KJ9239 (DMNH 517): Lancian, South Dakota
collected by S. Sacrison, K. Johnson 1987

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Bisonia niemii leaf
Johnson 2002 39 specimens
Johnson; HC200
Paleoaster inquirenda seed/fruit
Johnson 2002 2 specimens
HC7
Angiospermae - Arecaceae
Amesoneuron informal FU37 leaf
Johnson 2002 12 specimens
Hickey and Read; FU37
Angiospermae
Magnoliidae informal HC43 leaf
Johnson 2002 17 specimens
synonym of Magnoliid
HC43
Angiospermae - Laurales
? Laurales informal HC32 leaf
Johnson 2002 1 specimen
HC32
Angiospermae - Laurales - Lauraceae
Marmarthia trivialis leaf Johnson 1996
Johnson 2002 16 specimens
(Lesquereux) Johnson; HC105
Marmarthia pearsonii leaf Johnson 1996
Johnson 2002 21 specimens
Johnson; HC162
Angiospermae - Proteales - Platanaceae
Platanites marginata leaf (Lesquereux 1872)
Johnson 2002 44 specimens
recombined as Cissus marginata
HC1-106
Angiospermae - Proteales - Nelumbonaceae
Nelumbo informal FU62 leaf
Johnson 2002 1 specimen
FU62
Angiospermae - Rosales - Rhamnaceae
Ziziphus fibrillosus leaf
Johnson 2002 1 specimen
HC174
Rhamnus cleburni leaf Lesquereux 1872
Johnson 2002 32 specimens
HC62
Angiospermae - Fagales - Nothofagaceae
Dryophyllum tenneseensis leaf
Johnson 2002 10 specimens
HC44
Angiospermae - Malvales - Tiliaceae
Grewiopsis saportana leaf
Johnson 2002 1 specimen
HC2
Angiospermae
Dicotyledonae informal HC280 leaf
Johnson 2002 2 specimens
corrected as Dicotyledoneae
HC280
Dicotyledonae informal HC318 leaf
Johnson 2002 1 specimen
corrected as Dicotyledoneae
HC318
Angiospermae - Arales - Lemnaceae
Limnobiophyllum scutatum leaf Dawson 1875
Johnson 2002 1 specimen
(Dawson) Krassilov; FU38
Magnoliopsida
Quereuxia angulata leaf Newberry 1953
Johnson 2002 1 specimen
FU2
Pinopsida - Pinales - Taxaceae
Elatides longifolia leaf
Johnson 2002 21 specimens
HC165
Coniferales - Cupressaceae
Sequoia informal HC70 leaf
Johnson 2002 5 specimens
HC70
Reptilia - Tyrannosauridae
Tyrannosaurus rex Osborn 1905
1 individual
BHI 3033 ("Stan")
    = Tyrannosaurus regina Paul et al. 2022
Paul et al. 2022
synonym of Tyrannosaurus rex
see common names

Geography
Country:United States State/province:South Dakota County:Harding
Coordinates: 45.7° North, 103.6° West (view map)
Paleocoordinates:51.8° North, 77.3° West
Basis of coordinate:stated in text
Altitude:3090
Geographic resolution:small collection
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
*Period:Late/Upper Cretaceous *Epoch:Late/Upper Senonian
*International age/stage:Late/Upper Maastrichtian *Local age/stage:Lancian
Key time interval:Lancian Zone: HCIII
Age range of interval:72.10000 - 66.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Hell Creek
Stratigraphic resolution:bed
Stratigraphy comments: -16.2m to Hell Creek-Fort Union contact datum, -16.2m to K-T boundary based on calculations in Johnson (2002)
Marmarth Composite regional section, but no bed given
Lithology and environment
Primary lithology:paleosol/pedogenic poorly lithified sandstone
Includes fossils?Y
Lithology description: "Spatsiko" pedotype
Environment:fine channel fill Tectonic setting:cratonic basin
Taphonomy
Modes of preservation:body,adpression
Size of fossils:macrofossils
Spatial orientation:random
Associated major elements:many
Temporal resolution:snapshot
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,surface (in situ),mechanical,field collection
Collection size:228 specimens
Reason for describing collection:general faunal/floral analysis
Museum repositories:DMNH
Collectors:S. Sacrison, K. Johnson Collection dates:1987
Collection method comments: Specimens located at Denver Museum of Nature & Science
Metadata
Also known as:Stan T-rex site, KJ9239, 9308
Database number:48193
Authorizer:K. Johnson, M. Carrano Enterer:K. Johnson, M. Carrano
Modifier:M. Carrano Research group:paleobotany
Created:2005-03-15 17:56:31 Last modified:2021-12-15 10:11:01
Access level:the public Released:2005-03-15 17:56:31
Creative Commons license:CC BY
Reference information

Primary reference:
78884. G. J. Retallack. 1994. A pedotype approach to latest Cretaceous and earliest Tertiary paleosols in eastern Montana. Geological Society America Bulletin 106(11):1377-1397 [M. Carrano/M. Carrano]

Secondary references:

7941 K. R. Johnson. 2002. Megaflora of the Hell Creek and lower Fort Union Formations in the western Dakotas: Vegetational response to climate change, the Cretaceous-Tertiary boundary event, and rapid marine transgression. The Hell Creek Formation and the Cretaceous-Tertiary boundary in the northern Great Plains: An integrated continental record of the end of the Cretaceous. 361:329-391 [S. Wing/S. Wing/K. Johnson]
17042ETE P. L. Larson and R. A. Farrar. 1995. Cranial osteology and morphology of the Upper Cretaceous theropod, Tyrannosaurus rex. Journal of Vertebrate Paleontology 15(3, suppl.):40A [M. Carrano/M. Carrano/M. Carrano]
82296 G. S. Paul, W. S. Persons, and J. Van Raalte. 2022. The tyrant lizard king, queen and emperor: multiple lines of morphological and stratigraphic evidence support subtle evolution and probable speciation within the North American genus Tyrannosaurus. Evolutionary Biology 49(2):156-179 [M. Carrano/M. Carrano]
8329 P. Wilf, K. R. Johnson, and B. T. Huber. 2003. Correlated terrestrial and marine evidence for global climate changes before mass extinction at the Cretaceous-Paleogene boundary. Proceedings of the National Academy of Sciences 100:599-604 [P. Wilf/P. Wilf/P. Wilf]