Rohstoffbetriebe Oker quarry, bed 83, Langenberg: Late/Upper Kimmeridgian, Germany
collected by H. Ludtke 1998–

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Porifera indet. Grant 1836
Sander et al. 2006
Mammalia - Multituberculata - Pinheirodontidae
Teutonodon langenbergensis n. gen., n. sp. Martin et al. 2019
Martin et al. 2019 3 specimens
NLMH 105650 - holotype; referred material: NLMH 105656 and 105665
Mammalia - Multituberculata - Paulchoffatiidae
Paulchoffatiidae indet. Hahn 1969
Martin et al. 2019 2 specimens
NLMH 105655 and 105651 (the latter was previously attributed to Eobaataridae indet.)
Cimbriodon multituberculatus n. gen., n. sp. Martin et al. 2019
Martin et al. 2019 8 specimens
NLMH 105657 - holotype; referred material: NLMH 105658-105664
Reptilia - Testudines
Testudines indet. Batsch 1788
Sander et al. 2006
Reptilia - Theropoda
Theropoda indet. Marsh 1881
1 specimen
DFMMh/FV 705
Reptilia
cf. Ceratosauria indet. (Marsh 1884)
Evers and Wings 2020
chevron; DfMMh/FV/776
cf. Megalosauroidea indet. (Nopcsa 1928)
Evers and Wings 2020
fibula; DfMMh/FV/287
Reptilia - Megalosauridae
cf. Torvosaurus sp. Galton and Jensen 1979
Gerke and Wings 2016
two teeth; DFMMh/ FV1205, DFMMh/FV1206
Reptilia - Dromaeosauridae
Velociraptorinae indet. Barsbold 1983
van der Lubbe et al. 2009 6 specimens
DFMMh/FV 658, 707.1, 382, 383, 790.5, 530
Reptilia
Allosauroidea indet. Currie and Zhao 1994
Evers and Wings 2020
pedal ungual; DfMMh/FV1/19
Reptilia - Iguanodontidae
Iguanodontidae indet. Bonaparte 1850
2 specimens
tibia, fibula fragments
Reptilia
Sauropoda indet. (Marsh 1878)
11 individuals
(15 measurements)
    = Brachiosauridae indet. Riggs 1904
Windolf 2000
    = Europasaurus holgeri n. gen., n. sp. Sander et al. 2006
Sander et al. 2006
Reptilia - Diplodocidae
Diplodocidae indet. Marsh 1884
Windolf 2000
Reptilia
Pterodactylus sp. Cuvier 1809
Windolf 2000
Reptilia - Ctenochasmatidae
Ctenochasmatidae indet. Nopcsa 1928
Windolf 2000
new species
Reptilia
? Theriosuchus sp. Owen 1878
Windolf 2000 9 specimens
DFMMh/FV 200 and DFMMh/FV 605
    = Theriosuchus pusillus Owen 1878
Karl et al. 2006
    = Theriosuchus cf. pusillus Owen 1878
Tennant et al. 2016
    = Knoetschkesuchus langenbergensis n. gen., n. sp. Schwarz et al. 2017
Schwarz et al. 2017
+ DFMMh/FV 261, 790.12, 279, 790.11 and 325
Reptilia - Goniopholididae
Goniopholis sp. Owen 1842
Windolf 2000
Chondrichthyes - Rajiformes
Asterodermus informal sp. morphotype 15 Agassiz 1836
Thies 1995 1 specimen
nomen dubium belonging to Rhinobatoidei
Chondrichthyes - Hybodontiformes
Hybodontoidea informal morphotype 2
Thies 1995 1 specimen
Bivalvia
Bivalvia indet. Linnaeus 1758
Sander et al. 2006
Gastropoda
Gastropoda indet. Cuvier 1795
Sander et al. 2006
Ostracoda
Ostracoda indet. Latreille 1802
Sander et al. 2006
unclassified
Foraminifera indet. Eichwald 1830
Sander et al. 2006
Charales - Characeae
Characeae indet. Agardh 1824
Sander et al. 2006
see common names

Geography
Country:Germany State/province:Niedersachsen
Coordinates: 51.9° North, 10.5° East (view map)
Paleocoordinates:43.1° North, 18.4° East
Basis of coordinate:based on nearby landmark
Altitude:289 meters
Geographic resolution:outcrop
Time
Period:Jurassic Epoch:Late/Upper Jurassic
Stage:Kimmeridgian 10 m.y. bin:Jurassic 5
Key time interval:Late/Upper Kimmeridgian
Age range of interval:152.20000 - 149.20000 m.y. ago
Stratigraphy
Formation:Süntel
Local section:Oker Local bed:83
Stratigraphic resolution:bed
Stratigraphy comments: absolute age given as about 154 my = middle Kimmeridgian
given as bed 93 in error in Sander et al. 2006
Lithology and environment
Primary lithology:intraclastic lithified carbonaceous mudstone
Includes fossils?Y
Lithology description: "Bed 93 is a carbonate mudstone which near the bone accumulation contains large intraclasts (max. 200 cm3)."
Environment:estuary/bay
Geology comments: lagoon or bay, mixed salinity; "fully marine conditions and was surrounded by several large islands"
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:excellent
Associated major elements:many
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Spatial resolution:allochthonous
Collection methods and comments
Collection methods:salvage,selective quarrying,mechanical,field collection,survey of museum collection
Reason for describing collection:general faunal/floral analysis
Collectors:H. Ludtke Collection dates:1998–
Collection method comments: discovered 1998 by Holger Ludtke
Metadata
Also known as:bed 93
Database number:51859
Authorizer:M. Carrano, O. Rauhut Enterer:M. Carrano, O. Rauhut
Modifier:M. Carrano Research group:vertebrate
Created:2005-06-28 13:19:34 Last modified:2022-09-27 14:22:01
Access level:the public Released:2005-06-28 13:19:34
Creative Commons license:CC BY
Reference information

Primary reference:
33897.ETE R. Windolf. 1998. Dinosaurierfunde in Niedersachsen [Dinosaur finds in Lower Saxony]. Arbeitskreis Paläontologie Hannover 26:1-7 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

81292 C. Bolecsek and O. Wings. 2013. Europasaurus holgeri – the dwarfed dinosaur from the Late Jurassic of Germany. In F. Torcida Fernández-Baldor & P. Huerta (ed.), VI Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno. Libro de Resúmenes 46(47) [M. Carrano/M. Carrano/M. Carrano]
73618 S. W. Evers and O. Wings. 2020. Late Jurassic theropod dinosaur bones from the Langenberg Quarry (Lower Saxony, Germany) provide evidence for several theropod lineages in the central European archipelago. PeerJ 8:e8437 [O. Rauhut/O. Rauhut]
13807ETE M. Fastnacht. 2005. The first dsungaripterid pterosaur from the Kimmeridgian of Germany and the biomechanics of pterosaur long bones. Acta Palaeontologica Polonica 50(2):273-288 [M. Carrano/M. Carrano/M. Carrano]
73619 O. Gerke and O. Wings. 2016. Multivariate and cladistic analyses of isolated teeth reveal sympatry of theropod dinosaurs in the Late Jurassic of northern Germany. PLoS One 11(7):e0158334:1-52 [O. Rauhut/O. Rauhut/M. Carrano]
43567 H.-V. Karl, E. Groning, C. Brauckmann, D. Schwarz, and N. Knotschke. 2006. The Late Jurassic crocodiles of the Langenberg near Oker, Lower Saxony (Germany), and description of related materials (with remarks on the history of quarrying the “Langenberg Limestone” and “Obernkirchen Sandstone”). Clausthaler Geowissenschaften 5:59-77 [P. Mannion/J. Tennant]
48032 H.-V. Karl, E. Groning, C. Brauckmann and N. Knotschke. 2008. First remains of the head of Steneosaurus (Crocodylomorpha: Teleosauridae) from the Late Jurassic of Oker (Lower Saxony, Germany). Studia Geologica Salmanticensia 44(2):187-201 [P. Mannion/J. Tennant]
70732 J. N. Lallensack, P. M. Sander, N. Knötschke and O. Wings. 2015. Dinosaur tracks from the Langenberg Quarry (Late Jurassic, Germany) reconstructed with historical photogrammetry: evidence for large theropods soon after insular dwarfism. Palaeontologia Electronica 18(2):31A:1-34 [M. Carrano/M. Carrano]
69718 T. Martin, A. O. Averianov, J. A. Schultz, A. H. Schwemann, and O. Wings. 2019. Late Jurassic multituberculate mammals from Langenberg Quarry (Lower Saxony, Germany) and palaeobiogeography of European Jurassic multituberculates. Historical Biology [P. Mannion/P. Mannion]
16804ETE O. Mateus, T. Laven, and N. Knotschke. 2004. A dwarf between giants? A new Late Jurassic sauropod from Germany. Journal of Vertebrate Paleontology 24(3, suppl.):90A [M. Carrano/M. Carrano]
76658 T. S. Myers and A. R. Fiorillo. 2009. Evidence for gregarious behavior and age segregation in sauropod dinosaurs. Palaeogeography, Palaeoclimatology, Palaeoecology 274:96-104 [M. Carrano/M. Carrano]
16789ETE P. M. Sander, T. Laven, O. Mateus and N. Knoetschke. 2004. Insular dwarfism in a brachiosaurid sauropod from the Upper Jurassic of Germany. Journal of Vertebrate Paleontology 24(3):108A [M. Carrano/M. Carrano]
17682ETE P. M. Sander, O. Mateus, T. Laven and N. Knótschke. 2006. Bone histology indicates insular dwarfism in a new Late Jurassic sauropod dinosaur. Nature 441:739-741 [M. Carrano/M. Carrano]
61612 D. Schwarz, M. Raddatz, and O. Wings. 2017. Knoetschkesuchus langenbergensis gen. nov. sp. nov., a new atoposaurid crocodyliform from the Upper Jurassic Langenberg Quarry (Lower Saxony, northwestern Germany), and its relationships to Theriosuchus. PLoS ONE 12(2):e0160617:1-47 [P. Mannion/P. Mannion]
59838 J. P. Tennant, P. D. Mannion, and P. Upchurch. 2016. Evolutionary relationships and systematics of Atoposauridae (Crocodylomorpha: Neosuchia): implications for the rise of Eusuchia. Zoological Journal of the Linnean Society 177:854-936 [P. Mannion/P. Mannion]
37683 D. Thies. 1995. Placoid scales (Chondrichthyes: Elasmobranchii) from the Late Jurassic (Kimmeridgian) of northern Germany. Journal of Vertebrate Paleontology 15(3):463-481 [M. Clapham/M. Clapham]
70252 T. van der Lubbe, U. Richter, and N. Knötschke. 2009. Velociraptorine dromaeosaurid teeth from the Kimmeridgian (Late Jurassic) of Germany. Acta Palaeontologica Polonica 54(3):401-408 [M. Carrano/M. Carrano]
82714 R. Windolf. 2000. Dinosaur sites from the Upper Jurassic of northern Germany. In S. Sachs & R. Windolf (ed.), First Symposium on European Dinosaurs. Programme and Abstracts 26-27 [M. Carrano/M. Carrano]
76452 R. Windolf. 2000. Dinosaur sites in the Upper Jurassic of northern Germany. In M.-C. Buchy (ed.), 5th European Workshop on Vertebrate Palaeontology. Program. Abstracts. Excursion Guides 87-88 [M. Carrano/M. Carrano/M. Carrano]