Mazaterón: Bartonian, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri - Siluriformes - Bagridae
Bagridae indet. Bleeker 1852
Minwer-Barakat et al. 2012
Mammalia - Rodentia - Theridomyidae
Remys ibericus
Marigó et al. 2010
Pseudoltinomys crebrum n. sp. Peláez-Campomanes 1996
Peláez-Campomanes 1996
Mammalia - Rodentia - Pseudosciuridae
Sciuroides cf. siderolithicus (Picted and Humbert 1869)
Peláez-Campomanes 1996 21 elements
Teeth
Mammalia - Primates - Omomyidae
Pseudoloris cuestai n. sp. Minwer-Barakat et al. 2012
Minwer-Barakat et al. 2012
Mammalia - Primates - Adapidae
? Adapis sp. Cuvier 1822
Marigó et al. 2010
Mammalia - Primates - Notharctidae
Mazateronodon endemicus n. gen., n. sp. Marigó et al. 2010
Marigó et al. 2010
Mammalia - Perissodactyla - Lophiodontidae
Lophiodon sp. Cuvier 1822
Cuesta Ruiz-Colmenares 1996
11.816 (fragment of maxilla)
Mammalia - Perissodactyla - Palaeotheriidae
Leptolophus sp. Remy 1965
Cuesta Ruiz-Colmenares 1992
STUS 11565, incomplete left maxillary with P2–P3–(P4)–M1–(M2); STUS 8599, right M2; STUS 11826, right M2; STUS 11018, fragment of left M3 (labial wall not preserved); STUS 14247, right maxilla fragment with M2– M3; STUS 9201, right m3 (described in Perales-Gogenola et al. 2021)
Franzenium sp. n. sp. Casanovas and Santafe 1989
Cuesta Ruiz-Colmenares 1992
    = Franzenium durense n. sp. Cuesta 1993
Cuesta 1993
Palaeotherium sp. Cuvier 1804
    = Palaeotherium giganteum n. sp. Cuesta 1993
Cuesta 1993
? Plagiolophus sp. Pomel 1847
    = Plagiolophus sp. n. sp. Pomel 1847
Cuesta Ruiz-Colmenares 1992
    = Plagiolophus mazateronensis n. sp. Cuesta 1994
Cuesta Ruiz-Colmenares 1994
Paranchilophus remyi Casanovas and Santafé 1989
Cuesta Ruiz-Colmenares 1992
Cantabrotherium sp. n. sp. Casanova-Cladellas and Santafe-Llopis 1987
Cuesta Ruiz-Colmenares 1992
    = Cantabrotherium casanovasae n. sp. Cuesta 1993
Cuesta 1993
Iberolophus jimenezi n. sp. Badiola and Cuesta 2008
Badiola and Cuesta 2008 1 individual
US 11800, right hemimandible, exhibiting p1 and m1 alveoli, and the p2–p4 and m2–m3 series.
Mammalia - Xiphodontidae
cf. Leptotheridium indet. Stehlin 1910
Cuesta Ruiz-Colmenares 1993 1 specimen
13.625, M2
cf. Dichodon sp. Owen
Cuesta Ruiz-Colmenares 1993 1 specimen
11.814, fragment of mandible
Mammalia - Anoplotheriidae
Anoplotheriinae indet. Gray 1821
Cuesta Ruiz-Colmenares 1993 1 specimen
11.562, fragment of maxilla
Duerotherium sudrei n. gen., n. sp. Cuesta and Badiola 2009
Cuesta and Badiola 2009 1 individual
STUS 11562, a fragment of the left maxilla, showing the P3-M3 series.
cf. Dacrytherium sp. Filhol
Cuesta Ruiz-Colmenares 1993 1 specimen
13.624, fragment of maxilla
Mammalia - Creodonta
Carnivora indet. Bowdich 1821
    = Proviverra sp. Rütimeyer 1862
Cuesta Ruiz-Colmenares 1992
Reptilia
Diplocynodon sp. Pomel 1847
Iberosuchus sp. Antunes 1975
Squamata indet. (Oppel 1811)
Bolet 2023
IPS128812, fragment of tooth-bearing bone; IPS128815, indeterminate fragment of tooth-bearing bone; IPS128811, isolated unicuspid tooth
Reptilia - Lacertidae
Lacertidae indet. Bonaparte 1831
Minwer-Barakat et al. 2012
IPS128801, fragment of right maxilla; IPS12 8813, tooth attached to bone
cf. Lacertidae indet. Bonaparte 1831
Bolet 2023
Reptilia - Blanidae
cf. Blanidae indet. Kearney 2003
Bolet 2023
IPS49832, right dentary (Figure 1e); IPS128804, anterior portion of left dentary (Figure 1f) and IPS128799, posterior portion of right dentary
Reptilia - Anguidae
Anguidae indet. Gray 1825
Bolet 2023
IPS128803, partial premaxilla (Figure 2b); IPS128810, anterior portion of right dentary
cf. Melanosaurini indet. Sullivan 1979
Bolet 2023
IPS128809, osteoderm
Glyptosaurini indet. Sullivan 1979
Bolet 2023
IPS49833-49835, IPS128795-128797, IPS128805-128806, osteoderms
Anguinae indet. Gray 1825
Bolet 2023
IPS128798, partial skull bone (Figure 2c); IPS49838, partial left frontal bone (Figure 2d); IPS128807, a partial caudal vertebra (Figure 2e); IPS128809 (Figure 2f), fragment of osteoderm; IPS49836, and IPS49840 (fragmentary body osteoderms); IPS49844, a fragmentary vertebra; IPS49841, a tooth-bearing bone with one broken tooth.
Reptilia - Iguanidae
Iguanidae indet. Oppel 1811
Bolet 2023
IPS128802, fragment of maxilla (Figure 2a); IPS49839, tooth-bearing bone
Reptilia - Scincidae
cf. Scincidae informal indet. Morphotype 1
Bolet 2023
IPS128814, partial right maxilla.
cf. Scincidae informal indet. Morphotype 2
Bolet 2023
IPS128800, fragment of right maxilla.
Reptilia - Testudines - Podocnemididae
Neochelys sp. Bergounioux 1954
Reptilia - Testudines - Testudinidae
? Hadrianus sp. Cope 1872
    = Pelorochelon soriana n. gen., n. sp. Pérez-García et al. 2016
Pérez-García et al. 2016
STUS 14223, holotype, almost complete shell
Reptilia - Testudines - Trionychidae
    = Trionychidae indet. Gray 1825
Ortega et al. 2022
see common names

Geography
Country:Spain State/province:Castilla y Leon
Coordinates: 41.5° North, 2.1° West (view map)
Paleocoordinates:39.6° North, 5.3° West
Basis of coordinate:based on political unit
Altitude:926 meters
Geographic resolution:local area
Time
Period: Paleogene Epoch: Eocene
Stage: Bartonian 10 m.y. bin: Cenozoic 3
Key time interval: Bartonian
Age range of interval: 41.2 - 37.71 m.y. ago
Stratigraphy
Formation:Mazaterón
Stratigraphic resolution:group of beds
Stratigraphy comments: late Robiacian (Eocene European Land Mammal Age) in age, corresponding to the MP 15-16 Paleogene reference level
Lithology and environment
Primary lithology:gray marl
Secondary lithology: sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: The deposit is in carbonate facies, in small sandstone or conglomeratic lenses, extending 400 m laterally, and 5-8 m thick. the grey marls that contain the vertebrate fossils are included within the Mazaterón Formation, which consists of lacustrine-palustrine limestones and marls
Environment:lacustrine indet.
Geology comments: depositional setting is a shallow lacustrine system, which was subjected to constant flooding and drying processes (e.g., Alonso-Gavilán et al., 2004; Huerta and Armenteros, 2006). In terms of the European Land Mammal Ages, it is Robiacian (Mammal Paleogene [MP] 15– 16) in age (Cuesta, 1991, 1999; Antunes et al., 1997).
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Metadata
Also known as:La Solana
Database number:40358
Authorizer:J. Alroy, P. Holroyd, P. Mannion, J. Head, R. Butler Enterer:J. Wertheim, R. Butler, A. Garcia Selles, P. Holroyd, P. Mannion, G. Varnham
Modifier:M. Carrano Research group:vertebrate
Created:2004-06-22 11:24:41 Last modified:2022-06-17 13:47:07
Access level:the public Released:2004-06-22 11:24:41
Creative Commons license:CC BY
Reference information

Primary reference:
10894. E. Jiménez Fuentes, M. A. Cuesta, S. Martín Jesús, E. Mulas, E. Pérez Ramos and S. Jiménez-García. 1989. Primera nota sobre los vertebrados del Eoceno Superior de Matazarón y Deza (Soria). Stvdia Geologica Salmanticensia 5:129-133 [J. Alroy/J. Wertheim/J. Wertheim]

Secondary references:

60598 A. Badiola and M.-A. Cuesta. 2008. New endemic Eocene equoids from the Iberian Peninsula (western Europe). Journal of Vertebrate Paleontology 28(4):1149-1161 [R. Butler/R. Butler]
85236 A. Bolet. 2023. Lizards and amphisbaenians (Reptilia, Squamata) from the middle Eocene of Mazaterón (Soria, Spain). The Anatomical Record [J. Head/G. Varnham]
60663 M. A. Cuesta. 1993. Los Palaeotheriidae (Perissodactyla, Mammalia) del Eoceno de la Cuenca del Duero (Castilla y Leon, Espana). Estudios Geológicos 49:87-109 [R. Butler/R. Butler]
37053 M. A. Cuesta and A. Badiola. 2009. Duerotherium sudrei gen. et sp. nov., a new anoplotheriine artiodactyl from the Middle Eocene of the Iberian Peninsula. Journal of Vertebrate Paleontology 29(1):303-308 [M. Shalap/M. Shalap/R. Butler]
10895 M. A. Cuesta Ruiz-Colmenares. 1992. Creodontos del Paleógeno de Castilla y León. Vertebrados Fosiles de Castilla y Leon, Museo de Salamanca 101-104 [J. Alroy/J. Wertheim/J. Wertheim]
11035 M. A. Cuesta Ruiz-Colmenares. 1992. Perisodáctilos del Paleógeno de Castilla y León. Vertebrados fósiles de Castilla y León 111-120 [J. Alroy/J. Wertheim/J. Wertheim]
60665 M. A. Cuesta Ruiz-Colmenares. 1993. Cuatro formas de Artiodactyla (Mammalia) del yaciemiento eocene de Mazateron (Cuenca del Duero, Soria, España). Treb. Mus. Geol. Barcelona 3:81-90 [R. Butler/R. Butler]
11036 M. A. Cuesta Ruiz-Colmenares. 1993. Los Palaeotheriidae (Perissodactyla, Mammalia) del Eoceno de la Cuenca del Duero (Castilla y Leon, España). Estudios Geologicos 49:87-109 [J. Alroy/J. Wertheim/J. Wertheim]
11061 M. A. Cuesta Ruiz-Colmenares. 1994. Los Plagiolophinae (Remy, 1976) nuevo rango (Perissodactyla, Mammalia) del Eoceno de la Cuenca del Duero (Castilla y Leon, España). Estudios Geológicos 50:253-279 [J. Alroy/J. Wertheim/J. Wertheim]
61363 M. A. Cuesta Ruiz-Colmenares. 1996. Primeros hallazgos de Lophiodontidae (Perissodactyla, Mammalia) en el yacimiento eocénico de Mazaterón (Cuenca del Duero). Stvdia Geologica Salmanticensia 32:39-48 [P. Mannion/P. Mannion]
11034 M. A. Cuesta Ruiz-Colmenares. 1998. Presencia de Leptotheridium (Dacrytheriidae, Artiodactyla, Mammalia) en el yacimiento Eocenico de Caenes (Cuenca del Duero, Salamanca, España). Studia Geologica Salmanticensia 34:69-78 [J. Alroy/J. Wertheim/J. Wertheim]
46136 J. Marigó, R. Minwer-Barakat, and S. Moyà-Solà. 2010. New Anchomomyini (Adapoidea, Primates) from the Mazaterón Middle Eocene locality (Almazán Basin, Soria, Spain). Journal of Human Evolution 58(2):353-361 [J. Alroy/A. Garcia Selles]
46143 R. Minwer-Barakat, J. Marigó, and S. Moyà-Solà. 2012. Pseudoloris cuestai, a New Microchoerine (Primates, Omomyidae) from the Middle Eocene of the Iberian Peninsula. Journal of Vertebrate Paleontology 32(2):407-418 [J. Alroy/A. Garcia Selles]
80949 F. Ortega, I. Armenteros, A. Celis, F. Escaso, P. Huerta, S. M. Jesús, I. Narváez, A. Pérez-García, and J. L. Sanz. 2022. Crocodyliformes and Testudines from the Eocene of the Duero Basin (northwestern Spain): an update of their diversity and stratigraphic context. Historical Biology 34(8):1560-1581 [P. Mannion/G. Varnham/P. Mannion]
81942 F.-J. Ortega Coloma. 1992. Excavación, restauración y gestión de macrovertebrados fósiles [Excavation, restoration and management of macrovertebrate fossils]. In E. Jiménez-Fuentes (ed.), Vertebrados Fósiles de Castilla y León 13-25 [M. Carrano/M. Carrano]
60597 P. Peláez-Campomanes. 1996. Upper Eocene rodents from the Almazan basin (Soria, Spain). Eclogae Geologicae Helvetiae 89(3):1363-1385 [R. Butler/R. Butler]
81458 L. Perales-Gogenola, A. Badiola, A. Gómez-Olivencia and X. Pereda-Suberbiola. 2021. New Leptolophus (Palaeotheriidae) species from the Iberian Peninsula and early evidence of hypsodonty in an Eocene perissodactyl. Journal of Vertebrate Paleontology 41(1):e1912061 [P. Mannion/G. Varnham]
59713 A. Pérez-García, F. Ortega, and E. Jimenez Fuentes. 2016. Taxonomy, systematics, and diversity of the European oldest testudinids. Zoological Journal of the Linnean Society 177:648-675 [P. Holroyd/P. Holroyd/P. Mannion]