Los Alamitos vertebrates, middle section: Late/Upper Campanian - Early/Lower Maastrichtian, Argentina
collected by J. F. Bonaparte, J. E. Powell 1982-1983

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Chondrichthyes - Batoidei
Batoidei indet.
Cione 1987 12 elements
MACN-RN 148, twelve vertebrae
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteidae indet. Cuvier 1825
Cione 1987 207 elements
MACN-RN 150, 156, six vertebrae, skulls fragments and 200 dermal scales
cf. Atractosteus sp. Rafinesque 1820
Cione 1987 3 elements
MACN-RN 149, three teeth
Actinopteri - Perciformes
Percoidei indet. (Bleeker 1859)
Cione 1987 1 element
MACN-RN 155, tooth
Actinopteri - Spariformes - Sparidae
Sparidae indet. (Cuvier 1817)
Cione 1987 7 elements
"Lepidotes or Sparidae"
Actinopteri - Siluriformes
Siluriformes indet.
Cione 1987 75 elements
MACN-RN 154, 1139-1140, spines
Actinopteri - Siluriformes - Ariidae
cf. Ariidae indet. Berg 1958
Cione 1987 3 elements
MACN-RN 153, three pectoral spines fragments
Actinopteri - Siluriformes - Diplomystidae
cf. Diplomystidae indet. Eigenman 1890
Cione 1987 17 elements
MACN-RN 152, eight proximal fragments, eight central fragments and one pectoral spine
Amphibia - Temnospondyli - Pipidae
cf. Xenopus sp. Wagler 1827
Báez 1987 8 elements
    = Kuruleufenia xenopoides Gómez 2016
Suazo Lara and Gómez 2022
MACN-RN 159, fragment of skull, presacral, ilium, humeri, sphenethmoid, sacrourostyle, pelvis
Amphibia - Temnospondyli - Calyptocephalellidae
Leptodactylidae indet. Stirton 1953
Báez 1987 8 specimens
    = Calyptocephalellidae indet. Reig 1960
Suazo Lara and Gómez 2022
MACN-RN 160, maxillae, squamosal, humeri, ilium
Reptilia - Madtsoiidae
Madtsoiinae indet. Hoffstetter 1961
Pol et al. 2023 26 specimens
spelled with current rank as Madtsoiidae
MACN-RN 201-202, 204-227
Rionegrophis madtsoioides n. gen., n. sp. Albino 1986
Albino 1986 1 specimen
MACN-RN 32
Patagoniophis parvus n. gen., n. sp. Albino 1986
Albino 1986 3 specimens
MACN-RN 33-35
Alamitophis argentinus n. gen., n. sp. Albino 1986
Albino 1986 6 specimens
MACN-RN 27 (holotype) 27–31, 203
Reptilia
Aves indet. (Linnaeus 1758)
Agnolin and Martinelli 2009 5 specimens
MACN PV RN 1103 (distal end of indeterminate pedal phalanx), MACN PV RN 1104 (distal end of pedal phalanx, probably II-2), MACN PV RN 1105 (pedal ungual of digit I) and MACN PV RN 1106 (manual ungual of digit I); MACN PV RN 1107 (pedal phalanx 1 of right digit II without its proximal end)
cf. Neornithes indet. (Gadow 1893)
Agnolin and Martinelli 2009 1 specimen
MACN PV RN 1115 (distal end of right tarsometatarsus without metatarsal IV and distal trochlea of metatarsal II)
Ornithothoraces indet. Chiappe and Calvo 1994
Chiappe 1996 1 specimen
MACN-RN 976 (partial cervical vertebra)
Ornithurae indet. (Haeckel 1866)
Agnolin and Martinelli 2009 3 specimens
MACN PV RN 1111 (distal end of left tibiotarsus, without distal condyles), MACN PV RN 1112 (distal end of left tibiotarsus with damaged medial condyle) and MACN PV RN 1113 (distal end of left tibiotarsus with complete distal condyles)
Reptilia - Hesperornithiformes
cf. Hesperornithiformes indet. Fürbringer 1888
Agnolin and Martinelli 2009 1 specimen
MACN PV RN 1114 (distal end of left tibia)
Reptilia
Alamitornis minutus n. gen., n. sp. Agnolin and Martinelli 2009
Agnolin and Martinelli 2009 3 specimens
MACN PV RN 1108 - holotype (proximal third of a left humerus) and referred material: MACN PV RN 1109 (five proximal extremes of right humeri of different individuals) and MACN PV RN 1110 (distal half of a left femur of a juvenile specimen)
Reptilia - Titanosauridae
Aeolosaurus ? rionegrinus Powell 1987
Powell 1987 1 individual
MACN-RN 147
    = Titanosauridae indet. Lydekker 1885
Juárez Valieri and Calvo 2011
Titanosauridae indet. Lydekker 1885
Powell 1987 5 specimens
MACN-RN 1102, 1128
Reptilia - Hadrosauridae
Kritosaurus sp. Brown 1910
20 individuals
MACN-RN 142-146, 826, 977-995, 997-998
    = Kritosaurus australis n. sp. Bonaparte et al. 1984
Bonaparte et al. 1984
recombined as Huallasaurus australis
MACN-RN 2
Reptilia - Sphenodontidae
Sphenodontidae indet. Cope 1871
Apesteguía 2005 2 specimens
MACN Pv RN1099c, 1100
Kawasphenodon expectatus n. gen., n. sp. Apesteguía 2005
Apesteguía 2005 3 specimens
MACN Pv RN1098, 1099a-b
Reptilia
Niolamia sp. Ameghino 1899
de Broin 1987 20 specimens
MACN-RN 36-42, 65-76, 101
    = cf. Niolamia sp. Ameghino 1899
de Broin and de la Fuente 1993
additionally a fragment of nuchal (MACN RN 918), a short cervical plus long fragment of pleural (?) (MACN RN 910)
    = Meiolaniformes indet. Sterli and de la Fuente 2013
Sterli et al. 2013
The revision of the MACN collection allows us to corroborate the assignment of these specimens to Meiolaniformes by Broin (1987) and Broin and de la Fuente (1993)
Reptilia - Testudines
Chelidae informal indet. A Gray 1825
de Broin 1987 1 specimen
    = Palaeophrynops patagonicus n. gen., n. sp. Lapparent de Broin and de la Fuente 2001
Lapparent de Broin and de la Fuente 2001
MACN-RN 906. Anterior part of a carapace, morphology mostly seen in imprint, with the anterior border (partial peripherals 1 to 5, left and right), the nuchal, pleurals 1 and 2, neural 1 and 2 (partly); right lateral fragment with lateral part of pleurals 4 to 6 and peripherals 7 and 8; partial plastral anterior lobe, remains of 5 cervicals (only the 4th sufficiently preserved); left humerus and femur. Discovered by Dr Bonaparte. Lapparent de Broin & Fuente 2001 do not say which of the Los ALamitos chelid types P. patagonicus pertains to so I ahve arbitrarily placed in the 'indet A'
Reptilia - Testudines - Chelidae
Chelidae informal indet. D Gray 1825
de Broin 1987 33 specimens
The abundance is estimated, should be checked. MACN-RN 43-64, 77-100, 102-140
Chelidae informal indet. C Gray 1825
de Broin 1987 33 specimens
The abundance is estimated, should be checked. MACN-RN 43-64, 77-100, 102-140
    = cf. Yaminuechelys sp. de la Fuente et al. 2001
Maniel 2015
MACN-RN-43. Placa pygal plate. MACN-RN-45. 9th and 10th peripheral plates. MACN-RN-46.Posterior peripheral plate. MACN-RN-50. anterior peripheral plate. MACN-RN-63. posterior peripheral plate. MACN-RN-78. posterior peripheral plate. MACN-RN-86. 10th right peripheral plate. MACN-RN-88. posterior peripheral plate MACN-RN-123. neural fragment. MACN-RN-913. anterior peripheral plate. MACN-RN-914. Peripheral plate. MACN-RN-915. peripheral plate MACN-RN-926. xiphiplastron. MACN-RN927. xiphiplastron
Chelidae informal indet. B Gray 1825
de Broin 1987 33 specimens
The abundance is estimated, should be checked. MACN-RN 43-64, 77-100, 102-140
    = aff. Emydura sp. Bonaparte 1836
Maniel 2015
MACN-RN-907 and MACN-RN-908, two biconvex vertebra
Mammalia - Dryolestida - Casamiquelidae
Rougietherium tricuspes n. gen., n. sp. Bonaparte 2002
Bonaparte 2002 1 specimen
MACN-RN 162
Alamitherium bishopi n. gen., n. sp. Bonaparte 2002
Bonaparte 2002 3 specimens
MACN-RN 1034-1036
Mammalia - Dryolestida
Paraungulatum rectangularis n. gen., n. sp. Bonaparte 2002
Bonaparte 2002 3 specimens
MACN-RN 180 (left upper molar), 1141
Mammalia - Meridiolestida
Meridiolestida indet. Rougier et al. 2011
Gaetano et al. 2013 1 specimen
MACNPV-RN241, complete right upper premolariform
Mammalia - Meridiolestida - Mesungulatidae
Mesungulatidae indet. Bonaparte 1986
Gaetano et al. 2013 5 specimens
MACN-RN 239, complete left P1; MACN-RN 240, complete left p1, MACN-RN 1135-1137
Mesungulatum houssayi n. gen., n. sp. Bonaparte and Soria 1985
Bonaparte 1987 17 specimens
MACN-RN 1, 3-12, 141, 181-184, 244
Mammalia - Meridiolestida
Austrotriconodon mckennai n. gen., n. sp. Bonaparte 1986
Bonaparte 1986 6 specimens
MACN-RN 21 (holotype) 234-238
Mammalia - Dryolestidae
Groebertherium indet. Bonaparte 1986
Pol et al. 2023 16 specimens
MACN-RN 185-200
Groebertherium stipanicici n. gen., n. sp. Bonaparte 1986
Bonaparte 1986 5 specimens
MACN-RN 13-17
Groebertherium novasi n. gen., n. sp. Bonaparte 1986
Bonaparte 1986 2 specimens
MACN-RN 18-19
Mammalia
Paratheria indet. Thomas 1887
Bonaparte 1987
Ferugliotherium sp. Bonaparte 1986
Kielan-Jaworowska et al. 2004 2 specimens
MACN-RN 249 (anterior upper premolar), and framgnetary left dentary
Ferugliotherium windhauseni n. gen., n. sp. Bonaparte 1986
Bonaparte 1986 9 specimens
MACN-RN 20 (holotype worn right m2) and specimen referred by Krause et al 1992: MACN-RN 701A, B (partial left i1s), 701C (partial i1), 253 (partial left m1), 702A–D (partial I2s), 248-251, MACN-RN 975; incisors and premolars only tentatively referred
Mammalia - Sudamericidae
Gondwanatherium patagonicum n. gen., n. sp. Bonaparte 1986
Bonaparte 1986 18 specimens
MACN-RN 22 (holotype) 22-26, 228, 703-705, 970, 1024-1031
Dipnoi - Ptychoceratodontidae
Ptychoceratodus cionei n. sp. Apesteguia et al. 2007
Apesteguia et al. 2007 2 specimens
MACN PV RN157A and 157B, two lower tooth plates
Dipnoi - Ceratodontidae
Ameghinoceratodus iheringi (Ameghino 1906)
Apesteguia et al. 2007 1 specimen
MACN PV RN157C, right lower tooth plate
    = Atlantoceratodus patagonicus Agnolín 2010
Agnolín 2010
Ceratodus iheringi Ameghino 1906
Cione 1987 118 elements
recombined as Atlantoceratodus iheringi
MACN-RN 157-158, dental plates
Ptychoceratodus wichmanni n. sp. Apesteguía and Agnolin 2007
Apesteguia et al. 2007 1 specimen
MACN PV RN157A, left upper tooth plate
    = Metaceratodus wichmanni n. sp. Apesteguía and Agnolin 2007
Cione and Gouiric-Cavalli 2012
see common names

Geography
Country:Argentina State/province:Río Negro
Coordinates: 42.1° South, 66.3° West (view map)
Paleocoordinates:45.0° South, 52.3° West
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Key time interval:Late/Upper Campanian - Early/Lower Maastrichtian
Age range of interval:83.60000 - 66.00000 m.y. ago
Stratigraphy
Formation:Los Alamitos
Stratigraphic resolution:group of beds
Stratigraphy comments: middle section of formation includes vertebrate fossils distributed over about 30 meters of section
Mammals: Middle section of Los Alamitos Formation - green-coloured bed just below the concretionary top of the Cerrito del Mamifero, Alamitian provincial age (Campanian sensu lato)
Lithology and environment
Primary lithology:planar lamination,bioturbation,concretionary,yellow siltstone
Secondary lithology:bioturbation,intraclastic sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: "Pelitic facies: includes dominant massive, laminated or rippled (rare) siltstones and claystones, with yellowish (lower part of the sequences) to reddish, orange, brown and purple hues (upper part), and massive sandstones. This rocks show typical paquidermis solution pattern which allow its recognition in outcrops. The non-calcareous pelites contains (lower half) almost all the vertebrate remains"
Environment:lacustrine - large
Geology comments: dominant environment lacustrine, more brackish than fresh (clay flocculation in pelites). waters were oxygenated; lake permanent and shallow.
The paleoenvironment according to Andreis (1987; and personnal communication) is that of a lacustrine environment mixed with shallow marine sediments, as indicated by the presnce of glauconite in some levels of the sequence.
Taphonomy
Modes of preservation:body,trace
Degree of concentration:dispersed
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Articulated whole bodies:none
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:frequent
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,surface (float),surface (in situ),mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:J. F. Bonaparte, J. E. Powell Collection dates:1982-1983
Metadata
Also known as:Los Alamitos vertebrates, middle section - green-coloured bed (mammals), Cerro Cuadrado, Arroyo Verde
Database number:45087
Authorizer:M. Carrano, P. Mannion, E. Vlachos, R. Benson Enterer:M. Carrano, P. Mannion, F. Aspromonte, E. Vlachos, R. Benson, J. Tennant
Modifier:F. Aspromonte Research group:vertebrate
Collections that are a subset of this one:231685, 231687, 231688, 231689, 231690, 231691, 231692, 231693, 231694, 231695, 231696
Created:2004-10-04 11:03:49 Last modified:2023-08-28 14:55:42
Access level:the public Released:2004-10-04 11:03:49
Creative Commons license:CC BY
Reference information

Primary reference:

13925.ETE J. F. Bonaparte. 1984. Nuevas pruebas de la conexion fisica entre Sudamerica y Norteamerica en el Cretácico Tardio (Campaniano) [New evidences for the physical connection between South and North America in the Late Cretaceous (Campanian)]. Actas del III Congreso Argentino de Paleontologia y Bioestratigrafia 1:141-149 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

86689 F. Agnolín. 2010. A new species of the genus Atlantoceratodus (Dipnoiformes: Ceratodontoidei) from the Uppermost Cretaceous of Patagonia and a brief overview of fossil dipnoans from the Cretaceous and Paleogene of South America. Brazilian Geographical Journal 1(2):162-210 [E. Vlachos/F. Aspromonte]
37643 F. L. Agnolin and A. G. Martinelli. 2009. Fossil birds from the Late Cretaceous Los Alamitos Formation, Río Negro Province, Argentina. Journal of South American Earth Sciences 27:42-49 [P. Mannion/P. Mannion/M. Carrano]
17612 A. M. Albino. 1986. Nuevos Boidae Madtsoiinae en el Cretácico Tardio de Patagonia (Formación Los Alamitos, Río Negro, Argentina) [New Madtsoiinae Boinae in the Late Cretaceous of Patagonia (Los Alamitos Formation, Río Negro, Argentina)]. Actas del IV Congreso Argentino de Paleontología y Bioestratigrafía, Mendoza 2:15-21 [J. Head/J. Head/M. Carrano]
11736ETE A. M. Albino. 1987. The Late Cretaceous fauna of Los Alamitos, Patagonia, Argentina part V—the ophidians. Revista del Museo Argentina de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales: Paleontología 3(3):141-146 [M. Carrano/M. Carrano/M. Carrano]
11733ETE R. R. Andreis. 1987. The Late Cretaceous fauna of Los Alamitos, Patagonia, Argentina part I—stratigraphy and paleontology. Revista del Museo Argentina de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales: Paleontología 3(3):103-110 [M. Carrano/M. Carrano/M. Carrano]
34191ETE S. Apesteguía. 2005. A Late Campanian sphenodontid (Reptilia, Diapsida) from northern Patagonia. Comptes Rendus Palevol 4:663-669 [M. Carrano/M. Carrano]
72409 S. Apesteguia, F. L. Agnolin, and K. Claeson. 2007. Review of Cretaceous dipnoans from Argentina (Sarcopterygii: Dipnoi) with descriptions of new species. Revista del Museo Argentino de Ciencias Naturales nueva serie 9(1):27-40 [G. Lloyd/G. Lloyd/F. Aspromonte]
11734ETE A. M. Báez. 1987. The Late Cretaceous fauna of Los Alamitos, Patagonia, Argentina part III—anurans. Revista del Museo Argentina de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales: Paleontología 3(3):121-130 [M. Carrano/M. Carrano/F. Aspromonte]
30646ETE J. F. Bonaparte. 1984. Jurassic and Cretaceous terrestrial vertebrates of South America. National Geographic Society Research Reports (1975) 16:115-125 [M. Carrano/M. Carrano]
9952 J. F. Bonaparte. 1986. Sobre Mesungulatum houusayi y nuevos mamíferos Cretácicos de Patagonia, Argentina [On Mesungulatum houssayi and new Cretaceous mammals from Patagonia, Argentina]. Actas del IV Congreso Argentino de Paleontología y Biostratigrafía 2:48-61 [J. Alroy/J. Wertheim/M. Carrano]
13928ETE J. F. Bonaparte. 1986. A new and unusual Late Cretaceous mammal from Patagonia. Journal of Vertebrate Paleontology 6(3):264-270 [M. Carrano/M. Carrano/M. Carrano]
11737ETE J. F. Bonaparte. 1987. The Late Cretaceous fauna of Los Alamitos, Patagonia, Argentina part VIII—the mammals. Revista del Museo Argentina de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales: Paleontología 3(3):163-169 [M. Carrano/M. Carrano/M. Carrano]
38756 J. F. Bonaparte. 1990. New Late Cretaceous mammals from the Los Alamitos Formation, Northern Patagonia. National Geographic Research 6(1):63-93 [R. Benson/R. Benson]
16547ETE J. F. Bonaparte. 1996. Cretaceous tetrapods of Argentina. Münchner Geowissenschaften Abhandlungen 30:73-130 [M. Carrano/M. Carrano/M. Carrano]
70619 J. F. Bonaparte. 2002. New Dryolestida (Theria) from the Late Cretaceous of Los Alamitos, Argentina, and paleogeographical comments. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 224(3):339-371 [M. Carrano/M. Carrano/P. Wagner]
13926ETE J. F. Bonaparte, M. R. Franchi, J. E. Powell and E. Sepulveda. 1984. La Formación Los Alamitos (Campaniano-Maastrichtiano) del sudeste de Rio Negro, con descripcion de Kritosaurus australis n. sp. (Hadrosauridae). Significado paleogeografico de los vertebrados [The Los Alamitos Formation (Campanian-Maastrichtian) from the southeast of Rio Negro, with a description of Kritosaurus australis n. sp. (Hadrosauridae). Paleogeographic significance of the vertebrates]. Revista de la Asociación Geología Argentina 39(3-4):284-299 [M. Carrano/M. Carrano/M. Carrano]
11732ETE J. F. Bonaparte and G. Rougier. 1987. The Late Cretaceous fauna of Los Alamitos, Patagonia, Argentina part VII—the hadrosaurs. Revista del Museo Argentina de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales: Paleontología 3(3):155-161 [M. Carrano/M. Carrano/M. Carrano]
37378 C. R. A. Candeiro. 2010. Record of the genus Aeolosaurus (Sauropoda, Titanosauria) in the Late Cretaceous of South America: paleogeographic implications. Estudios Geológicos 66(2):243-253 [M. Carrano/M. Carrano]
12947ETE L. M. Chiappe. 1996. Early avian evolution on the southern hemisphere: the fossil record of birds in the Mesozoic of Gondwana. Memoirs of the Queensland Museum 39(3):533-554 [M. Carrano/M. Carrano/M. Carrano]
48178 L. M. Chiappe. 2007. Aves. In Z. B. Gasparini, L. Sangado & R. A. Coria (eds.), Patagonian Mesozoic Reptiles 257-270 [M. Carrano/M. Carrano]
11738ETE A. L. Cione. 1987. The Late Cretaceous fauna of Los Alamitos, Patagonia, Argentina part II—the fishes. Revista del Museo Argentina de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales: Paleontología 3(3):111-120 [M. Carrano/M. Carrano/M. Carrano]
86688 A. L. Cione and S. Gouiric-Cavalli. 2012. Metaceratodus kaopen comb. nov. and M. wichmanni comb. nov., two Late Cretaceous South American species of an austral lungfish genus (Dipnoi). Alcheringa 36(2):203-216 [E. Vlachos/F. Aspromonte]
75729 P. Cruzado-Caballero, L. S. Filippi, A. H. Méndez, A. C. Garrido, and I. Díaz-Martínez. 2018. First ornithopod remains from the Bajo de la Carpa Formation (Santonian, Upper Cretaceous), northern Patagonia, Argentina. Cretaceous Research 83:182-193 [M. Carrano/M. Carrano/M. Carrano]
11739ETE F. de Broin. 1987. The Late Cretaceous fauna of Los Alamitos, Patagonia, Argentina part IV—Chelonia. Revista del Museo Argentina de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales: Paleontología 3(3):131-139 [M. Carrano/M. Carrano/M. Carrano]
42415 F. de Broin and M. de la Fuente. 1993. Les tortues fossiles d'Argentine: synthese [The fossil turtles of Argentina: synthesis]. Annales de Paléontologie (Invertebrés-Vertebrés) 79(3):169-232 [R. Benson/R. Benson/M. Carrano]
49730 L. C. Gaetano, C. A. Marsicano, and G. W. Roughier. 2013. A revision of the putative Late Cretaceous triconodonts from South America. Cretaceous Research 46:90-100 [P. Mannion/J. Tennant]
23486ETE J. R. Horner, D. B. Weishampel, and C. A. Forster. 2004. Hadrosauridae. In D. B. Weishampel, H. Osmolska, and P. Dodson (eds.), The Dinosauria (2nd edition). University of California Press, Berkeley 438-463 [M. Carrano/M. Carrano/M. Carrano]
34687ETE R. D. Juárez Valieri and J. O. Calvo. 2011. Revision of MUCPv 204, a Senonian basal titanosaur from northern Patagonia. In J. O. Calvo, J. D. Porfiri, B. J. González Riga, D. Dos Santos (eds.), Dinosaurios y Paleontología desde América Latina. Anales del III Congreso Latinoamericano de Paleontología. Editorial de la Universidad Nacional de Cuyo, Neuquén 143-152 [M. Carrano/M. Carrano]
14365 Z. Kielan-Jaworowska, R. L. Cifelli, and Z.-X. Luo. 2004. Mammals from the age of dinosaurs: Origins, evolution, and structure. Columbia University Press, New York 1-630 [R. Whatley/R. Whatley/R. Whatley]
38915 D. W. Krause, Z. Kielan-Jaworowska, and J. F. Bonaparte. 1992. Ferugliotherium Bonaparte, the first known multituberculate from South America. Journal of Vertebrate Paleontology 12(3):351-376 [R. Benson/R. Benson]
17570 F. d.e. Lapparent de Broin and M. S. de la Fuente. 2001. Oldest world Chelidae (Chelonii, Pleurodira), from the Cretaceous of Patagonia, Argentina. C. R. Acad. Sci. Paris, Sciences de la Terre et des planètes 333:463-470 [P. Holroyd/P. Holroyd]
85727 I. Maniel. 2015. Anatomía, Diversidad y Sistemática de las tortugas Pleurodira del Cretácico de Patagonia. UNIVERSIDAD DE BUENOS AIRES Facultad de Ciencias Exactas y Naturales Departamento de Ecología, Genética y Evolución 1-496 [E. Vlachos/E. Vlachos/E. Vlachos]
38236 A. G. Martinelli, D. Riff, and R. P. Lopes. 2011. Discussion about the occurrence of the genus Aeolosaurus Powell 1987 (Dinosauria, Titanosauria) in the Upper Cretaceous of Brazil. Gaea - Journal of Geoscience 7(1):34-40 [P. Mannion/P. Mannion/M. Carrano]
85586 D. Pol, E. Vlachos, and F. Aspromonte. 2023. The end of the dinosaur era in Patagonia (NatGeo Project). [E. Vlachos/E. Vlachos/E. Vlachos]
11740ETE J. E. Powell. 1987. The Late Cretaceous fauna of Los Alamitos, Patagonia, Argentina part VI—the titanosaurids. Revista del Museo Argentina de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales: Paleontología 3(3):147-153 [M. Carrano/M. Carrano/M. Carrano]
13712ETE J. E. Powell. 2003. Revision of South American titanosaurid dinosaurs: palaeobiological, palaeobiogeographical and phylogenetic aspects. Records of the Queen Victoria Museum Launceston 111:1-173 [M. Carrano/M. Carrano/M. Carrano]
32771ETE A. Prieto-Marquez and G. C. Salinas. 2010. A re-evaluation of Secernosaurus koerneri and Kritosaurus asutralis (Dinosauria, Hadrosauridae) from the Late Cretaceous of Argentina. Journal of Vertebrate Paleontology 30(3):813-837 [M. Carrano/M. Carrano/M. Carrano]
47125 J. Sterli, M. S. de la Fuente, and I. A. Cerda. 2013. A new species of meiolaniform turtle and a revision of the Late Cretaceous Meiolaniformes of South America. Ameghiniana 50(2):240-256 [R. Benson/R. Benson]
85526 F. Suazo Lara and R. O. Gómez. 2022. In the shadow of dinosaurs: Late Cretaceous frogs are distinct components of a widespread tetrapod assemblage across Argentinean and Chilean Patagonia. Cretaceous Research 131:105085 [P. Mannion/P. Mannion]