PPP# 168 - Escudo de Veraguas [Escudo de Veraguas Fm]: Piacenzian, Panama

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
? Turbonillidae indet.
1 specimen
Turbonillidae indet.
4 specimens
Anthozoa - Scleractinia - Caryophylliidae
Asterosmilia profunda (Duncan 1864)
Gastropoda - Trochidae
Microgaza sp. Dall 1881
7 specimens
Calliostoma sp. Swainson 1840
1 specimen
Gastropoda - Epitoniidae
Epitonium (Asperiscala) sp. (de Boury 1909)
1 specimen
Gastropoda - Eulimidae
Eulimidae indet. Philippi 1853
2 specimens
Scalenostoma sp. Deshayes 1863
1 specimen
Gastropoda - Pseudomelaniidae
? Compsodrillia sp. Woodring 1928
1 specimen
Miraclathurella sp. Woodring 1928
1 specimen
? Crassispira (? Crassispira) sp. Swainson 1840
1 specimen
? Lioglyphostoma sp. Woodring 1928
2 specimens
Gastropoda - Neogastropoda - Olividae
Olivella (Minioliva) sp. Olsson 1956
12 specimens
Olivella (Macgintiella) sp. Olsson 1956
1 specimen
Oliva (Oliva) sp. Bruguière 1789
2 specimens
Gastropoda - Neogastropoda - Buccinidae
Metula sp. Adams and Adams 1853
2 specimens
Antillophos (Antillophos) sp. Woodring 1928
63 specimens
Gastropoda - Neogastropoda - Nassariidae
Nassarius sp. Duméril 1806
1 specimen
sensu-lato
Gastropoda - Neogastropoda - Fasciolariidae
Fusinus sp. Rafinesque 1815
1 specimen
Gastropoda - Neogastropoda - Columbellidae
Cotonopsis (Cotonopsis) sp. Olsson 1942
2 specimens
Gastropoda - Neogastropoda - Conidae
Conus sp. Linnaeus 1758
12 specimens
Gastropoda - Neogastropoda - Drilliidae
Agladrillia (Eumetadrillia) sp. (Woodring 1928)
1 specimen
Agladrillia (Agladrillia) sp. Woodring 1928
6 specimens
Gastropoda - Neogastropoda - Clathurellidae
? Glyphostoma sp. Gabb 1872
1 specimen
Gastropoda - Neogastropoda - Mangeliidae
Vitricythara sp. Fargo 1953
1 specimen
Brachycythara sp. Woodring 1928
2 specimens
Ithycythara sp. Woodring 1928
1 specimen
Gastropoda - Neogastropoda - Terebridae
Strioterebrum sp. Sacco 1891
7 specimens
Gastropoda - Neogastropoda - Turridae
Polystira sp. Woodring 1928
1 specimen
Gastropoda - Neogastropoda - Marginellidae
Marginellidae indet. Fleming 1828
6 specimens
Volvarina sp. (Hinds 1844)
14 specimens
? Dentimargo sp. Cossmann 1899
5 specimens
Gastropoda - Neogastropoda - Cystiscidae
? Persicula sp. Schumacher 1817
1 specimen
Gastropoda - Neogastropoda - Muricidae
Typhinae indet. Cossmann 1903
2 specimens
Haustellum sp. Schumacher 1817
4 specimens
Gastropoda - Cassidae
Sconsia sp. Gray 1847
1 specimen
Gastropoda - Naticidae
Naticidae indet. Guilding 1834
7 specimens
Natica (Naticarius) sp. (Dumeril 1806)
3 specimens
original and current combination Naticarius
Gastropoda - Rissoidae
Rissoidae indet. Gray 1847
2 specimens
Alvania sp. Risso 1826
1 specimen
Gastropoda - Atlantidae
Atlanta sp. Lesueur 1817
3 specimens
Gastropoda - Triphoridae
Triphoridae indet. Gray 1847
1 specimen
Gastropoda - Cocculinidae
? Cocculina sp. Dall 1882
1 specimen
Gastropoda - Heterostropha - Acteonidae
Acteon sp. Montfort 1810
4 specimens
Gastropoda - Heterostropha - Pyramidellidae
Pyramidellidae indet. Gray 1840
1 specimen
Eulimella (Ebalina) sp. (Thiele 1929)
14 specimens
Gastropoda - Heterostropha - Architectonicidae
Pseudotorinia sp. Sacco 1892
2 specimens
Architectonica sp. Röding 1798
5 specimens
Gastropoda - Opisthobranchia - Cylichnidae
Acteocina sp. Gray 1847
1 specimen
Gastropoda - Opisthobranchia - Retusidae
Volvulella (Paravolvulella) sp. (Harry 1967)
1 specimen
Volvulella (Volvulella) sp. Newton 1891
3 specimens
Sulcoretusa sp. Burch 1945
3 specimens
Gastropoda - Opisthobranchia - Cavoliniidae
Cavolinia sp. Avilgaard 1791
1 specimen
Styliola sp. Gray 1850
3 specimens
Bivalvia - Arcida - Arcidae
Anadara (Rasia) sp. (Gray 1857)
3 specimens
Bivalvia - Pectinida - Pectinidae
Amusium sp. (Röding 1798)
1 specimen
Bivalvia - Poromyida - Cuspidariidae
Cuspidaria (Cuspidaria) sp. Dall 1886
3 specimens
Bivalvia - Poromyida - Verticordiidae
Haliris (Haliris) sp. Dall 1886
1 specimen
Trigonulina sp. d'Orbigny 1946
2 specimens
Bivalvia - Cardiida - Montacutidae
Mysella (Mysella) sp. Angas 1877
1 specimen
Bivalvia - Cardiida - Kelliellidae
Kelliella sp. Sars 1865
1 specimen
Bivalvia - Cardiida - Tellinidae
Eurytellina sp. Fischer 1887
1 specimen
spelled with current rank as Tellina (Eurytellina)
Bivalvia - Cardiida - Semelidae
Abra sp. Lamarck 1818
6 specimens
Bivalvia - Nuculida - Nuculidae
Leionucula sp. Quenstedt 1930
4 specimens
Nucula (Lamellinucula) sp. (Schenck 1944)
3 specimens
Bivalvia - Nuculanida - Nuculanidae
Saccella sp. Woodring 1925
22 specimens
Scaphopoda - Gadilida - Entalinidae
Bathoxiphus sp. Pilsbry 1897
9 specimens
Scaphopoda - Gadilida - Gadilidae
Gadilopsis sp. (Woodring 1925)
21 specimens
Scaphopoda - Dentaliida - Dentaliidae
Dentalium sp. Linnaeus 1758
8 specimens
sensu-lato
see common names

Geography
Country:Panama State/province:Bocas
Coordinates: 9.1° North, 81.5° West (view map)
Paleocoordinates:9.8° North, 81.1° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Pliocene
Stage:Piacenzian 10 m.y. bin:Cenozoic 6
Key time interval:Piacenzian
Age range of interval:3.60000 - 2.58800 m.y. ago
Age estimate:3.6 Ma (age-depth)3.6 to 3.5 Ma (other)
Stratigraphy
Geological group:Bocas del Toro Formation:Escudo de Veraguas
Stratigraphic resolution:bed
Stratigraphy comments: STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), planktic foraminifera suggest 1.8 Ma near the top and 3.5 Ma at the base. PPP dataset indicates an age range between 3.6 and 3.5 Ma.
Lithology and environment
Primary lithology:burrows,bioturbation,fine poorly lithified argillaceous sandstone
Secondary lithology:burrows,bioturbation poorly lithified argillaceous siltstone
Includes fossils?Y
Includes fossils?Y
Lithology description: GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, prevasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.
Environment:deep-water indet.
Geology comments: ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:bulk,sieve,field collection
Reason for describing collection:paleoecologic analysis
Museum repositories:BMNH,NMB,USNM
Collection method comments: COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 8/7/87. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),
Taxonomic list comments:NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Metadata
Also known as:NMB Loc. 17621; 87-09-01
Database number:61336
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2006-06-05 20:32:37 Last modified:2013-11-25 13:37:23
Access level:the public Released:2006-06-05 20:32:37
Creative Commons license:CC BY
Reference information

Primary reference:

17667. L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]