USGS 6019b, Canal Zone: Aquitanian, Panama
collected by D.F. MacDonald, T.W. Vaughan 1911

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Scaphopoda - Dentaliida - Dentaliidae
Dentalium (Fissidentalium) uscarianum Olsson 1922
Woodring 1973 f category
recombined as Fissidentalium uscarianum
Gastropoda - Heterostropha - Architectonicidae
Architectonica (Architectonica) nobilis (Röding 1798)
Woodring 1970 r category
original and current combination Architectonica nobilis
Gastropoda - Opisthobranchia - Cylichnidae
Scaphander (Scaphander) cryus
Woodring 1970 r category
Gastropoda - Cerithiidae
Bittium scotti Brown and Pilsbry 1913
Woodring 1970 c category
Gastropoda - Turritellidae
Turritella venezuelana Hodson 1926
r category
Woodring changes identification from previous monograph
Turritella (? Bactrospira) amaras
a category
Woodring changes identification from previous monograph
Gastropoda - Neotaenioglossa - Calyptraeidae
Crepidula sp. Lamarck 1799
r category
Crucibulum sp. Schumacher 1817
Calyptraea cf. centralis (Conrad 1841)
c category
Gastropoda - Naticidae
? Neverita sp. Risso 1826
a category
Gastropoda - Rissoidae
Rissoina (? Zebinella) sp. (Mörch 1876)
r category
Gastropoda - Neogastropoda - Columbellidae
? Mitrella sp. Risso 1826
Woodring 1964 r category
Gastropoda - Neogastropoda - Fasciolariidae
? Fusinus sp. Rafinesque 1815
Woodring 1964 r category
Gastropoda - Neogastropoda - Nassariidae
Nassarius (? Uzita) praeambiguus (Brown and Pilsbry 1913)
Woodring 1964 c category
Gastropoda - Neogastropoda - Melongenidae
Melongena sp. Schumacher 1817
Woodring 1964 r category
Gastropoda - Neogastropoda - Olividae
? Olivella sp. Swainson 1831
Woodring 1964 r category
Oliva (Oliva) sp. Bruguière 1789
Woodring 1970 f category
Gastropoda - Neogastropoda - Marginellidae
Prunum (Microspira) aff. apalachee (Gardner 1947)
Woodring 1970 r category
Marginella (Eratoidea) sp. (Weinkauff 1879)
Woodring 1970 r category
original and current combination Eratoidea
Bivalvia - Nuculanida - Nuculanidae
Adrana sp. Adams and Adams 1858
Woodring 1973 r category
Bivalvia - Pectinida - Pectinidae
Lepidopecten proterus Woodring 1982
Woodring 1982 a category
Bivalvia - Arcida - Arcidae
Anadara (Rasia) dariensis (Brown and Pilsbry 1911)
Woodring 1973 c category
Subspecies: Anadara (Rasia) dariensis progonica
Anadara (Grandiarca) balboai
Woodring 1973 a category
Bivalvia - Cardiida - Cardiidae
Trigoniocardia (Apiocardia) cf. simrothi (Dall 1900)
Woodring 1982 c category
Trachycardium (Dallocardia) dominicense (Gabb 1873)
Woodring 1982 c category
Subspecies: Trachycardium (Dallocardia) dominicense dominicense
Laevicardium laevigatum (Linnaeus 1758)
Woodring 1982 r category
replaced by Laevicardium serratum
Bivalvia - Cardiida - Donacidae
Donax gonoides
Woodring 1982 f category
Bivalvia - Cardiida - Tellinidae
Tellina (Eurytellina) ophiaca Woodring 1982
Woodring 1982 c category
Bivalvia - Cardiida - Psammobiidae
Gari (Gobraeus) sp. (Brown 1844)
Woodring 1982 r category
Bivalvia - Cardiida - Veneridae
Lirophora (Lirophora) cf. quirosensis (Hodson 1927)
Woodring 1982 r category
Ventricolaria harrisiana (Olsson 1922)
Woodring 1982 c category
Chione (Chionopsis) eurylopas Woodring 1982
Woodring 1982 c category
Chione (Chionopsis) propinqua (Spieker 1922)
Woodring 1982 a category
Pitar (Pitar) aff. quirosanus (Hodson 1927)
Woodring 1982 r category
Tivela (Tivela) mactroides (Born 1778)
Woodring 1982 r category
Clementia (Clementia) dariena (Conrad 1855)
Woodring 1982 r category
Subspecies: Clementia (Clementia) dariena dariena
Bivalvia - Cardiida - Mactridae
Harvella sp. (Gray 1853)
Woodring 1982 r category
Bivalvia - Pholadida - Corbulidae
Bothrocorbula (Hexacorbula) cruziana (Olsson 1932)
Woodring 1982 a category
Varicorbula cf. vieta (Guppy 1866)
Woodring 1982 c category
Bivalvia - Lucinida - Lucinidae
Lucinisca cf. calhounensis (Dall 1903)
Woodring 1982 r category
Bivalvia - Carditida - Carditidae
Carditamera cf. tegea (Dall 1903)
Woodring 1982 f category
Malacostraca - Decapoda - Portunidae
Callinectes sp. Stimpson 1860
Rathbun 1918
Malacostraca - Decapoda - Panopeidae
Panopeus sp. Milne Edwards 1834
Rathbun 1918
Malacostraca - Decapoda - Callianassidae
Callianassa scotti Brown and Pilsbry 1913
Rathbun 1918
recombined as Neocallichirus scotti
see common names

Geography
Country:Panama
Coordinates: 9.1° North, 79.7° West (view map)
Paleocoordinates:8.1° North, 77.3° West
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Aquitanian 10 m.y. bin:Cenozoic 5
Key time interval:Aquitanian
Age range of interval:23.03000 - 20.44000 m.y. ago
Age estimate:21.6 Ma (Sr isotope)
Stratigraphy
Formation:Culebra
Stratigraphic resolution:bed
Stratigraphy comments: LITHOSTRATIGRAPHY: From the Culebra Formation, which unconformably overlies the Las Cascadas agglomerate; reassigned to La Boca Fm in subsequent publications of Woodring (e.g., 1973, 1982), but further emended by Kirby et al. (2009) to the lower member of the Culebra Formation. AGE: Early Miocene in text; direct date extrapolated from Kirby et al. (1999), =Aquitanian.
Lithology and environment
Primary lithology:planar lamination,fine poorly lithified "shale"
Secondary lithology: carbonaceous mudstone
Includes fossils?Y
Includes fossils?Y
Lithology description: Dark, stratified tuff and clay
Environment:marine indet.
Geology comments: No paleoenvironmental data reported.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:selective quarrying,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:USNM
Collectors:D.F. MacDonald, T.W. Vaughan Collection dates:1911
Collection method comments: Collected by MacDonald and Vaughan (1911). Material is also held (but not specified for any specific collection) at Stanford University, Acad. Nat. Sci. Phil., Corenell University, PRI, U. Cal. and Cal. Acad. Sci. Locality now unaccessible (submerged).
Taxonomic list comments:Exhaustive for gastropoda.
Metadata
Also known as:Woodring Loc. 99b
Database number:42631
Authorizer:A. Hendy, M. Clapham Enterer:A. Hendy, M. Clapham
Modifier:M. Clapham Research group:marine invertebrate
Created:2004-08-03 11:02:29 Last modified:2022-07-20 12:01:31
Access level:the public Released:2004-08-03 11:02:29
Creative Commons license:CC BY
Reference information

Primary reference:

11306. W. P. Woodring. 1957. Geology and Paleontology of Canal Zone and adjoining parts of Panama. United States Geological Survey Professional Paper 306(A) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

43691 M. X. Kirby, D. S. Jones, and B. J. MacFadden. 2009. Lower Miocene stratigraphy along the Panama Canal and its bearing on the Central American Peninula. PLoS One 3(7):1-14 [A. Hendy/A. Hendy]
43320decapod M. J. Rathbun. 1918. Decapod crustaceans from the Panama region. Bulletin of the United States National Museum 103:123-184 [C. Schweitzer/S. Yost/M. Clapham]
13232 W. P. Woodring. 1964. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Columbellidae to Volutidae). United States Geological Survey Professional Paper 306(C):241-297 [A. Miller/A. Hendy/A. Hendy]
13236 W. P. Woodring. 1970. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). United States Geological Survey Professional Paper 306(D):299-452 [A. Miller/A. Hendy/A. Hendy]
13235 W. P. Woodring. 1973. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae). United States Geological Survey Professional Paper 306(E):453-539 [A. Miller/A. Hendy/A. Hendy]
13359 W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy]