Woodring loc. 138a [lower Gatun Fm]: Tortonian, Panama

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Murex (Murex) recurvirostris (Broderip 1832)
Woodring 1959 R category
Subspecies: Murex (Murex) recurvirostris recurvirostris
    = Haustellum messorium Sowerby 1841
Hendy et al. 2008
Scaphopoda - Gadilida - Gadilidae
Cadulus (Platyschides) epetrion
Woodring 1973 A category
Gastropoda - Heterostropha - Architectonicidae
Heliacus (Astronacus) stonemanae (Maury 1917)
Woodring 1959 R category
Architectonica (Architectonica) nobilis (Röding 1798)
Woodring 1959 R category
original and current combination Architectonica nobilis
Subspecies: Architectonica (Architectonica) nobilis nobilis
Gastropoda - Pseudomelaniidae
Carinodrillia (Carinodrillia) zooki (Brown and Pilsbry 1911)
Woodring 1970 R category
recombined as Carinodrillia zooki
Gastropoda - Epitoniidae
Epitonium (Epitonium) cf. foliaceicostum
Woodring 1959 R category
Epitonium (Asperiscala) sp. (de Boury 1909)
Woodring 1959 R category
Gastropoda - Cerithiidae
Alabina asperoides
Woodring 1959 C category
Subspecies: Alabina asperoides asperoides
Bittium nugatorium Brown and Pilsbry 1911
Woodring 1959 C category
Gastropoda - Turritellidae
Turritella matarucana Hodson 1926
Turritella gatunensis Conrad 1857
Subspecies: Turritella gatunensis gatunensis
Turritella (Torcula) alitilira
original and current combination Torcula
Subspecies: Turritella (Torcula) alitilira alitilira
Gastropoda - Eulimidae
Niso (Niso) mesata n. sp. Woodring 1970
Woodring 1970 R category
Balcis (Balcis) lipara n. sp. Woodring 1970
Woodring 1970 F category
Eulima nobilis Guppy 1896
Woodring 1970 F category
Gastropoda - Neotaenioglossa - Calyptraeidae
Calyptraea centralis (Conrad 1841)
Crepidula plana Say 1822
Crucibulum (Dispotaea) springvaleense (Rutsch 1942)
Gastropoda - Personidae
Distorsio (Rhysema) decussata (Valenciennes 1832)
Woodring 1959 R category
recombined as Distorsio decussata
Subspecies: Distorsio (Rhysema) decussata gatunensis
Gastropoda - Cypraeidae
Cypraea (Muracypraea) henekeni (Sowerby 1850)
Woodring 1959 F category
    = Muracypraea woodringi Landau and Groves 2011
Landau and Groves 2011
Gastropoda - Strombidae
Strombus gatunensis Toula 1908
Woodring 1959 R category
Gastropoda - Neogastropoda - Volutidae
Enaeta ecnomia n. sp. Woodring 1964
Woodring 1964 R category
Gastropoda - Neogastropoda - Muricidae
Murex (? Murex) polynematicus (Brown and Pilsbry 1911)
Woodring 1959 R category
recombined as Vokesimurex polynematicus
Eupleura thompsoni n. sp. Woodring 1959
Woodring 1964 R category
Gastropoda - Neogastropoda - Cystiscidae
Persicula (Rabicea) couviana (Maury 1925)
Woodring 1970 R category
synonym of Persicula propeobesa
Subspecies: Persicula (Rabicea) couviana stenygra
Gastropoda - Neogastropoda - Mitridae
Mitra (Tiara) longa (Gabb 1873)
Woodring 1964 R category
recombined as Subcancilla longa
Subspecies: Mitra (Tiara) longa longa
Gastropoda - Neogastropoda - Olividae
Oliva (Oliva) gatunensis (Toula 1908)
Woodring 1964 C category
original and current combination Oliva gatunensis
Olivella (Niteoliva) terryi Olsson 1956
Woodring 1964 A category
Gastropoda - Neogastropoda - Olivellidae
Agaronia testacea (Lamarck 1811)
Woodring 1964 R category
Subspecies: Agaronia testacea hadra
Gastropoda - Neogastropoda - Nassariidae
Nassarius (? Uzita) praeambiguus (Brown and Pilsbry 1913)
Woodring 1964 C category
Nassarius (Nassarius) parapristus (Gardner 1944)
Woodring 1964 A category
Subspeciees: Nassarius (Nassarius) parapristus conarus
Calophos ectyphus n. sp. Woodring 1964
Woodring 1964 C category
Gastropoda - Neogastropoda - Buccinidae
Cymatophos veatchi (Olsson 1922)
Woodring 1964 C category
Subspecies: Cymatophos veatchi veatchi
Antillophos (Antillophos) candei (d'Orbigny 1842)
Woodring 1964 R category
recombined as Antillophos candeanus
Subspecies: Antillophos (Antillophos) candei gatunensis
Strombinophos mimicus n. sp. Woodring 1964
Woodring 1964 R category
Hanetia dalli (Brown and Pilsbry 1911)
Woodring 1964 F category
Subspecies: Hanetia dalli protera
Gastropoda - Neogastropoda - Columbellidae
Anachis (Costoanachis) mira (Dall 1896)
Woodring 1964 F category
Subspecies: Anachis (Costoanachis) mira fugax
Mitrella (Columbellopsis) aff. fenestrata (Adams 1850)
Woodring 1964 R category
synonym of Columbellopsis nycteis
Strombina (Strombina) lessepsiana (Brown and Pilsbry 1911)
Woodring 1964 C category
Gastropoda - Neogastropoda - Drilliidae
Crassispira (Crassispira) consors (Sowerby 1850)
Woodring 1970 R category
recombined as Hindsiclava consors
Subspecies: Crassispira (Crassispira) consors consors
Gastropoda - Neogastropoda - Turridae
Pleurofusia (Cruziturricula) fusinus (Brown and Pilsbry 1911)
Woodring 1970 F category
recombined as Cruziturricula fusinus
Subspecies: Pleurofusia (Cruziturricula) fusinus fusinus
Pleurolira (Polystira) tenagos Gardner 1947
Woodring 1970 R category
    = Polystira sp. Woodring 1928
Todd and Johnson 2013
Scobinella morierei (Cossmann 1913)
Woodring 1970 R category
Gastropoda - Neogastropoda - Conidae
Conus aemulator Brown and Pilsbry 1911
Woodring 1970 R category
Subspecies: Conus aemulator aemulator
Conus burckhardti Bose 1906
Woodring 1970 R category
Subspecies: Conus burckhardti harrisi
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria (Euclia) codazzii (Anderson 1929)
Woodring 1970 R category
recombined as Euclia codazzii
Cancellaria (Euclia) dinota n. sp. Woodring 1970
Woodring 1970 R category
recombined as Euclia dinota
Cancellaria (Cancellaria) epistomifera (Guppy 1876)
Woodring 1970 R category
Subspecies: Cancellaria (Cancellaria) epsitomera dariena
    = Bivetiella dariena Toula 1908
Landau et al. 2012
Cancellaria (Narona) barystoma n. sp. Woodring 1970
Woodring 1970 R category
recombined as Narona barystoma
Aphera islacolonis (Maury 1917)
Woodring 1970 R category
    = Aphera aphrodite Landau et al. 2012
Landau et al. 2012
Cancellaria (Pyruclia) cibarcola (Anderson 1929)
Woodring 1970 F category
Subspecies: Cancellaria (Pyruclia) cibarcola cibarcola
Gastropoda - Naticidae
Stigmaulax guppiana (Toula 1909)
Natica (Naticarius) stenopa n. sp.
Polinices stanislasmeunieri Maury 1917
Gastropoda - Vitrinellidae
Anticlimax (Anticlimax) gatunensis
Cyclostremiscus (Ponocyclus) pentagonus (Gabb 1873)
Gastropoda - Tornidae
Teinostoma (Pseudorotella) stemonium n. sp. Woodring 1957
Teinostoma (Idioraphe) angulatum
spelled with current rank as Teinostoma (Idioraphie)
Subpecies: Teinostoma (Idioraphe) angulatum trochalum
Gastropoda - Caecidae
Caecum cf. regulare Carpenter 1858
Woodring 1959 R category
recombined as Caecum (Caecum) regulare
Gastropoda - Skeneidae
Solariorbis (Solariorbis) strongylus n. sp.
Solariorbis (Hapalorbis) hyptius n. sp.
Subspecies: Solariorbis (Hapalorbis) hyptius hyptius
Bivalvia - Nuculanida - Nuculanidae
Saccella acrita (Dall 1908)
Woodring 1973 F category
Subspecies: Saccella acrita epacra
Saccella balboae (Brown and Pilsbry 1911)
Woodring 1973 R category
Bivalvia - Pectinida - Pectinidae
Leptopecten ecnomius Woodring 1982
Woodring 1982 C category
Bivalvia - Arcida - Glycymerididae
Glycymeris (Tucetona) secticostata
Woodring 1973 R category
original and current combination Tucetona
Subspecies: Glycymeris (Tucetona) secticostata schencki
Bivalvia - Arcida - Noetiidae
Noetia (Noetia) reversa (Sowerby 1833)
Woodring 1973 R category
recombined as Noetia reversa
Subspecies: Noetia (Noetia) reversa macdonaldi
Bivalvia - Arcida - Arcidae
Anadara (Potiarca) chavezi
Woodring 1973 C category
Anadara (Tosarca) veatchi (Olsson 1922)
Woodring 1973 R category
Subspecies: Anadara (Tosarca) veatchi veatchi
Anadara (Rasia) fissicosta (Spieker 1922)
Woodring 1973 F category
Anadara (Rasia) dariensis (Brown and Pilsbry 1911)
Woodring 1973 C category
Subspecies: Anadara (Rasia) dariensis dariensis
Anadara (Rasia) sechurana (Olsson 1932)
Woodring 1973 C category
Bivalvia - Lucinida - Lucinidae
Bellucina actinus (Dall 1903)
Woodring 1982 A category
Bivalvia - Pholadida - Corbulidae
Bothrocorbula (Hexacorbula) gatunensis (Toula 1909)
Woodring 1982 A category
Caryocorbula oropendula (Olsson 1922)
Woodring 1982 A category
Subspecies: Caryocorbula oropendula stena
Varicorbula disparilis (d'Orbigny 1853)
Woodring 1982 R category
Bivalvia - Cardiida - Mactridae
Mulinia aff. lateralis (Say 1822)
Woodring 1982 R category
Bivalvia - Cardiida - Kelliellidae
Alveinus rotundus Dall 1900
Woodring 1982 A category
Bivalvia - Cardiida - Veneridae
Macrocallista (Paradione) maculata (Linnaeus 1758)
Woodring 1982 C category
recombined as Macrocallista maculata
Clementia (Clementia) dariena (Conrad 1855)
Woodring 1982 R category
Subspecies: Clementia (Clementia) dariena dariena
Cyclinella cyclica (Guppy 1866)
Woodring 1982 R category
Lirophora (Panchione) mactropsis (Conrad 1855)
Woodring 1982 F category
Chione (Chionopsis) tegulum (Brown and Pilsbry 1911)
Woodring 1982 C category
Bivalvia - Cardiida - Chamidae
Arcinella arcinella (Linnaeus 1767)
Woodring 1982 R category
Chama berjadinensis Hodson 1927
Woodring 1982 R category
Bivalvia - Cardiida - Cardiidae
Trachycardium (Dallocardia) baiterum n. sp. Woodring 1982
Woodring 1982 F category
Bivalvia - Cardiida - Tellinidae
Tellina (Eurytellina) punicea (Born 1778)
Woodring 1982 R category
recombined as Eurytellina punicea
Bivalvia - Cardiida - Semelidae
Ervilia valhosierr Gardner 1928
Woodring 1982 A category
Bivalvia - Carditida - Crassatellidae
Eucrassatella (Hybolophus) elassa Woodring 1982
Woodring 1982 A category
Crassinella martinicensis (d'Orbigny 1853)
Woodring 1982 R category
see common names

Geography
Country:Panama
Coordinates: 9.3° North, 79.8° West (view map)
Paleocoordinates:9.1° North, 78.5° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Tortonian 10 m.y. bin:Cenozoic 6
Key time interval:Tortonian
Age range of interval:11.63000 - 7.24600 m.y. ago
Age estimate:11.1 Ma (other)
Stratigraphy
Formation:Gatun Member:lower Gatun
Stratigraphic resolution:bed
Stratigraphy comments: Same stratigraphic position as previous collection (52173). Lower part of the Gatun Formation. The Gatun Formation is sibdivided into three faunal zones (proposed by Thomson and Keen, 1946): lower, middle, and upper. AGE: Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 11.1 Ma (Hendy, unpublished data).
Lithology and environment
Primary lithology:fine poorly lithified silty sandstone
Includes fossils?Y
Lithology description: SPECIFIC: Soft, silty fine-grained sandstone. GENERAL: Massive medium-to very fine-grained sandstone and siltstone. Somewhat calcareous, or marly, somewhat tuffaceous, and with a clay-like matrix. The lower part of the formation consists principally of medium-very fine-grained sandstone.
Environment:marine indet.
Geology comments: No specific environmental data presented.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:selective quarrying,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:USNM
Collection method comments: Collected by Thomson (1942). Material is also held (but not specified for any specific collection) at Stanford University, Acad. Nat. Sci. Phil., Cornell University, PRI, U. Cal. and Cal. Acad. Sci.
Taxonomic list comments:Exhaustive for gastropoda and bivalvia.
Metadata
Also known as:Stanford #2656
Database number:52174
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2005-07-28 14:29:37 Last modified:2013-11-25 13:37:23
Access level:group members Released:2006-07-28 14:29:37
Creative Commons license:CC BY
Reference information

Primary reference:
11306. W. P. Woodring. 1957. Geology and Paleontology of Canal Zone and adjoining parts of Panama. United States Geological Survey Professional Paper 306(A) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

19909 A. J. W. Hendy, D. P. Buick, K. V. Bulinski, C. A. Ferguson, and A. I. Miller. 2008. Unpublished census data from Atlantic coastal plain and circum-Caribbean Neogene assemblages and taxonomic opinions. [A. Hendy/A. Hendy/A. Hendy]
43491 B. Landau, R. E. Petit, and C. M. Silva. 2012. New Cancellariidae (Mollusca, Gastropoda) from the Miocene Gatun Formation of Panama, with eleven new species. Journal of Paleontology 86(6):907-930 [A. Hendy/A. Hendy]
43452 B. M. Landau and L. T. Groves. 2011. Cypraeidae (Mollusca: Gastropoda) from the early Miocene Cantaure Formation of northern Venezuela. Novaplex 12(1-2):1-38 [A. Hendy/A. Hendy]
48647 J. A. Todd and K. G. Johnson. 2013. Dissecting a marine snail species radiation (Conoidea: Turridae: Polystira) over 12 million years in the southwestern Caribbean. Bulletin of Marine Science 89(4) [A. Hendy/A. Hendy]
11372 W. P. Woodring. 1959. Geology and paleontology of Canal Zone and adjoining parts of Panama: Description of Tertiary mollusks (gastropods: Vermetidae to Thaididae). United States Geological Survey Professional Paper 306(B):147-239 [A. Miller/A. Hendy/A. Hendy]
13232 W. P. Woodring. 1964. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Columbellidae to Volutidae). United States Geological Survey Professional Paper 306(C):241-297 [A. Miller/A. Hendy/A. Hendy]
13236 W. P. Woodring. 1970. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). United States Geological Survey Professional Paper 306(D):299-452 [A. Miller/A. Hendy/A. Hendy]
13235 W. P. Woodring. 1973. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae). United States Geological Survey Professional Paper 306(E):453-539 [A. Miller/A. Hendy/A. Hendy]
13359 W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy]