. W. Krause, J. J. W. Sertich, R. R. Rogers, S. C. Kast, A. H. Rasoamiaramanana and G. A. Buckley. 2010. Overview of the discovery, distribution, and geological context of Simosuchus clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar. Society of Vertebrate Paleontology Memoir 10:4-12 [P. Mannion/P. Mannion]
32931ETE T. C. LaDuke, D. W. Krause, J. D. Scanlon and N. J. Kley. 2010. A Late Cretaceous (Maastrichtian) snake assemblage from the Maevarano Formation, Mahajanga Basin, Madagascar. Journal of Vertebrate Paleontology 30(1):109-138 [M. Carrano/M. Carrano]
34251ETE P. M. O'Connor and C. A. Forster. 2010. A Late Cretaceous (Maastrichtian) avifauna from the Maevarano Formation, Madagascar. Journal of Vertebrate Paleontology 30(4):1178-1201 [M. Carrano/M. Carrano]
54399 A. C. Pritchard, J. A. McCartney, D. W. Krause and N. J. Kley. 2014. New snakes from the Upper Cretaceous (Maastrichtian) Maevarano Formation, Mahajanga Basin, Madagascar. Journal of Vertebrate Paleontology 34(5):1080-1093 [P. Mannion/P. Mannion]
23421ETE E. M. Roberts, R. R. Rogers, and B. Z. Foreman. 2007. Continental insect borings in dinosaur bone: examples from the Late Cretaceous of Madagascar and Utah. Journal of Paleontology 81(1):201-208 [M. Carrano/M. Carrano]
10610ETE R. R. Rogers, J. H. Hartman, and D. W. Krause. 2000. Stratigraphic analysis of Upper Cretaceous rocks in the Mahajanga Basin, northwestern Madagascar: implications for ancient and modern faunas. Journal of Geology 108:275-301 [M. Carrano/M. Carrano/M. Carrano]
28071ETE A. H. Turner and G. A. Buckley. 2008. Mahajangasuchus insignis (Crocodyliformes: Mesoeucrocodylia) cranial anatomy and new data on the origin of the eusuchian-style palate. Journal of Vertebrate Paleontology 28(2):382-408 [M. Carrano/M. Carrano]