Pui, Barbat River: Early/Lower Maastrichtian, Romania
collected by D. Grigorescu 1983

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli - Alytidae
Amphibia indet. Linnaeus 1758
    = ? Alytidae indet. Fitzinger 1843
Venczel et al. 2016
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpeton sp. Estes and Hoffstetter 1976
Venczel et al. 2016
Reptilia - Hypsilophodontidae
Hypsilophodontidae indet. Dollo 1882
Reptilia - Rhabdodontidae
cf. Rhabdodon sp. (Matheron 1869)
teeth
    = Rhabdodon robustus Nopcsa 1900
Grigorescu et al. 1999
recombined as Zalmoxes robustus
Reptilia
Choi et al. 2020
Reptilia - Dromaeosauridae
? Coeluridae indet. Marsh 1881
1 specimen
tooth
    = Deinonychosauria indet. Colbert and Russell 1969
Buffetaut et al. 1986
    = Dromaeosauridae indet. Colbert and Russell 1969
Grigorescu et al. 1999
Reptilia - Titanosauridae
Titanosauridae indet. Lydekker 1885
Weishampel et al. 1991 1 specimen
FGGUB 1007
Reptilia
? Titanosauridae indet. Lydekker 1885
1 specimen
tooth
    = Magyarosaurus dacus Nopcsa 1915
Grigorescu et al. 1999
Reptilia - Crocodyliformes
Crocodilia indet. Owen 1842
    = Crocodyliformes indet. Hay 1930
Puértolas-Pascual et al. 2016
Reptilia - Squamata
Squamata indet. Oppel 1811
Reptilia - Xantusiidae
Paracontogenys sp. Schatzinger 1975
Csiki et al. 2010
Reptilia - Globauridae
Contogenys sp. Estes 1969
Csiki et al. 2010
Reptilia
Testudinata indet. (Oppel 1811)
Reptilia - Testudines - Dortokidae
Rabi et al. 2013
new genus and species
Mammalia - Multituberculata
Multituberculata indet. Cope 1884
2 incisor morphotypes
Mammalia - Multituberculata - Kogaionidae
cf. Paracimexomys sp. Archibald 1982
3 specimens
    = Barbatodon transylvanicum n. gen., n. sp. Radulesco and Samson 1987
Radulesco and Samson 1987
corrected as Barbatodon transylvanicus
ISB IS.001, IS.003; referred material: FGGUB M.1635 and UBB P-Mt1-4
Acipenseriformes
Acipenseriformes indet. Berg 1940
Actinopteri - Characiformes - Characidae
Characidae indet.
Gastropoda - Heterostropha - Anadromidae
Lychnus sp. Matheron 1832
Gastropoda - Cyclophoridae
cf. Cyclophorus sp. Montfort 1810
Gastropoda - Pomatiasidae
cf. Bauxia sp. Caziot 1890
see common names

Geography
Country:Romania State/province:Hunedoara
Coordinates: 45.5° North, 23.1° East (view map)
Paleocoordinates:37.9° North, 19.8° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
Key time interval:Early/Lower Maastrichtian
Age range of interval:72.10000 - 66.00000 m.y. ago
Stratigraphy
Formation:Sînpetru
Stratigraphic resolution:bed
Stratigraphy comments: Customarily referred to the Sînpetru Formation, but it could represent a distinct stratigraphic unit, informally dubbed the ‘Bărbat Formation’ (see Csiki-Sava et al. 2016)

Most of the Pui localities are from the middle-upper part of the Bărbat River succession and are likely to be latest early to early late Maastrichtian (see Csiki-Sava et al. 2016)

Ţabără and Slimani, 2019: The Sînpetru Formation was first assigned to the upper Maastrichtian on the basis of the presence of the pollen Pseudopapilopollis praesubhercynicus (see Antonescu et al., 1983). This assignment was later questioned by Van Itterbeeck et al. (2005), who suggested a lower/upper Maastrichtian boundary interval. Palaeomagnetic studies indicate that the Sînpetru Formation was laid down between the beginning of Chron 32n.1 and the end of Chron 31n (68.7–71.5 Ma) (Panaiotu and Panaiotu, 2010). The age of this formation is more difficult to establish due to its low content in palynomorphs and long-ranging taxa such as Deltoidospora minor, Gleicheniidites senonicus and Laevigatosporites ovatus. All taxa identified by us from the Sînpetru Formation were also previously recorded from the same deposits by Van Itterbeeck et al. (2005) and Csiki et al. (2008). Therefore, we agree the early/late Maastrichtian age of this formation, as suggested by Van Itterbeeck et al. (2005) on the basis of a more conclusive palynological content.
Lithology and environment
Primary lithology:fine,gray,green silty sandstone
Includes fossils?Y
Lithology description: "fine, grey-greenish silty sandstone"
Environment:"channel"
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:excellent
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:frequent
Collection methods and comments
Collection methods:bulk,sieve,field collection
Reason for describing collection:taxonomic analysis
Collectors:D. Grigorescu Collection dates:1983
Metadata
Also known as:Pui 1983; Bărbat River
Database number:67811
Authorizer:M. Carrano, P. Holroyd, P. Mannion Enterer:M. Carrano, P. Holroyd, P. Mannion
Modifier:G. Varnham Research group:vertebrate
Created:2006-12-11 15:51:45 Last modified:2022-02-09 08:17:14
Access level:the public Released:2006-12-11 15:51:45
Creative Commons license:CC BY
Reference information

Primary reference:
55945. D. Grigorescu, J.-L. Hartenberger, C. Radulescu, P. Samson, and J. Sudre. 1985. Découverte de Mammifères et Dinosaures dans le Crétacé supérieur de Pui (Roumaine) [Discovery of mammals and dinosaurs in the Upper Cretaceous of Pui (Romania)]. Comptes Rendus de l'Académie des Sciences à Paris, Série II 301(19):1365-1368 [M. Carrano/M. Carrano]

Secondary references:

25139ETE E. Buffetaut, B. Marandat, and B. Sigé. 1986. Découverte de dents de deinonychosaures (Saurischia, Theropoda) dans le Crétacé superieur du Sud de la France [Discovery of deinonychosaur teeth (Saurischia, Theropoda) in the Upper Cretaceous of southern France]. Comptes Rendus de l'Académie des Sciences à Paris, Série II 303:1393-1396 [M. Carrano/M. Carrano]
81824 S. Choi, M. Moreno-Azanza, Z. Csiki-Sava, E. Prondvai, and Y.-N. Lee. 2020. Comparative crystallography suggests maniraptoran theropod affinities for latest Cretaceous European ‘geckoid’ eggshell. Papers in Palaeontology 6(2):265-292 [M. Carrano/M. Carrano]
37257 Z. Csiki, D. Grigorescu, and M. Rücklin. 2005. A new multituberculate specimen from the Maastrichtian of Pui, Romania and reassessment of affinities of Barbatodon. Acta Palaeontologica Romaniae 5:73-86 [M. Carrano/M. Carrano]
77788 Z. Csiki, D. Grigorescu, V. Codrea and F. Therrien. 2010. Taphonomic modes in the Maastrichtian continental deposits of the Haţeg Basin, Romania—palaeoecological and palaeobiological inferences. Palaeogeography, Palaeoclimatology, Palaeoecology 293(3-4):375-390 [M. Carrano/M. Carrano]
76092 Z. Csiki-Sava, M. Vremir, S. Vasile, S. L. Brusatte, G. Dyke, D. Naish, M. A. Norell and R. Totoianu. 2016. The East Side Story – The Transylvanian latest Cretaceous continental vertebrate record and its implications for understanding Cretaceous–Paleogene boundary events. Cretaceous Research 57:662-698 [P. Mannion/P. Mannion]
19377ETE D. Grigorescu, M. Venczel, Z. Csiki and R. Limberea. 1999. New latest Cretaceous microvertebrate fossil assemblages from the Haţeg Basin (Romania). Geologie en Mijnbouw 78:301-314 [M. Carrano/M. Carrano/P. Mannion]
76284 E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
59665 M. Rabi, M. Vremir, and H. Tong. 2013. Preliminary Overview of Late Cretaceous Turtle Diversity in Eastern Central Europe (Austria, Hungary, and Romania). In D. B. Brinkman, P. A. Holroyd, and J. D. Gardner (eds.), Morphology and Evolution of Turtles 307-336 [P. Holroyd/P. Holroyd]
29885ETE C. Radulesco and P. Samson. 1987. Précisions sur les affinités des Multituberculés (Mammalia) du Crétacé Supérieur de Roumanie [Precise details on the affinities of the multituberculates (Mammalia) from the Upper Cretaceous of Romania]. Comptes Rendus de l'Académie des Sciences à Paris, Série II 304(1):1825-1830 [M. Carrano/M. Carrano/M. Carrano]
60309 T. Smith and V. Codrea. 2015. Red iron-pigmented tooth enamel in a multituberculate mammal from the Late Cretaceous Transylvanian "Hateg Island". PLoS ONE 10(7):e0132550:1-16 [M. Uhen/L. Shadan/P. Mannion]
65871 A. Solomon, V. Codrea, M. Venczel, M. Dumbravă, and T. Smith. 2016. New Remains of the Multituberculate Mammal Barbatodon from the Upper Cretaceous of the Hațeg Basin (Romania). Journal of Mammalian Evolution 23(4):319-335 [P. Mannion/P. Mannion]
79666 D. Ţabără and H. Slimani. 2022. Palynological and palynofacies analyses of Upper Cretaceous deposits in the Haţeg Basin, southern Carpathians, Romania. Cretaceous Research 104(104185) [P. Mannion/G. Varnham/G. Varnham]
65862 M. Venczel, J. D. Gardner, V. A. Codrea, Z. Csiki-Sava, S. Vasile and A. A. Solomon. 2016. New insights into Europe’s most diverse Late Cretaceous anuran assemblage from the Maastrichtian of western Romania. Palaeobiodiversity and Palaeoenvironments 96(1):61-95 [P. Mannion/P. Mannion]
19380ETE D. B. Weishampel, D. Grigorescu, and D. B. Norman. 1991. The dinosaurs of Transylvania. National Geographic Research & Exploration 7(2):196-215 [M. Carrano/M. Carrano]
13939ETE D. B. Weishampel, C.-M. Jianu, Z. Csiki and D. B. Norman. 2003. Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the latest Cretaceous of Romania. Journal of Systematic Palaeontology 1(2):65-123 [M. Carrano/M. Carrano/M. Carrano]