CBI-14, Dzharakuduk II: Middle Turonian - Late/Upper Turonian, Uzbekistan
collected by L. A. Nessov and Uzbek-Russian-British-American-Canadian (URBAC) expedition 1974–1994, 1997–2006
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
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Sheikhia sp.
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Nessov 1995 | 1 specimen | ||||||||
Desertiana mira
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Nessov 1985 | |||||||||
Mammalia
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Theria indet.
Parker and Haswell 1897
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Averianov 2000 | 3 specimens | ||||||||
Mammalia
- Deltatheroida
- Deltatheridiidae
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Sulestes sp.
Nessov 1985
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Averianov 2000 | 1 specimen | ||||||||
= Sulestes karakshi
Nessov 1985
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Averianov et al. 2010 | |||||||||
Sulestes karakshi
Nessov 1985
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Averianov et al. 2010 | 25 specimens | ||||||||
URBAC 00-001, left m2; URBAC 00-057, right dentary fragment; URBAC 03-009, left m2; URBAC 03-043, right dentary fragment; URBAC 03-090, left dentary fragment; URBAC 03-104, left dentary fragment; URBAC 04-149, left maxilla fragment; URBAC 04-164, left maxilla fragment; URBAC 04-169, right m3; URBAC 04-217, left m4; URBAC 04-291, right m2; URBAC 04-344, left m2; URBAC 06-006, right dentary; URBAC 06-019, right m3; URBAC 98–012, right m3; URBAC 99-017, left M2; ZIN 88467, right dentary fragment; URBAC03-222, right dentary; ZIN 89008, left dentary fragment; ZIN 90311, right dentary fragment; CCMGE 5/12455, left dentary fragment; ZIN 90312, left maxilla fragment; CCMGE 40/12000, right maxilla; ZIN 90309, left dentary fragment; ZIN 90310, right dentary fragment; URBAC 04-346, left m4 | ||||||||||
? Deltatheroides kizylkumensis
Nessov 1993
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Averianov 2000 | 1 specimen | ||||||||
synonym of Sulestes karakshi | ||||||||||
Mammalia
- Kennalestidae
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Eoungulatum kudukensis n. gen., n. sp.
Nessov et al. 1998
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Nessov et al. 1998 | 1 specimen | ||||||||
CCMGE 2/12455 (holotype left M2) | ||||||||||
cf. Eoungulatum kudukensis
Nessov et al. 1998
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Nessov et al. 1998 | 3 specimens | ||||||||
CCMGE 16/12953 (left M3), 17/12953 (right M2), 18/12953 (left M3) | ||||||||||
Mammalia
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Sazlestes tis n. gen., n. sp.
Nessov 1997
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Nessov 1997 | |||||||||
including holotype (ZIN C.82553) | ||||||||||
Mammalia
- Zhelestidae
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"Zhelestidae" indet.
Nessov 1990
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Averianov 2000 | 4 specimens | ||||||||
incl. 1 gen. et sp. nov. | ||||||||||
Sorlestes budan n. gen., n. sp.
Nessov 1985
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Nessov 1985 | 8 specimens | ||||||||
synonym of Zhelestes temirkazyk | ||||||||||
CCMGE 3/12176 and extra specimens listed by Nessov et al 1998 | ||||||||||
Parazhelestes sp.
Nessov 1993
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Averianov 2000 | 4 specimens | ||||||||
sp. nov. | ||||||||||
= Parazhelestes minor
Nessov et al. 1998
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Nessov et al. 1998 | |||||||||
synonym of Parazhelestes mynbulakensis | ||||||||||
CCMGE 11/12176, 21/12953, 22/12953, 12/12953 | ||||||||||
Parazhelestes robustus n. gen., n. sp.
Nessov 1993
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Nessov et al. 1998 | 3 specimens | ||||||||
CCMGE 70/12455 (holotype left maxilla), CMGE 10/12953 (left M1), 10/12953 (right M2) | ||||||||||
Aspanlestes cf. aptap
Nessov 1985
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Nessov 1985 | 1 specimen | ||||||||
= ? Aspanlestes sp.
Nessov 1985
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Nessov 1995 | |||||||||
= Aspanlestes aptap
Nessov 1985
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Averianov 2000 | |||||||||
? Zhelestes bezelgen n. sp.
Nessov 1987
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Nessov 1987 | 1 specimen | ||||||||
synonym of Aspanlestes aptap | ||||||||||
CCMGE 1/12455 | ||||||||||
Mammalia
- Zalambdalestidae
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? Kulbeckia sp.
Nessov 1993
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Averianov 2000 | 40 specimens | ||||||||
= Kulbeckia kulbecke
Nessov 1993
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Archibald and Averianov 2003 | |||||||||
~40 specimens listed | ||||||||||
Mammalia
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Paranyctoides aralensis n. sp.
Nessov 1993
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Averianov 2000 | 1 specimen | ||||||||
synonym of Paranyctoides quadrans | ||||||||||
Marsasia sp.
Nessov 1997
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Averianov and Kielan-Jaworowska 1999 | 1 specimen | ||||||||
ZIN C.83131 (left dentary fragment) | ||||||||||
= Paranyctoides aralensis
Nessov 1993
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Averianov et al. 2010 | |||||||||
synonym of Paranyctoides quadrans | ||||||||||
Mammalia
- Periptychidae
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Periptychidae indet.
Cope 1882
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Nessov 1995 | 1 specimen | ||||||||
Mammalia
- Multituberculata
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Uzbekbaatar kizylkumensis n. gen., n. sp.
Kielan-Jaworowska and Nessov 1992
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Kielan-Jaworowska and Nessov 1992 | 11 specimens | ||||||||
including CCMGE 100/ 12455, right p4 (holotype) and specimens referred by Averianov & Archibald (2006); four other cimolodontan multituberculate specimens from this locality may or may not be referrable | ||||||||||
Reptilia
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Buckantaus crassidens n. gen., n. sp.
Nessov 1985
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1 specimen | |||||||||
TsNIGR 26/12000 | ||||||||||
Reptilia
- Testudines
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Khunnuchelys kizylkumensis
Brinkman et al. 1993
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Brinkman et al. 1993 | 2 individuals | ||||||||
CCMGE 303/12458, 8a/12458 | ||||||||||
Reptilia
- Plesiosauria
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? Plesiosauria indet.
de Blainville 1835
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Nessov 1985 | 1 specimen | ||||||||
Reptilia
- Ornithischia
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Ornithischia indet.
Seeley 1888
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Nessov 1995 | 2 specimens | ||||||||
Reptilia
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Turanoceratops tardabilis
Nessov et al. 1989
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Nessov 1995 | 5 specimens | ||||||||
One element - CCGME 628/12457 (partial braincase) - now referred to Titanosauria indet. | ||||||||||
Reptilia
- Ceratopsidae
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? Ceratopsidae indet.
Marsh 1888
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Nessov 1995 | 2 specimens | ||||||||
Reptilia
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Gilmoreosaurus arkhangelskyi
Nessov 1995
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Nessov 1995 | 7 specimens | ||||||||
CCMGE 565/12457, 10/12454, 595/12176 | ||||||||||
= Levnesovia transoxiana
Sues and Averianov 2009
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Sues and Averianov 2009 | |||||||||
Reptilia
- Hadrosauridae
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Hadrosauridae indet.
Cope 1869
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Nessov 1995 | 2 specimens | ||||||||
Cionodon kyzylkumensis
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Nessov 1995 | 1 specimen | ||||||||
nomen dubium belonging to Hadrosauridae | ||||||||||
Reptilia
- Theropoda
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Theropoda indet.
Marsh 1881
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Nessov 1995 | 3 specimens | ||||||||
Reptilia
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Euronychodon asiaticus n. sp.
Nessov 1995
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Nessov 1995 | 22 specimens | ||||||||
synonym of Paronychodon | ||||||||||
CCMGE 9/12454; ZIN PH 301/16, 1052/16, 1053/16, 1055/16, 1056/16, 1057/16, 1058/16, 1060/16, 1062/16, 1063/16, 1064/16, 1065/16, 1066/16, 1069/16, 1070/16, 1072/16, 1149/16, 1150/16, 1151/16, 1152/16, 1833/16 | ||||||||||
Reptilia
- Dromaeosauridae
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Dromaeosauridae indet.
Colbert and Russell 1969
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Nessov 1995 | 1 specimen | ||||||||
Asiamericana asiatica n. gen., n. sp.
Nessov 1995
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Nessov 1995 | 61 specimens | ||||||||
CCMGE 460/12457; ZIN PH 1073/16, 1074/16, 1075/16, 1076/16, 1077/16, 1078/16, 1079/16, 1080/16, 1081/16, 1082/16, 1083/16, 1084/16, 1085/16, 1086/16, 1087/16, 1088/16, 1089/16, 1091/16, 1092/16, 1093/16, 1094/16, 1095/16, 1096/16, 1097/16, 1098/16, 1099/16, 1100/16, 1102/16, 1103/16, 1104/16, 1105/16, 1108/16, 1110/16, 1111/16, 1112/16, 1113/16, 1114/16, 1116/16, 1117/16, 1118/16, 1120/16, 1122/16, 1123/16, 1124/16, 1126/16, 1129/16, 1130/16, 1131/16, 1132/16, 1133/16, 1134/16, 1135/16, 1136/16, 1137/16, 1139/16, 1141/16, 1142/16, 1143/16, 1144/16, 1145/16 | ||||||||||
Itemirus medullaris
Kurzanov 1976
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Sues and Averianov 2014 | 8 specimens | ||||||||
ZIN PH 2338/16, 2352/16, 2353/16, 2351/16, 90/16, 13/16, 2269/16, 182/16 | ||||||||||
Reptilia
- Alexornithiformes
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Alexornithiformes indet.
Brodkorb 1976
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Panteleyev 1998 | 1 specimen | ||||||||
ZIN PO 4821, coracoid | ||||||||||
Reptilia
- Alexornithiformes
- Alexornithidae
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Sazavis prisca n. gen., n. sp.
Nessov 1989
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Nessov and Yarkov 1989 | 1 specimen | ||||||||
PO 3472 | ||||||||||
Ichthyornis maltshevskyi n. sp.
Nessov 1986
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Nessov 1986 | 1 specimen | ||||||||
recombined as Lenesornis maltshevskyi | ||||||||||
PO 3434 | ||||||||||
Explorornis informal sp. 1
Panteleyev 1998
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Panteleyev 1998 | 1 specimen | ||||||||
ZIN PO 4818, coracoid | ||||||||||
Explorornis informal sp. 2
Panteleyev 1998
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Panteleyev 1998 | 1 specimen | ||||||||
ZIN PO 4817, coracoid | ||||||||||
Explorornis nessovi n. gen., n. sp.
Panteleyev 1998
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Panteleyev 1998 | 1 specimen | ||||||||
ZIN PO 4819, coracoid | ||||||||||
Reptilia
- Alexornithiformes
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Catenoleimus anchoretus n. gen., n. sp.
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Panteleyev 1998 | 1 specimen | ||||||||
ZIN PO 4606, coracoid | ||||||||||
Reptilia
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Ichthyornis minusculus n. sp.
Nessov 1990
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Nessov 1990 | 1 specimen | ||||||||
PO 3941 | ||||||||||
? Neornithes indet.
(Gadow 1893)
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Hope 2002 | 1 specimen | ||||||||
PO 3434a | ||||||||||
Reptilia
- Caenagnathidae
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Caenagnathasia martinsoni
Currie et al. 1994
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Sues and Averianov 2015 | 1 specimen | ||||||||
ZIN PH 934/16, dorsal vertebra | ||||||||||
Reptilia
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Tyrannosauroidea indet.
Walker 1964
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Averianov and Sues 2012 | 9 specimens | ||||||||
ZIN PH 1036/16, 1034/16, 1033/16, 1039/16, 684/16, 708/16, 756/16, 734/16, 120/16 | ||||||||||
? Alectrosaurus sp.
Gilmore 1933
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Nessov 1995 | |||||||||
Reptilia
- Ornithomimidae
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Ornithomimidae indet.
Marsh 1890
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Sues and Averianov 2016 | 13 specimens | ||||||||
USNM 538119; ZIN PH 429/16, 129/16, 134/16, 152/16, 611/16, 825/16, 1424/16, 989/16, 576/16, 126/16, 852/16, 821/16 | ||||||||||
Reptilia
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Segnosauria indet.
Barsbold and Perle 1980
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Nessov 1995 | 1 specimen | ||||||||
synonym of Therizinosauroidea | ||||||||||
Sauropoda indet.
Marsh 1878
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Nessov 1985 | 1 specimen | ||||||||
= Titanosauria indet.
Bonaparte and Coria 1993
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Sues et al. 2015 | |||||||||
CCGME 628/12457, incomplete braincase (originally referred to Turanoceratops tardabilis by Nessov [1995:fig. 5]); numerous isolated teeth in ZIN PH collection (including ZIN PH 253/16, ZIN PH 254/16, ZIN PH 255/16, and ZIN PH 256/16); IZANUZ 335, fragment of cervical vertebra; IZANUZ 611 and USNM 538133, dorsal vertebral centra; USNM 538127, anterior caudal vertebra; ZIN PH 962/16, posterior caudal vertebra; ZIN PH 1462/16, fragment of proximal portion of right ulna; ZIN PH 278/16, manual phalanx V-1; IZANUZ 248 and IZANUZ 636, metatarsalia I; IZANUZ 27 and ZIN PH 672/16, metatarsalia II; CCMGE 663/12457 and ZIN PH 280/16, ungual phalanges of pedal digit I; ZIN PH 480/16, ungual phalanx of pedal digit II or III | ||||||||||
Dzharatitanis kingi n. gen., n. sp.
Averianov and Sues 2021
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Averianov and Sues 2021 | 1 specimen | ||||||||
USNM 538127 (holotype), caudal vertebra | ||||||||||
Amphibia
- Temnospondyli
- Discoglossidae
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Estesina elegans
Ročec and Nessov 1993
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Ročec and Nessov 1993 | 1 specimen | ||||||||
LU-N6/335 (maxilla) | ||||||||||
Saevesoederberghia egredia n. gen., n. sp.
Ročec and Nessov 1993
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Ročec and Nessov 1993 | 1 specimen | ||||||||
LU-N6/375 (holotype partial maxilla) | ||||||||||
Aralobatrachus robustus
Nessov 1981
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Ročec and Nessov 1993 | 10 elements | ||||||||
LU-N6/353, 367, 409, 435, 314 (maxilla), 299 (urostyle), 343, 419, 427, 296 (humeri), | ||||||||||
Procerobatrachus paulus n. gen., n. sp.
Ročec and Nessov 1993
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Ročec and Nessov 1993 | 1 specimen | ||||||||
LU-N6/412 (holotype partial maxilla) | ||||||||||
cf. Kizylkuma sp.
Nessov 1981
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Ročec and Nessov 1993 | 3 specimens | ||||||||
LU-N6/319, 322, 423 (humeri) | ||||||||||
Kizylkuma antiqua
Nessov 1981
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Ročec and Nessov 1993 | 3 specimens | ||||||||
LU-N6/271, 332, 398 (maxilla) | ||||||||||
Amphibia
- Temnospondyli
- Gobiatidae
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Gobiatoides parvus n. gen., n. sp.
Ročec and Nessov 1993
|
Ročec and Nessov 1993 | 1 specimen | ||||||||
LU-N6/344 (holotype partial right maxilla) | ||||||||||
Gobiates asiaticus n. sp.
Ročec and Nessov 1993
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Ročec and Nessov 1993 | 1 specimen | ||||||||
LU-N6/370 (holotype partial squamosal) | ||||||||||
Gobiates dzhyrakudukensis n. sp.
Ročec and Nessov 1993
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Ročec and Nessov 1993 | 2 specimens | ||||||||
LU-N6/341 (holotype maxilla), 357 | ||||||||||
Gobiates tatarinovi n. sp.
Ročec and Nessov 1993
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Ročec and Nessov 1993 | 1 specimen | ||||||||
LU-N6/405 | ||||||||||
Gobiates kizylkumensis n. sp.
Ročec and Nessov 1993
|
Ročec and Nessov 1993 | 1 specimen | ||||||||
LU-N6/363 (holotype partial squamosal) | ||||||||||
Amphibia
- Temnospondyli
- Urodela
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Nesovtriton mynbulakensis n. gen., n. sp.
Skutschas 2009
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Skutschas 2009 | 6 specimens | ||||||||
ZIN PH 1/85, 5/49, 3/85, 4/85, 5/85, 8/85 | ||||||||||
Actinopteri
- Amiiformes
- Amiidae
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Amia limosa
Nessov 1985
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nomen dubium belonging to Amiinae | ||||||||||
Actinopteri
- Lepisosteiformes
- Lepisosteidae
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Lepisosteus turanensis
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Nessov 1985 | 1 specimen | ||||||||
Actinopteri
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? Elopidae indet.
Bonaparte 1846
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1 specimen | |||||||||
TsNIGR 12/12000 | ||||||||||
= Elopomorpha indet.
Greenwood et al. 1966
|
Nessov 1995 | |||||||||
Actinopteri
- Albuliformes
- Albulidae
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Albula lapidosa
Nessov 1985
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Actinopteri
- Ichthyodectiformes
- Ichthyodectidae
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? Ichthyodectidae indet.
Crook 1892
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Actinopteri
- Aspidorhynchiformes
- Aspidorhynchidae
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Belonostomus aciculiferus n. sp.
Nessov 1985
|
1 specimen | |||||||||
TsNIGR 12/12000 | ||||||||||
Acipenseriformes
- Polyodontidae
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Psephuroides kazakhorum n. gen., n. sp.
Nessov 1997
|
Nessov 1997 | 25 specimens | ||||||||
including holotype | ||||||||||
see common names |
Geography
Country: | Uzbekistan | State/province: | Navoi |
Coordinates: | 42.1° North, 62.7° East (view map) | ||
Paleocoordinates: | 36.8° North, 57.0° East | ||
Basis of coordinate: | based on nearby landmark | ||
Geographic resolution: | small collection |
Time
Period: | Cretaceous | Epoch: | Late/Upper Cretaceous |
Stage: | Turonian | 10 m.y. bin: | Cretaceous 6 |
Key time interval: | Middle Turonian - Late/Upper Turonian | ||
Age range of interval: | 93.90000 - 89.80000 m.y. ago |
Stratigraphy
Formation: | Bissekty | ||||
Local section: | Dzharakuduk | Local bed: | 57 m | ||
Local order: | bottom to top | ||||
Stratigraphic resolution: | bed | ||||
Stratigraphy comments: Approx. 57 m above the base of the formation |
Lithology and environment
Primary lithology: | "cross stratification" sandstone |
Secondary lithology: | conglomerate |
Lithology description: Formation comprises medium-grained, poorly lithified, cross-bedded fluvial sandstones and clast-supported, well-cemented infraformational conglomerates | |
Environment: | fluvial indet. |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils,mesofossils |
Preservation of anatomical detail: | good |
Fragmentation: | occasional |
Collection methods and comments
Collection methods: | surface (float),field collection | ||
Reason for describing collection: | taxonomic analysis | ||
Museum repositories: | USNM | ||
Collectors: | L. A. Nessov and Uzbek-Russian-British-American-Canadian (URBAC) expedition | Collection dates: | 1974–1994, 1997–2006 |
Metadata
Also known as: | Dzarakuduk; Dzhara-Kuduk, Dzhirakuduk, Dzhyrakuduk, Kul’beke, TsBI-14 | ||
Database number: | 58168 | ||
Authorizer: | M. Carrano, R. Benson, P. Mannion | Enterer: | K. Maguire, P. Mannion, M. Carrano, R. Benson |
Modifier: | M. Carrano | Research group: | vertebrate |
Created: | 2006-01-25 13:56:42 | Last modified: | 2023-07-18 15:07:30 |
Access level: | the public | Released: | 2006-01-25 13:56:42 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
38131. | L. A. Nessov. 1985. Redkie kostnye ryby, nazemiye yashcheritsy i mlekopitayushchie zony limanov i primorskikh nizmennostey mela Kyzylkumov [Rare bony fishes, terrestrial lizards, and mammals in the estuarine and coastal lowland zone of the Kyzyl-Kum Cretaceous]. Yearbook of the All-Union Paleontological Association 28:199-219 [M. Carrano/M. Carrano] |
Secondary references:
39419 | J. D. Archibald and A. O. Averianov. 2003. The Late Cretaceous placental mammal Kulbeckia. Journal of Vertebrate Paleontology 23(2):404-419 [R. Benson/R. Benson] | |
39260 | A. Averianov and Z. Kielan-Jaworowska. 1999. Marsupials from the Late Cretaceous of Uzbekistan. Acta Palaeontologica Polonica 44(1):71-81 [R. Benson/R. Benson] | |
75738 | A. Averianov and H.-D. Sues. 2021. First rebbachisaurid sauropod dinosaur from Asia. PLoS ONE 16(2):e0246620 [B. Gee/B. Gee/P. Mannion] | |
18356 | ETE | A. O. Averianov. 2000. Mammals from the Mesozoic of Kirgizstan, Uzbekistan, Kazakhstan and Tadzhikistan. In M. J. Benton, M. A. Shishkin, D. M. Unwin, E. N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia 627-652 [M. Carrano/M. Carrano/P. Wagner] |
39418 | A. O. Averianov and J. D. Archibald. 2006. New specimens of the multituberculate mammal Uzbekbaatar from the Late Cretaceous of Uzbekistan. Acta Palaeontologica Polonica 51(2):377-380 [R. Benson/R. Benson] | |
39417 | A. O. Averianov, J. D. Archibald, and E. G. Ekdale. 2010. New material of the Late Cretaceous deltatheroidan mammal Sulestes from Uzbekistan and phylogenetic reassessment of the metatherian-eutherian dichotomy. Journal of Systematic Palaeontology 8(3):301-330 [R. Benson/R. Benson] | |
65870 | A. O. Averianov and D. Archibald. 2016. New evidence on the stem placental mammal Paranyctoides from the Upper Cretaceous of Uzbekistan. Palaeontologia Polonica 67:25-33 [P. Mannion/P. Mannion] | |
71016 | A. O. Averianov and H.-D. Sues. 2012. Skeletal remains of Tyrannosauroidea (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan. Cretaceous Research 34:284-297 [M. Carrano/M. Carrano] | |
85320 | A. O. Averianov and H.-D. Sues. 2017. Review of Cretaceous sauropod dinosaurs from Central Asia. Cretaceous Research 69:184-197 [M. Carrano/M. Carrano] | |
19683 | ETE | D. B. Brinkman, L. A. Nessov, and J.-H. Peng. 1993. Khunnuchelys gen. nov., a new trionychid (Testudines: Trionychidae) from the Late Cretaceous of Inner Mongolia and Uzbekistan. Canadian Journal of Earth Sciences 30(10-11):2214-2223 [M. Carrano/M. Carrano/E. Vlachos] |
29037 | S. Hope. 2002. The Mesozoic radiation of Neornithes. In L. M. Chiappe and L. M. Witmer (eds.), Mesozoic Birds: Above the Heads of Dinosaurs. University of California Press, Berkeley 339-388 [J. Alroy/J. Alroy/M. Carrano] | |
38914 | Z. Kielan-Jaworowska and L. A. Nessov. 1992. Multituberculate mammals from the Cretaceous of Uzbekistan. Acta Palaeontologica Polonica 37(1):1-17 [R. Benson/R. Benson] | |
18352 | ETE | E. N. Kurochkin. 2000. Mesozoic birds of Mongolia and the former USSR. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 533-559 [M. Carrano/M. Carrano/M. Carrano] |
38153 | L. A. Nessov. 1985. Novye mlekopitayushchie Mela Kyzylkumov [New mammals from the Cretaceous of Kyzyl-Kum]. Vestnik Leningradskogo Universiteta, Geologiia, Geografiia, series 7 17:8-18 [M. Carrano/M. Carrano] | |
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