SH-20, Khodzhakul: Early/Lower Cenomanian, Uzbekistan
collected by A. D. Arkhangelsky, L. A. Nessov 1914; 1978, 1981–1982, 1985
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
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Teredolithes limanicus
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Nessov 1985 | |||||||||
Equisetopsida
- Araucariaceae
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? Araucariaceae indet.
Henckel and Hochstetter 1865
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Nessov 1985 | 1 specimen | ||||||||
= Araucariaceae indet.
Henckel and Hochstetter 1865
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Nessov 1995 | |||||||||
Angiospermae
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Angiospermae indet.
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Ročec and Nessov 1993 | |||||||||
Coniferales
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Coniferales indet.
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Ročec and Nessov 1993 | |||||||||
Cephalopoda
- Ammonoidea
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Ammonoidea indet.
Zittel 1884
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Ročec and Nessov 1993 | |||||||||
Gastropoda
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Gastropoda indet.
Cuvier 1795
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Ročec and Nessov 1993 | |||||||||
Bivalvia
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Bivalvia indet.
Linnaeus 1758
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Ročec and Nessov 1993 | |||||||||
Malacostraca
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Malacostraca indet.
(Latreille 1802)
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Ročec and Nessov 1993 | |||||||||
Chondrichthyes
- Hybodontiformes
- Distobatidae
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Glickmanodus rarus n. gen., n. sp.
Nessov et al. 1997
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Nessov 1995 | |||||||||
including holotype: CCMGE 677/12457 | ||||||||||
Chondrichthyes
- Hybodontiformes
- Polyacrodontidae
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Polyacrodus torosus
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Nessov 1995 | 1 element | ||||||||
Polyacrodus ex gr. illingworthi
(Dixon 1850)
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Nessov 1995 | 1 element | ||||||||
Chondrichthyes
- Batoidea
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Batoidea indet.
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Ročec and Nessov 1993 | |||||||||
Chondrichthyes
- Lamniformes
- Anacoracidae
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Eoanacorax dalinkevichiusi
Glikman and Shvazhaite 1971
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Ročec and Nessov 1993 | |||||||||
recombined as Squalicorax dalinkevichiusi | ||||||||||
Chondrichthyes
- Lamniformes
- Paraisuridae
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Paraisurus sp.
Glikman 1957
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Ročec and Nessov 1993 | |||||||||
Chondrichthyes
- Holocephali
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Holocephali indet.
Bonaparte 1832
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Ročec and Nessov 1993 | |||||||||
Chondrichthyes
- Chimaeriformes
- Callorhynchidae
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? Elasmodus sp.
Egerton
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Averianov 2001 | |||||||||
Acipenseriformes
- Polyodontidae
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Polyodontidae indet.
Bonaparte 1838
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Nessov 1995 | 1 element | ||||||||
Hesperopsephurus kyatensis n. gen., n. sp.
Nessov 1997
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Nessov 1997 | 26 specimens | ||||||||
including holotype CCMGE 32/12177 | ||||||||||
Amphibia
- Temnospondyli
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Anura indet.
(Fischer von Waldheim 1813)
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Ročec and Nessov 1993 | |||||||||
Amphibia
- Temnospondyli
- Pelobatidae
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? Pelobatidae indet.
Lataste 1879
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Nessov 1982 | |||||||||
Amphibia
- Temnospondyli
- Urodela
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Mynbulakia nongratis
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Nessov 1982 | |||||||||
synonym of Eoscapherpeton | ||||||||||
Horezmia gracile n. gen., n. sp.
Nessov 1981
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Nessov 1981 | |||||||||
ZIN PHA 77-2 | ||||||||||
Mammalia
- Bobolestidae
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Bobolestes zenge n. gen., n. sp.
Nessov 1985
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Nessov 1985 | 1 specimen | ||||||||
from "Khodzhakul, SKH-20"; CCMGE 2/12176 | ||||||||||
Mammalia
- Zhelestidae
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cf. Sheikhdzheilia rezvyii
Averianov and Archibald 2005
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Averianov and Archibald 2005 | |||||||||
from "Khodzhakul, SKH-20" | ||||||||||
Mammalia
- Pappotheriidae
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Pappotheriidae sp.
Slaughter 1965
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Nessov 1988 | |||||||||
Teeth | ||||||||||
Reptilia
- Testudines
- Adocidae
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Adocus kizylkumensis
Nesov 1981
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Nessov 1981 | |||||||||
Reptilia
- Testudines
- Lindholmemydidae
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Mongolemys sp.
Khosatzky and Mlynarski 1971
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Nessov 1981 | |||||||||
= Mongolemys cf. occidentalis
Nessov 1984
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Nessov 1984 | |||||||||
recombined as Khodzhakulemys occidentalis | ||||||||||
CCMGE 15/12086, fragment of nuchal | ||||||||||
Reptilia
- Testudines
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Kizylkumemys schultzi
Nessov 1977
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Nessov 1981 | |||||||||
= Kizylkumemys sp.
Nessov 1977
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Nessov 1984 | |||||||||
Reptilia
- Testudines
- Nanhsiungchelyidae
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Basilemys sp.
Hay 1902
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Nessov 1981 | |||||||||
= Nanhsiungchelyidae indet.
Yeh 1966
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Nessov 1982 | |||||||||
Reptilia
- Testudines
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cf. Kirgizemys sp.
Nessov and Khozatsky 1973
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Nessov 1984 | |||||||||
Oxemys gutta n. gen., n. sp.
Nessov 1977
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Nessov 1984 | |||||||||
TsNIGR Museum, no. 2/11478 | ||||||||||
Chelonia indet.
(Latreille 1800)
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Ročec and Nessov 1993 | |||||||||
synonym of Testudines | ||||||||||
Reptilia
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Iguania indet.
Cuvier 1817
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Gao and Nessov 1998 | 2 specimens | ||||||||
CMGE 40/12000 (frontal), 41/12000 (partial parietal) | ||||||||||
Gekkota indet.
Cuvier 1817
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Nessov 1995 | 1 specimen | ||||||||
Oxia karakalpakensis n. gen., n. sp.
Nessov 1985
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Nessov 1985 | 4 individuals | ||||||||
Dentaries: CMGE 28/12000 (holotype; Nessov 1985); 29/12000, 43/12000, 44/12000 | ||||||||||
Hodzhakulia magna n. gen., n. sp.
Nessov 1985
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Nessov 1985 | 2 individuals | ||||||||
Maxillae: CMGE30/12000 (holotype, right), 31/12000 (left); right dentaries: 32/12000, 32a/12000 | ||||||||||
Varanoidea indet.
Gray 1827
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Nessov 1982 | |||||||||
Scincomorpha indet.
Camp 1923
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Gao and Nessov 1998 | 1 specimen | ||||||||
CMGE 42/12000 (partial right dentary) | ||||||||||
Reptilia
- Crocodylia
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Crocodylia indet.
(Owen 1842)
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Ročec and Nessov 1993 | |||||||||
Reptilia
- Ankylosauridae
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? Shamosaurinae indet.
Tumanova 1983
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Nessov 1995 | 2 specimens | ||||||||
also listed as Shamosauridae | ||||||||||
Reptilia
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Kulceratops kulensis n. gen., n. sp.
Nessov 1995
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Nessov 1995 | 1 specimen | ||||||||
nomen dubium belonging to Ceratopsia | ||||||||||
CCMGE 495/12457 | ||||||||||
? Gilmoreosaurus atavus n. sp.
Nessov 1995
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Nessov 1995 | 1 specimen | ||||||||
nomen dubium belonging to Euornithopoda | ||||||||||
CCMGE 576/12457 | ||||||||||
Reptilia
- Hypsilophodontidae
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? Hypsilophodontidae indet.
Dollo 1882
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Nessov 1995 | |||||||||
Reptilia
- Hadrosauridae
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Hadrosauridae indet.
Cope 1869
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2 specimens | |||||||||
Reptilia
- Coeluridae
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Coeluridae indet.
Marsh 1881
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Nessov 1995 | |||||||||
Reptilia
- Dromaeosauridae
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Richardoestesia asiatica
(Nessov 1995)
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Sues and Averianov 2013 | 1 specimen | ||||||||
original and current combination Asiamericana asiatica | ||||||||||
ZIN PH 1222/16 | ||||||||||
"Laelaps" cf. explanatus
Cope 1876
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Nessov 1995 | |||||||||
recombined as Dromaeosaurus explanatus | ||||||||||
"possibly this is a peculiar species of Alectrosaurus" | ||||||||||
Reptilia
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Paronychodon asiaticus
(Nessov 1995)
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Sues and Averianov 2013 | 1 specimen | ||||||||
synonym of Paronychodon | ||||||||||
ZIN PH 1071/16 | ||||||||||
Aves indet.
Linnaeus 1758
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Ročec and Nessov 1993 | |||||||||
Reptilia
- Gruiformes
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Horezmavis eocretacea n. gen., n. sp.
Nessov and Borkin 1983
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Nessov and Borkin 1983 | 1 specimen | ||||||||
Reptilia
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Neosauropoda indet.
Bonaparte 1986
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Nessov 1995 | 12 specimens | ||||||||
"Diplodocidae or Titanosauridae"; teeth: ZIN PH 2428/16, 2429/16, 2430/16 | ||||||||||
= Titanosauria indet.
Bonaparte and Coria 1993
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Averianov and Sues 2017 | |||||||||
Actinopteri
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Holostei indet.
(Müller 1846)
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Nessov 1982 | |||||||||
Actinopteri
- Dapediidae
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Dapedius sp.
(Leach 1822)
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Nessov 1982 | |||||||||
original and current combination Dapedium | ||||||||||
Actinopteri
- Lepisosteiformes
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Lepidotes sp.
Agassiz 1832
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Nessov 1982 | |||||||||
Actinopteri
- Aulopiformes
- Enchodontidae
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Enchodontidae indet.
Woodward 1901
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Nessov 1985 | 1 specimen | ||||||||
Actinopteri
- Aulopiformes
- Eurypholidae
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Eurypholis sp.
Pictet 1850
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Nessov 1995 | 2 specimens | ||||||||
Actinopteri
- Ichthyodectiformes
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Sultanuvaisia antiqua
Nessov 1981
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Nessov 1982 | |||||||||
Actinopteri
- Aspidorhynchiformes
- Aspidorhynchidae
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Aspidorhynchidae indet.
Bleeker 1859
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Nessov 1982 | 2 specimens | ||||||||
see common names |
Geography
Country: | Uzbekistan | State/province: | Navoi |
Coordinates: | 40.1° North, 65.8° East (view map) | ||
Paleocoordinates: | 35.9° North, 60.5° East | ||
Basis of coordinate: | based on nearby landmark | ||
Altitude: | 405 meters | ||
Geographic resolution: | small collection |
Time
Period: | Cretaceous | Epoch: | Late/Upper Cretaceous |
Stage: | Cenomanian | 10 m.y. bin: | Cretaceous 5 |
Key time interval: | Early/Lower Cenomanian | ||
Age range of interval: | 99.60000 - 93.50000 m.y. ago |
Stratigraphy
Formation: | Khodzhakul | ||||
Stratigraphic resolution: | bed | ||||
Stratigraphy comments: lower/middle part of the formation. Upper Albian in Averianov (2001), but Early Cenomanian in more recent papers. |
Lithology and environment
Primary lithology: | pebbly calcareous sandstone |
Secondary lithology: | pebbly sandy claystone |
Includes fossils? | Y |
Lithology description: "limonite pebbles lying amongst a suite of calcareous sandstones and sandy clays" | |
Environment: | coastal indet. |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils,mesofossils |
Preservation of anatomical detail: | medium |
Disassociated major elements: | some |
Disassociated minor elements: | some |
Fragmentation: | occasional |
Collection methods and comments
Collection methods: | field collection,survey of museum collection | ||
Reason for describing collection: | general faunal/floral analysis | ||
Collectors: | A. D. Arkhangelsky, L. A. Nessov | Collection dates: | 1914; 1978, 1981–1982, 1985 |
Metadata
Also known as: | Khodja-Kull; SKh-20, CX-20 | ||
Database number: | 58208 | ||
Authorizer: | M. Carrano, J. Alroy, M. Clapham, R. Benson, P. Mannion | Enterer: | K. Maguire, M. Carrano, J. Alroy, M. Clapham, R. Benson, P. Mannion |
Modifier: | M. Carrano | Research group: | vertebrate |
Created: | 2006-01-27 11:55:54 | Last modified: | 2023-07-18 14:55:22 |
Access level: | the public | Released: | 2006-01-27 11:55:54 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
25546. | ETE | A. N. Riabinin. 1931. Ostatki dinozavrov iz verkhnego mela nizov'ev r. Amu-Dar'i [On the dinosaurian remains from the Upper Cretaceous of the lower parts of the Amu-Daria River]. Zapiski Rossiyskogo Mineralogicheskogo Obshchestva 60(1):114-118 [M. Carrano/M. Carrano] |
Secondary references:
28807 | A. Averianov and J. D. Archibald. 2005. Mammals from the mid-Cretaceous Khodzhakul Formation, Kyzylkum Desert, Uzbekistan. Cretaceous Research 26(4):593-608 [J. Alroy/J. Alroy] | |
73859 | A. O. Averianov. 2001. Systematics of the Cretaceous-Paleogene chimaeroid fish of the genus Elasmodus (Chondrichthyes, Holocephali). Paleontological Journal 35:289-300 [M. Clapham/M. Clapham] | |
85320 | A. O. Averianov and H.-D. Sues. 2017. Review of Cretaceous sauropod dinosaurs from Central Asia. Cretaceous Research 69:184-197 [M. Carrano/M. Carrano] | |
44060 | I. G. Danilov. 1999. A new linholmemydid genus (Testudines: Lindholmemydidae) from the mid-Cretaceous of Uzbekistan. Russian Journal of Herpetology 6(1):63-71 [R. Benson/R. Benson] | |
41748 | I. G. Danilov and E. V. Syromyatnikova. 2008. New materials on turtles of the family Nanhsiungchelyidae from the Cretaceous of Uzbekistan and Mongolia, with a review of the Nanhsiungchelyid record in Asia. Proceedings of the Zoological Institute RAS 312(1/2):3-25 [R. Benson/R. Benson/P. Mannion] | |
37483 | K. -Q. Gao and L. A. Nessov. 1998. Early Cretaceous squamates from the Kyzylkum Desert, Uzbekistan. Neues Jahrbuch für Geologie und Paläontologie 207(3):289-309 [R. Benson/R. Benson] | |
18352 | ETE | E. N. Kurochkin. 2000. Mesozoic birds of Mongolia and the former USSR. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 533-559 [M. Carrano/M. Carrano/M. Carrano] |
64195 | L. A. Nessov. 1981. Amfibii i reptilii v ekosistemakh Mela sredney Azii [Amphibia and reptiles in Cretaceous ecosystems of central Asia]. In I. V. Darevsky (ed.), The Problems of Herpetology. Fifth Herpetological Conference. Abstracts 91-92 [M. Carrano/M. Carrano/M. Carrano] | |
38127 | L. A. Nessov. 1982. Drevneishie mlekopitaiushchie SSSR [Ancient mammals of the USSR]. Palyentologicheskogo Obshchyestva 25:228-243 [M. Carrano/M. Carrano] | |
41769 | L. A. Nessov. 1984. Data on late Mesozoic turtles from the USSR. Studia Geologica Salamanticensia, vol. especial 1 (Studia Palaeocheloniologica I) 1:215-223 [R. Benson/R. Benson] | |
38131 | L. A. Nessov. 1985. Redkie kostnye ryby, nazemiye yashcheritsy i mlekopitayushchie zony limanov i primorskikh nizmennostey mela Kyzylkumov [Rare bony fishes, terrestrial lizards, and mammals in the estuarine and coastal lowland zone of the Kyzyl-Kum Cretaceous]. Yearbook of the All-Union Paleontological Association 28:199-219 [M. Carrano/M. Carrano] | |
38153 | L. A. Nessov. 1985. Novye mlekopitayushchie Mela Kyzylkumov [New mammals from the Cretaceous of Kyzyl-Kum]. Vestnik Leningradskogo Universiteta, Geologiia, Geografiia, series 7 17:8-18 [M. Carrano/M. Carrano] | |
19868 | L. A. Nessov. 1988. Late Mesozoic amphibians and lizards of Soviet Middle Asia. Acta Zoologica Cracoviensia 31(14):475-486 [E. Fara/E. Fara] | |
16510 | ETE | L. A. Nessov. 1995. Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii [Dinosaurs of northern Eurasia: new data about assemblages, ecology, and paleobiogeography]. Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg 1-156 [M. Carrano/K. Maguire/M. Carrano] |
39261 | L. A. Nessov. 1997. In L. B. Goloneva & A. O. Averianov (ed.), Nemorskiye Pozvonochnyye Melovogo Perioda Severnoy Yevrazii [Cretaceous Non-Marine Vertebrates of Northern Eurasia] 1-218 [R. Benson/R. Benson/M. Carrano] | |
60768 | L. A. Nessov and L. Y. Borkin. 1983. New findings of birds' bones from the Cretaceous of Mongolia and Middle Asia. Trudy Zoologicheskogo Instituta 116:108-110 [M. Uhen/M. Uhen] | |
14517 | ETE | D. B. Norman and H.-D. Sues. 2000. Ornithopods from Kazakhstan, Mongolia and Siberia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 462-479 [M. Carrano/K. Maguire/M. Carrano] |
18862 | Z. Ročec and L. A. Nessov. 1993. Cretaceous anurans from Central Asia. Palaeontographica Abteilung A 226:1-54 [E. Fara/E. Fara/M. Uhen] | |
61991 | M. J. Ryan. 1997. Middle Asian dinosaurs. In P. J. Currie & K. Padian (ed.), Encyclopedia of Dinosaurs 442-444 [M. Carrano/M. Carrano] | |
46400 | H.-D. Sues and A. Averianov. 2013. Enigmatic teeth of small theropod dinosaurs from the Upper Cretaceous (Cenomanian–Turonian) of Uzbekistan. Canadian Journal of Earth Sciences 50(3):306-314 [P. Mannion/J. Tennant/M. Carrano] | |
68327 | K. B. Yuryev. 1954. Kratkiy obzor nakhodok dinozavrov na territorii SSSR [A brief reivew of dinosaur finds in the USSR]. Uchenyye Zapiski, Leningradskogo Ordena Lenina Gosudarstvennogo Universiteta. Seriya Biologicheskikh Nauk 181(38):183-197 [M. Carrano/M. Carrano] |