SH-20, Khodzhakul: Early/Lower Cenomanian, Uzbekistan
collected by A. D. Arkhangelsky, L. A. Nessov 1914; 1978, 1981–1982, 1985

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Teredolithes limanicus
Nessov 1985
Equisetopsida - Araucariaceae
? Araucariaceae indet. Henckel and Hochstetter 1865
Nessov 1985 1 specimen
    = Araucariaceae indet. Henckel and Hochstetter 1865
Nessov 1995
Angiospermae
Angiospermae indet.
Ročec and Nessov 1993
Coniferales
Coniferales indet.
Ročec and Nessov 1993
Cephalopoda - Ammonoidea
Ammonoidea indet. Zittel 1884
Ročec and Nessov 1993
Gastropoda
Gastropoda indet. Cuvier 1795
Ročec and Nessov 1993
Bivalvia
Bivalvia indet. Linnaeus 1758
Ročec and Nessov 1993
Malacostraca
Malacostraca indet. (Latreille 1802)
Ročec and Nessov 1993
Chondrichthyes - Hybodontiformes - Distobatidae
Glickmanodus rarus n. gen., n. sp. Nessov et al. 1997
Nessov 1995
including holotype: CCMGE 677/12457
Chondrichthyes - Hybodontiformes - Polyacrodontidae
Polyacrodus torosus
Nessov 1995 1 element
Polyacrodus ex gr. illingworthi (Dixon 1850)
Nessov 1995 1 element
Chondrichthyes - Batoidea
Batoidea indet.
Ročec and Nessov 1993
Chondrichthyes - Lamniformes - Anacoracidae
Eoanacorax dalinkevichiusi Glikman and Shvazhaite 1971
Ročec and Nessov 1993
recombined as Squalicorax dalinkevichiusi
Chondrichthyes - Lamniformes - Paraisuridae
Paraisurus sp. Glikman 1957
Ročec and Nessov 1993
Chondrichthyes - Holocephali
Holocephali indet. Bonaparte 1832
Ročec and Nessov 1993
Chondrichthyes - Chimaeriformes - Callorhynchidae
? Elasmodus sp. Egerton
Averianov 2001
Acipenseriformes - Polyodontidae
Polyodontidae indet. Bonaparte 1838
Nessov 1995 1 element
Hesperopsephurus kyatensis n. gen., n. sp. Nessov 1997
Nessov 1997 26 specimens
including holotype CCMGE 32/12177
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Ročec and Nessov 1993
Amphibia - Temnospondyli - Pelobatidae
? Pelobatidae indet. Lataste 1879
Nessov 1982
Amphibia - Temnospondyli - Urodela
Mynbulakia nongratis
Nessov 1982
synonym of Eoscapherpeton
Horezmia gracile n. gen., n. sp. Nessov 1981
Nessov 1981
ZIN PHA 77-2
Mammalia - Bobolestidae
Bobolestes zenge n. gen., n. sp. Nessov 1985
Nessov 1985 1 specimen
from "Khodzhakul, SKH-20"; CCMGE 2/12176
Mammalia - Zhelestidae
cf. Sheikhdzheilia rezvyii Averianov and Archibald 2005
Averianov and Archibald 2005
from "Khodzhakul, SKH-20"
Mammalia - Pappotheriidae
Pappotheriidae sp. Slaughter 1965
Nessov 1988
Teeth
Reptilia - Testudines - Adocidae
Adocus kizylkumensis Nesov 1981
Nessov 1981
Reptilia - Testudines - Lindholmemydidae
Nessov 1981
    = Mongolemys cf. occidentalis Nessov 1984
Nessov 1984
recombined as Khodzhakulemys occidentalis
CCMGE 15/12086, fragment of nuchal
Reptilia - Testudines
Kizylkumemys schultzi Nessov 1977
Nessov 1981
    = Kizylkumemys sp. Nessov 1977
Nessov 1984
Reptilia - Testudines - Nanhsiungchelyidae
Basilemys sp. Hay 1902
Nessov 1981
    = Nanhsiungchelyidae indet. Yeh 1966
Nessov 1982
Reptilia - Testudines
cf. Kirgizemys sp. Nessov and Khozatsky 1973
Nessov 1984
Oxemys gutta n. gen., n. sp. Nessov 1977
Nessov 1984
TsNIGR Museum, no. 2/11478
Chelonia indet. (Latreille 1800)
Ročec and Nessov 1993
synonym of Testudines
Reptilia
Iguania indet. Cuvier 1817
Gao and Nessov 1998 2 specimens
CMGE 40/12000 (frontal), 41/12000 (partial parietal)
Gekkota indet. Cuvier 1817
Nessov 1995 1 specimen
Oxia karakalpakensis n. gen., n. sp. Nessov 1985
Nessov 1985 4 individuals
Dentaries: CMGE 28/12000 (holotype; Nessov 1985); 29/12000, 43/12000, 44/12000
Hodzhakulia magna n. gen., n. sp. Nessov 1985
Nessov 1985 2 individuals
Maxillae: CMGE30/12000 (holotype, right), 31/12000 (left); right dentaries: 32/12000, 32a/12000
Varanoidea indet. Gray 1827
Nessov 1982
Scincomorpha indet. Camp 1923
Gao and Nessov 1998 1 specimen
CMGE 42/12000 (partial right dentary)
Reptilia - Crocodylia
Crocodylia indet. (Owen 1842)
Ročec and Nessov 1993
Reptilia - Ankylosauridae
? Shamosaurinae indet. Tumanova 1983
Nessov 1995 2 specimens
also listed as Shamosauridae
Reptilia
Kulceratops kulensis n. gen., n. sp. Nessov 1995
Nessov 1995 1 specimen
nomen dubium belonging to Ceratopsia
CCMGE 495/12457
? Gilmoreosaurus atavus n. sp. Nessov 1995
Nessov 1995 1 specimen
nomen dubium belonging to Euornithopoda
CCMGE 576/12457
Reptilia - Hypsilophodontidae
? Hypsilophodontidae indet. Dollo 1882
Nessov 1995
Reptilia - Hadrosauridae
Hadrosauridae indet. Cope 1869
2 specimens
Reptilia - Coeluridae
Coeluridae indet. Marsh 1881
Nessov 1995
Reptilia - Dromaeosauridae
Richardoestesia asiatica (Nessov 1995)
Sues and Averianov 2013 1 specimen
original and current combination Asiamericana asiatica
ZIN PH 1222/16
"Laelaps" cf. explanatus Cope 1876
Nessov 1995
recombined as Dromaeosaurus explanatus
"possibly this is a peculiar species of Alectrosaurus"
Reptilia
Paronychodon asiaticus (Nessov 1995)
Sues and Averianov 2013 1 specimen
synonym of Paronychodon
ZIN PH 1071/16
Aves indet. Linnaeus 1758
Ročec and Nessov 1993
Reptilia - Gruiformes
Horezmavis eocretacea n. gen., n. sp. Nessov and Borkin 1983
Nessov and Borkin 1983 1 specimen
Reptilia
Neosauropoda indet. Bonaparte 1986
Nessov 1995 12 specimens
"Diplodocidae or Titanosauridae"; teeth: ZIN PH 2428/16, 2429/16, 2430/16
    = Titanosauria indet. Bonaparte and Coria 1993
Averianov and Sues 2017
Actinopteri
Holostei indet. (Müller 1846)
Nessov 1982
Actinopteri - Dapediidae
Dapedius sp. (Leach 1822)
Nessov 1982
original and current combination Dapedium
Actinopteri - Lepisosteiformes
Lepidotes sp. Agassiz 1832
Nessov 1982
Actinopteri - Aulopiformes - Enchodontidae
Enchodontidae indet. Woodward 1901
Nessov 1985 1 specimen
Actinopteri - Aulopiformes - Eurypholidae
Eurypholis sp. Pictet 1850
Nessov 1995 2 specimens
Actinopteri - Ichthyodectiformes
Sultanuvaisia antiqua Nessov 1981
Nessov 1982
Actinopteri - Aspidorhynchiformes - Aspidorhynchidae
Aspidorhynchidae indet. Bleeker 1859
Nessov 1982 2 specimens
see common names

Geography
Country:Uzbekistan State/province:Navoi
Coordinates: 40.1° North, 65.8° East (view map)
Paleocoordinates:35.9° North, 60.5° East
Basis of coordinate:based on nearby landmark
Altitude:405 meters
Geographic resolution:small collection
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Cenomanian 10 m.y. bin:Cretaceous 5
Key time interval:Early/Lower Cenomanian
Age range of interval:99.60000 - 93.50000 m.y. ago
Stratigraphy
Formation:Khodzhakul
Stratigraphic resolution:bed
Stratigraphy comments: lower/middle part of the formation. Upper Albian in Averianov (2001), but Early Cenomanian in more recent papers.
Lithology and environment
Primary lithology:pebbly calcareous sandstone
Secondary lithology:pebbly sandy claystone
Includes fossils?Y
Lithology description: "limonite pebbles lying amongst a suite of calcareous sandstones and sandy clays"
Environment:coastal indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:medium
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:occasional
Collection methods and comments
Collection methods:field collection,survey of museum collection
Reason for describing collection:general faunal/floral analysis
Collectors:A. D. Arkhangelsky, L. A. Nessov Collection dates:1914; 1978, 1981–1982, 1985
Metadata
Also known as:Khodja-Kull; SKh-20, CX-20
Database number:58208
Authorizer:M. Carrano, J. Alroy, M. Clapham, R. Benson, P. Mannion Enterer:K. Maguire, M. Carrano, J. Alroy, M. Clapham, R. Benson, P. Mannion
Modifier:M. Carrano Research group:vertebrate
Created:2006-01-27 11:55:54 Last modified:2023-07-18 14:55:22
Access level:the public Released:2006-01-27 11:55:54
Creative Commons license:CC BY
Reference information

Primary reference:

25546.ETE A. N. Riabinin. 1931. Ostatki dinozavrov iz verkhnego mela nizov'ev r. Amu-Dar'i [On the dinosaurian remains from the Upper Cretaceous of the lower parts of the Amu-Daria River]. Zapiski Rossiyskogo Mineralogicheskogo Obshchestva 60(1):114-118 [M. Carrano/M. Carrano]

Secondary references:

28807 A. Averianov and J. D. Archibald. 2005. Mammals from the mid-Cretaceous Khodzhakul Formation, Kyzylkum Desert, Uzbekistan. Cretaceous Research 26(4):593-608 [J. Alroy/J. Alroy]
73859 A. O. Averianov. 2001. Systematics of the Cretaceous-Paleogene chimaeroid fish of the genus Elasmodus (Chondrichthyes, Holocephali). Paleontological Journal 35:289-300 [M. Clapham/M. Clapham]
85320 A. O. Averianov and H.-D. Sues. 2017. Review of Cretaceous sauropod dinosaurs from Central Asia. Cretaceous Research 69:184-197 [M. Carrano/M. Carrano]
44060 I. G. Danilov. 1999. A new linholmemydid genus (Testudines: Lindholmemydidae) from the mid-Cretaceous of Uzbekistan. Russian Journal of Herpetology 6(1):63-71 [R. Benson/R. Benson]
41748 I. G. Danilov and E. V. Syromyatnikova. 2008. New materials on turtles of the family Nanhsiungchelyidae from the Cretaceous of Uzbekistan and Mongolia, with a review of the Nanhsiungchelyid record in Asia. Proceedings of the Zoological Institute RAS 312(1/2):3-25 [R. Benson/R. Benson/P. Mannion]
37483 K. -Q. Gao and L. A. Nessov. 1998. Early Cretaceous squamates from the Kyzylkum Desert, Uzbekistan. Neues Jahrbuch für Geologie und Paläontologie 207(3):289-309 [R. Benson/R. Benson]
18352ETE E. N. Kurochkin. 2000. Mesozoic birds of Mongolia and the former USSR. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 533-559 [M. Carrano/M. Carrano/M. Carrano]
64195 L. A. Nessov. 1981. Amfibii i reptilii v ekosistemakh Mela sredney Azii [Amphibia and reptiles in Cretaceous ecosystems of central Asia]. In I. V. Darevsky (ed.), The Problems of Herpetology. Fifth Herpetological Conference. Abstracts 91-92 [M. Carrano/M. Carrano/M. Carrano]
38127 L. A. Nessov. 1982. Drevneishie mlekopitaiushchie SSSR [Ancient mammals of the USSR]. Palyentologicheskogo Obshchyestva 25:228-243 [M. Carrano/M. Carrano]
41769 L. A. Nessov. 1984. Data on late Mesozoic turtles from the USSR. Studia Geologica Salamanticensia, vol. especial 1 (Studia Palaeocheloniologica I) 1:215-223 [R. Benson/R. Benson]
38131 L. A. Nessov. 1985. Redkie kostnye ryby, nazemiye yashcheritsy i mlekopitayushchie zony limanov i primorskikh nizmennostey mela Kyzylkumov [Rare bony fishes, terrestrial lizards, and mammals in the estuarine and coastal lowland zone of the Kyzyl-Kum Cretaceous]. Yearbook of the All-Union Paleontological Association 28:199-219 [M. Carrano/M. Carrano]
38153 L. A. Nessov. 1985. Novye mlekopitayushchie Mela Kyzylkumov [New mammals from the Cretaceous of Kyzyl-Kum]. Vestnik Leningradskogo Universiteta, Geologiia, Geografiia, series 7 17:8-18 [M. Carrano/M. Carrano]
19868 L. A. Nessov. 1988. Late Mesozoic amphibians and lizards of Soviet Middle Asia. Acta Zoologica Cracoviensia 31(14):475-486 [E. Fara/E. Fara]
16510ETE L. A. Nessov. 1995. Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii [Dinosaurs of northern Eurasia: new data about assemblages, ecology, and paleobiogeography]. Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg 1-156 [M. Carrano/K. Maguire/M. Carrano]
39261 L. A. Nessov. 1997. In L. B. Goloneva & A. O. Averianov (ed.), Nemorskiye Pozvonochnyye Melovogo Perioda Severnoy Yevrazii [Cretaceous Non-Marine Vertebrates of Northern Eurasia] 1-218 [R. Benson/R. Benson/M. Carrano]
60768 L. A. Nessov and L. Y. Borkin. 1983. New findings of birds' bones from the Cretaceous of Mongolia and Middle Asia. Trudy Zoologicheskogo Instituta 116:108-110 [M. Uhen/M. Uhen]
14517ETE D. B. Norman and H.-D. Sues. 2000. Ornithopods from Kazakhstan, Mongolia and Siberia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 462-479 [M. Carrano/K. Maguire/M. Carrano]
18862 Z. Ročec and L. A. Nessov. 1993. Cretaceous anurans from Central Asia. Palaeontographica Abteilung A 226:1-54 [E. Fara/E. Fara/M. Uhen]
61991 M. J. Ryan. 1997. Middle Asian dinosaurs. In P. J. Currie & K. Padian (ed.), Encyclopedia of Dinosaurs 442-444 [M. Carrano/M. Carrano]
46400 H.-D. Sues and A. Averianov. 2013. Enigmatic teeth of small theropod dinosaurs from the Upper Cretaceous (Cenomanian–Turonian) of Uzbekistan. Canadian Journal of Earth Sciences 50(3):306-314 [P. Mannion/J. Tennant/M. Carrano]
68327 K. B. Yuryev. 1954. Kratkiy obzor nakhodok dinozavrov na territorii SSSR [A brief reivew of dinosaur finds in the USSR]. Uchenyye Zapiski, Leningradskogo Ordena Lenina Gosudarstvennogo Universiteta. Seriya Biologicheskikh Nauk 181(38):183-197 [M. Carrano/M. Carrano]