PBDB Collection

Haikou [Yuanshan Fm (formerly Qiongzhusi)]: Atdabanian, China

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

unclassified

	Cyathocephalus bispinosus

	Diplopyge forcipatus

	Diplopyge minutus

	Glossocaris oculatus

	Jianshania furcatus

	Mafangia subscalaria

	Mafangocaris multinodus

	Ovalicephalus mirabilis

	Pterotrum triacanthus

	Pigmaclypeatus daziensis

	Syrrhaptis intestinalis

	Paucipoda haikouensis

	Xioheiqingella peculiaris

	Protosagitta spinosa

	Haikouella lanceolata

	Haikouella jianshanensis

	Zhongxiniscus intermedius

	Zhongjianichthys rostratus

	Anthotrum robustus

	Calathites spinalis

	Dinomisculus venustus

	Macrocephalus elongatus

	Phacatrum tubifer

	Phasganula longa

	Pristioites bifarius

	Rhipitrus clavifer

	Megaspirellus houi
Nectocarididae

	Petalilium latus Luo and Hu 1999
Scleroctenophora

	Trigoides aclis Luo et al. 1999
unclassified

	Xidazoon stephanus n. gen., n. sp. Shu et al. 1999	Shu et al. 1999

	Xidazoon stephanus Shu et al. 1999

	Xianguangia sinica Chen and Erdtmann 1991

	Cambrorhytium elongatum
Hexactinellida - Reticulosa - Protospongiidae

	Saetaspongia densa Mehl and Reitner 1993
Demospongiae - Protomonaxonida - Choiidae

	Choiaella radiata Rigby and Hou 1995

	Choia sp. Walcott 1920

	Allantospongia mica Rigby and Hou 1995
Demospongiae - Protomonaxonida - Halichondritidae

	Halichondrites sp. Walcott 1920
Demospongiae - Protomonaxonida - Hamptoniidae

	Hamptonia bowerbanki Walcott 1920
Demospongiae - Protomonaxonida - Hazeliidae

	Hazelia palmata Walcott 1920
Demospongiae - Protomonaxonida - Leptomitidae

	Leptomitus teretiusculus Chen et al. 1989

	Paraleptomitella dictyodroma Chen et al. 1989
Demospongiae - Vauxiidae

	Vauxia progracilenta n. sp.	Wei et al. 2021

	Vauxia paraleioia	Wei et al. 2021
Agnatha

	Myllokunmingia fengjiaoa n. gen., n. sp. Wagner 2023	Shu et al. 1999

	Myllokunmingia fengjiaoa Wagner 2023

	Haikouichthys ercaicunensis n. gen., n. sp. Luo et al. 1999	Shu et al. 1999

	Haikouichthys ercaicunensis Luo et al. 1999
unclassified

	Cathaymyrus haikouensis
Vetulicolida - Vetulicolidae

	Vetulicola cuneatus

	Vetulicola rectangulata
Vetulicolida - Vetulicolata - Didazoonidae

	Didazoon haoae

	Pomatrum ventralis Luo et al. 1999
unclassified

	Cheungkongella ancestralis n. gen., n. sp. Shu et al. 2001	Shu et al. 2001

	Cheungkongella ancestralis Shu et al. 2001
Eldoniidae

	Stellostomites eumorphus Sun and Hou 1987
unclassified

	Phlogites brevis

	Phlogites longus Luo and Hu 1999
Hyolithida - Sulcavitidae

	Linevitus billingsi (Walcott 1886)

	Ambrolinevitus ventricosus Qian 1978
Hyolithida

	Glossolites magnus
Lingulata - Lingulida - Botsfordiidae

	Diandongia pista Rong 1974
Lingulata - Lingulida - Lingulellotretidae

	Lingulellotreta malongensis (Rong 1974)
Lingulata - Lingulida - Obolidae

	Lingulella chengjiangensis Jin et al. 1993
	recombined as Eoglossa chengjiangensis
Lingulata - Discinida - Disciniidae

	Heliomedusa orienta Sun and Hou 1987	Zhang et al. 2003

	Heliomedusa orienta Sun and Hou 1987
unclassified

	Conicula striata Luo et al. 1999

	Cotyledion tylodes Luo and Hu 1999
Amiskwidae

	Amiskwia sinica
unclassified

	Onychodictyon ferrox	Chen et al. 2002
Parvancorinidae

	Primicaris larvaformis Zhang et al. 2003
Bradoriida

	Kunmingella bispinosus

	Kunmingella douvillei Mansuy 1912

	Acanthomeridion serratum Hou et al. 1989

	Combinivalvula chengjiangensis Hou 1987
unclassified

	Jianfengia multisegmentalis Hou 1987
Radiodonta - Amplectobeluidae

	Amplectobelua trispinata
Radiodonta - Anomalocarididae

	Parapeytoia yunnanensis Hou et al. 1995

	Anomalocaris saron Hou et al. 1995
Isoxyida - Isoxyidae

	Isoxys curvirostratus Vannier and Chen 2000

	Isoxys auritus Jiang 1982

	Isoxys paradoxus Hou 1987
Mollisoniida

	Urokodia aequalis Hou et al. 1989
Fuxianhuiida - Fuxianhuiidae

	Fuxianhuia protensa Hou 1987
Waptiida - Waptiidae

	Waptia sp. Walcott 1912	Shu et al. 1999

	Chuandianella ovata (Li 1975)
unclassified

	Branchiocaris yunnanensis Hou 1987

	Ercaicunia multinodosa Luo et al. 1999
Megacheira - Fortiforcipida - Fortiforcipidae

	Fortiforceps foliosa Hou and Bergström 1997
Megacheira - Leanchoiliida - Leanchoiliidae

	Leanchoilia illecebrosa (Hou 1987)

	Leanchoilia asiatica Luo and Hu 1997
	synonym of Leanchoilia illecebrosa

	Apiocephalus elegans Luo and Hu 1999
	synonym of Leanchoilia illecebrosa

	Dianchia mirabilis Luo and Hu 1997
	synonym of Leanchoilia illecebrosa
unclassified

	Megadictyon sinicum

	Megadictyon haikouensis Luo and Hu 1999
Trilobita - Redlichiida - Yunnanocephalidae

	Yunnanocephalus sp. Kobayashi 1936	Shu et al. 1999

	Yunnanocephalus yunnanensis (Mansuy 1912)
Trilobita - Redlichiida - Redlichiidae

	Kuanyangia pustulosa (Lu 1941)

	Eoredlichia sp. Chang 1950	Shu et al. 1999

	Eoredlichia intermedia (Lu 1940)
	recombined as Eoredlichia intermediata
Nektaspidida - Naraoiidae

	Naraoia sp. Walcott 1912	Shu et al. 1999

	Naraoia longicaudata Zhang and Hou 1985

	Naraoia spinosa Zhang and Hou 1985
Nektaspidida

	Sinoburius lunaris Hou et al. 1991

	Xandarella sp. Hou et al. 1991

	Cindarella eucalla Chen et al. 1996
unclassified

	Kuamaia lata Hou 1987

	Saperion glumaceum Hou et al. 1991

	Squamacula clypeata Hou and Bergström 1997

	Retifacies abnormalis Hou et al. 1989

	Tanglangia longicaudata

	Ercaia minuscula
Odaraiida - Pectocariidae

	? Odaraia eurypetala
	synonym of Pectocaris
Luolishaniida

	Luolishania longicruris Hou and Chen 1989

	Xishania longiusula Hu 2002
	recombined as Facivermis longiusula

	Facivermis yunnanicus Hou and Chen 1989
unclassified

	Hallucigenia fortis Hou and Bergström 1995	Chen et al. 2002

	Cardiodictyon catenulum Hou et al. 1991

	Diania cactiformis n. gen., n. sp. Liu et al. 2011	Liu et al. 2011

	Corynetis brevis Luo and Hu 1999
Ottoidae

	Banffia confusa
unclassified

	Sicyophorus rara Luo and Hu 1999

	? Maotianshania cylindrica Sun and Huo 1987
Cricocosmiidae

	Tabelliscolex hexagonus Han et al. 2003

	Cricocosmia jiningensis

	Tylotites petiolaris Luo and Hu 1999
Palaeoscolecida - Cricocosmida - Palaeoscolecidae

	Palaeoscolex sinensis Hou and Sun 1988
	replaced by Mafangscolex sinensis
unclassified

	Paraselkirkia sinica

	Acosmia maotiania Chen and Zhou 1997

	Fuxianospira gyrata

	Sinocylindra yunnanensis Chen and Erdtmann 1991

	Sandaokania latinodosa Luo and Hu 1999

	Lagenula striolata Luo and Hu 1999

	Discoides abnormis Luo et al. 1999
see common names

Geography

Country:	China	State/province:	Yunnan
Coordinates:	24.8° North, 102.6° East (view map)
Paleocoordinates:	35.1° North, 161.1° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	local area

Time

Period:	Cambrian	Epoch:	Series 2
Stage:	Stage 3	10 m.y. bin:	Cambrian 2
Key time interval:	Atdabanian	Trilobite zone:	Eoredlichia
Age range of interval:	521.00000 - 515.60000 m.y. ago

Stratigraphy

Formation:	Yuanshan
Stratigraphic resolution:	group of beds
Stratigraphy comments: LITHOSTRATIGRAPHY: Formerly from the Yuanshan Mbr, Qiongzhusi Fm. Yuanshan is raised to formation rank in recent publications (Steiner 2005, 2007). Wade-Giles romanization of Qiongzhusi is Chiungchussu. AGE: Eoredlichia Zone. Refined to Atdabanian by A. Hendy (2/2009)

Lithology and environment

Primary lithology:	planar lamination,gray,green,yellow lithified carbonaceous mudstone
Includes fossils?	Y
Lithology description: LITHOLOGY: Dark grey-green (yellow, when weathered), finely laminated mudstones, containing organic hash in great quantity. LITHIFICATION: Lithified, on the basis of figures.
Environment:	marine indet.
Geology comments: "benthic sediment flows... are believed to have been the the principal method of burial" (Shu et al. 1999b) "presence of agglutinated quartz grains, substantially coarser than the surrounding matrix... suggests... that the animal inhabited a higher-energy, sandy sea floor and was transported" (Shu et al. 2001)

Taphonomy

Modes of preservation:	body,adpression,soft parts
Lagerst�tten type:	conservation
Size of fossils:	macrofossils
Spatial orientation:	random
Spatial resolution:	allochthonous

Collection methods and comments

Collection methods:	surface (in situ),hydrochloric,field collection
Reason for describing collection:	taxonomic analysis
Collection method comments: COLLECTOR: Multiple collectors. REPOSITORY: Not stated in text.
Taxonomic list comments:COVERAGE: Exhaustive for all groups. NOMENCLATURE: From a combination of authoritative and review publications, with modern nomenclature and species-resolution identifications.

Metadata

Also known as:	Chengjiang biota; Jingmacun, Kunming
Database number:	68083
Authorizer:	J. Alroy, A. Hendy, W. Kiessling, P. Wagner	Enterer:	J. Alroy, P. Wagner, A. Hendy, M. Domansky
Modifier:	P. Wagner	Research group:	marine invertebrate
Created:	2007-01-02 11:35:02	Last modified:	2022-12-20 19:57:56
Access level:	the public	Released:	2007-01-02 11:35:02
Creative Commons license:	CC BY

Reference information

Primary reference:

29233.

M. Steiner, M. Zhu, Y. Zhao and B.-D. Erdtmann. 2005. Lower Cambrian Burgess Shale-type fossil associations of South China. Palaeogeography, Palaeoclimatology, Palaeoecology 220:129-152 [A. Hendy/A. Hendy]

Secondary references:

29234		L. Chen, H. Luo, S. Hu, J. Yin, Z. Jiang, Z. Wu, F. Li and A. Chen. 2002. Early Cambrian Chengjiang fauna in eastern Yunnan, CHina. Yunnan Science and Technology Press, Kunming 1-199 [A. Hendy/A. Hendy]
39908		J. Liu, M. Steiner, J. A. Dunlop, H. Keupp, D. Shu, Q. Ou, J. Han, Z. Zhang, and X. Zhang. 2011. An armoured Cambrian lobopodian from China with arthropod-like appendages. Nature 470(7335):526-530 [W. Kiessling/M. Domansky]
19551		D. Shu, S. Conway Morris, X.-L. Zhang, L. Chen, Y. Li and J. Han. 1999. A pipiscid-like fossil from the Lower Cambrian of south China. Nature 400:746-749 [J. Alroy/J. Alroy]
19552		D. Shu, H.-L. Luo, S. Conway Morris, X.-L. Zhang, S.-X. Hu, L. Chen, J. Han, M. Zhu, Y. Li and L.-Z. Chen. 1999. Lower Cambrian vertebrates from south China. Nature 402:42-46 [J. Alroy/J. Alroy]
19548		D.-G. Shu, L. Chen, J. Han and X.-L. Zhang. 2001. An Early Cambrian tunicate from China. Nature 411:472-473 [J. Alroy/J. Alroy]
83424		F. Wei, Y. Zhao, A. Chen, X. Hou, and P. Cong. 2021. New vauxiid sponges from the Chengjiang Biota and their evolutionary significance. Journal of the Geological Society 178(5):jgs2020-162 [P. Wagner/P. Wagner]
29209		X.-G. Zhang, X.-G. Hou, and C. C. Emig. 2003. Evidence of lophophore diversity in Early Cambrian Brachiopoda. Proceedings of the Royal Society, London B (Suppl.) 270:S65-S68 [A. Hendy/A. Hendy]