PBDB Collection

Khodzhakul I: Early/Lower Cenomanian, Uzbekistan

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Reptilia - Testudines - Trionychidae

	Trionyx cf. kyrgyzensis Nessov 1995	Vitek and Danilov 2014
	recombined as Petrochelys kyrgyzensis

	Trionyx dissolutus Vitek and Danilov 2014	Vitek and Danilov 2014
Reptilia - Testudines - Adocidae

	Adocidae indet. Cope 1869	Danilov et al. 2011
	orig. Ferganemys sp.

	Adocus kizylkumensis n. sp. Nesov 1981		6 specimens
	Includes holotype: ZIN PHT S75-15, lateral part of undetermined costal; plus ZIN PH 13/87, neural 1; ZIN PH 8/87, neural 4(?); ZIN PH S78-1, costal 6; ZIN PH 12/87, peripheral 4; ZIN PH 9/87, bridge peripheral
Reptilia - Testudines - Nanhsiungchelyidae

	Nanhsiungchelyidae indet. Yeh 1966	Danilov and Syromyatnikova 2008
Reptilia

	Segnosauria indet. Barsbold and Perle 1980	Nessov 1995	1 specimen
	synonym of Therizinosauroidea

	Ornithocheirus sp. Seeley 1869	Averianov 2007	1 specimen
	ZIN PH 50/44, rostrum fragment
	= Pterodactyloidea indet. Plieninger 1901	Rodrigues and Kellner 2013
	Partial tip of the rostrum (specimen ZIN PH no. 50/44)
Reptilia - Azhdarchidae

	Azhdarchidae indet. Nessov 1984	Nessov 1990	4 specimens
	ZIN PH 61/44, jaw fragment. ZIN PH 80/44, fragment left coracoid. ZIN PH 55/44, fragment of wing phalanx. ZIN PH 44/44, partial pedal phalanx
Reptilia - Ornithocheiridae

	Ornithocheiridae indet. Seeley 1870	Averianov 2007	5 specimens
	ZIN PH 46/44, 47/44, jaw fragments. ZIN PH 41, 44, teeth. ZIN PH 4/44, partial scapula
see common names

Geography

Country:	Uzbekistan	State/province:	Karakalpakstan
Coordinates:	40.1° North, 65.8° East (view map)
Paleocoordinates:	35.9° North, 60.5° East
Basis of coordinate:	based on nearby landmark
Altitude:	405 meters
Geographic resolution:	small collection

Time

Period:	Cretaceous	Epoch:	Late/Upper Cretaceous
Stage:	Cenomanian	10 m.y. bin:	Cretaceous 5
Key time interval:	Early/Lower Cenomanian
Age range of interval:	100.50000 - 93.90000 m.y. ago

Stratigraphy

Formation:	Khodzhakul
Stratigraphic resolution:	bed
Stratigraphy comments: lower part of the formation in the lower middle Albian; also assigned to lower Cenomanian

Lithology and environment

Primary lithology:	not reported
Environment:	terrestrial indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils
Abundance in sediment:	rare
Articulated whole bodies:	none
Associated major elements:	none

Collection methods and comments

Collection methods:	field collection
Reason for describing collection:	general faunal/floral analysis
Collection method comments: Discovered by Sh. Zhuraev in 1978

Metadata

Database number:	58466
Authorizer:	M. Carrano, P. Mannion	Enterer:	K. Maguire, P. Mannion, J. Tennant, M. Carrano
Modifier:	M. Carrano	Research group:	vertebrate
Created:	2006-02-08 12:56:39	Last modified:	2019-06-19 16:54:39
Access level:	the public	Released:	2006-02-08 12:56:39
Creative Commons license:	CC BY

Reference information

Primary reference:

64195.

L. A. Nessov. 1981. Amfibii i reptilii v ekosistemakh Mela sredney Azii [Amphibia and reptiles in Cretaceous ecosystems of central Asia]. In I. V. Darevsky (ed.), The Problems of Herpetology. Fifth Herpetological Conference. Abstracts 91-92 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

31142		A. O. Averianov. 2007. Mid-Cretaceous ornithocheirids (Pterosauria, Ornithocheiridae) from Russia and Uzbekistan. Paleontological Journal 41:79-86 [R. Butler/R. Butler/R. Butler]
37404	ETE	I. G. Danilov, V. B. Sukhanov, and E. V. Syromyatnikova. 2011. New Asiatic materials on turtles of the family Adocidae with a review of the adocid record in Asia. Proceedings of the Zoological Institute, Russian Academy of Sciences 315(2):101-132 [M. Carrano/M. Carrano/P. Mannion]
41748		I. G. Danilov and E. V. Syromyatnikova. 2008. New materials on turtles of the family Nanhsiungchelyidae from the Cretaceous of Uzbekistan and Mongolia, with a review of the Nanhsiungchelyid record in Asia. Proceedings of the Zoological Institute RAS 312(1/2):3-25 [R. Benson/R. Benson/P. Mannion]
31047		L. A. Nessov. 1990. Flying reptiles of the Jurassic and Cretaceous of the USSR and the significance of their remains for the reconstruction of paleogeographic conditions. Bulletin of Leningrad University, Series 7, Geology and Geography 4(28):3-10 [R. Butler/R. Butler/R. Butler]
16510	ETE	L. A. Nessov. 1995. Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii [Dinosaurs of northern Eurasia: new data about assemblages, ecology, and paleobiogeography]. Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg 1-156 [M. Carrano/K. Maguire/M. Carrano]
47175		T. Rodrigues and A. W. A. Kellner. 2013. Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys 308:1-112 [P. Mannion/J. Tennant]
41724		E. V. Syromyatnikova and I. G. Danilov. 2009. New material and a revision of turtles of the genus Adocus (Adocidae) from the Late Cretaceous of middle Asia and Kazakhstan. Proceedings of the Zoological Institute RAS 313(1):74-94 [R. Benson/R. Benson]
31032		D. M. Unwin and N. N. Bakhurina. 2000. Pterosaurs from Russia, Middle Asia and Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 420-433 [R. Butler/R. Butler/M. Carrano]
50301		N. S. Vitek and I. G. Danilov. 2014. Soft-shelled turtles (Trionychidae) from the Cenomanian of Uzbekistan. Cretaceous Research 49:1-12 [P. Mannion/J. Tennant]