PBDB Collection

Mont Berru: Thanetian, France

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Amphibia - Temnospondyli - Batrachosauroididae

	Palaeoproteus gallicus Estes et al. 1967	Estes et al. 1967
	MNHN CR 6705-6708, 6722 (atlantes), 6709-6711 (trunk vertebrae) and 6712 (dentary)
Reptilia - Remiornithidae

	Remiornis heberti Lemoine 1881	Martin 1992
	Referred material (MNHN BR)
Reptilia - Gastornithiformes - Gastornithidae

	Gastornis minor Lemoine 1878	Martin 1992
	Referred material

	Gastornis parisiensis Hébert 1855	Martin 1992
	Referred material (MNHN BR)

	Gastornis russelli n. sp. Martin 1992	Martin 1992
	MNHN R 3560 - type (left tarsometatarsus); MNHN R 2583 (a bill fragment) may also belong to this species (Martin 1992)
Reptilia - Strigiformes

	Berruornis orbisantiqui n. gen., n. sp. Mourer-Chauviré 1994	Mourer-Chauviré 1994
	MNHN R4155 - holotype (incomplete right tarsometatarsus); referred material: MNHN BR 11186 (distal left tibiotarsus), BR 11195 (proximal right tarsometatarsus), BR 12482-12483 (right tarsometatarsus) and BR 14571 (right tarsometatarsus)
Reptilia

	Asiatosuchus sp. Mook 1940	Martin et al. 2014
	= Asiatosuchus depressifrons Blainville 1855	Delfino et al. 2019
	IRSNB R357-363, IRSNB Vert 26857,MNHN F BR 15070 (and more)

	Diplocynodon remensis n. sp. Martin et al. 2014	Martin et al. 2014
	MHNH F BR 4020 - type (a nearly complete skull); referred material (MNHN)

	Laurasichersis relicta n. gen., n. sp. Pérez-García 2020	Pérez-García 2020
	MNHN.F BR13513 - holotype
Reptilia - Testudines - Trionychidae

	Trionyx sp. Geoffroy Saint-Hilaire 1809	Broin 1977
Reptilia - Testudines

	Gallica lapparentiana n. gen., n. sp. Pérez-García 2020	Pérez-García 2020
	MNHN.F BR13516 - holotype
Reptilia - Compsemydidae

	Berruchelus russelli n. gen., n. sp. Pérez-García 2012	Pérez-García 2012
	MNHN.F.BR 9110 - holotype (nuchal plate); referred material (MNHN): neurals (BR 2780, BR 9097, BR 9098, BR 15256), costals (BR 2827, BR 4181, BR 4288, BR 9038, BR 9043, BR 9062, BR 9092, BR 9101–BR 9103, BR 9105, BR 9112, BR 9115, BR 9118, BR 13430, BR 13433–BR 13647, BR 13651, BR 13652, BR 13713, BR 15002, BR 15005, BR 15009, BR 15043, BR 15059, BR 15151, BR 15157–BR 15159, BR 15169, BR 15408, BR 17468–BR 17470), peripherals (BR 872, BR 2676, BR 2690, BR 2775, BR 2776, BR 2793, BR 2834, BR 2861, BR 2728, BR 2734, BR 3655, BR 3658, BR 4182, BR 4217, BR 4218, BR 4289, BR 9093, BR 9099, BR 9100, BR 9104, BR 9109, BR 9113, BR 9114, BR 9116, BR 9117, BR 13447, BR 13455–BR 13457, BR 13709, BR 13728, BR 15010, BR 15126, BR 15255), suprapygal (BR 9096), pygal (BR 15010), epiplastra (BR 9091, BR 13454, BR 13725, BR 17471), entoplastra (BR 4206, BR 4212, BR 13726, BR 15010), hyoplastra (BR 1657, BR 4221, BR 9040, BR 9041, BR 9042, BR 9056, BR 9106, BR 13648, BR 13650, BR 15000, BR 15001, BR 15189, BR 40309), mesoplastra (BR 9036, BR 13649, BR 15003), hypoplastra (BR 2696, BR 2746, BR 4189, BR 4193, BR 4196, BR 9035, BR 9037, BR 9039, BR 9057, BR 9094, BR 9107), xiphiplastra (BR 13723, BR 15004), and seven fragments of indeterminate plates, numbered collectively as BR 15004; material originally preliminarily assigned to Compsemys (Broin 1977)
Mammalia - Macroscelidea - Louisinidae

	Louisina mirabilis n. gen., n. sp. Russell 1964	Hooker and Russell 2012
	MNHN.F.BR143 - holotype (left dentary with crowns of M1–3 and alveoli of P2–4); paratypes: right dentary fragment with M2–3 (MNHN.F.BR-4406); referred material: right maxilla, M1–2, MNHN.F.BR-7-Lass; RM1, MNHN.F.BR-3-L; LM1, MNHN.F.I-729; RM3, MNHN. F.I-728

	Louisina marci Hooker and Russell 2012	Hooker and Russell 2012
	Two RM1, MNHN.F.BR-13-Pn, I-683; five LM2s, MNHN.F.I-583, I-709, I-714, I-718, BR-12-Bn; RM2, MNHN.F.I-707, LM3, IRSNB.CR-48-B-Wouters; RP3, MNHN.F.I-686; LP4, IRSNB.CR-34-B-Wouters; LM1, MNHN.F.I-721; two RM2s, MNHN.F.I-705, IRSNB. CR-42-B-Wouters

	Thryptodon brailloni Hooker and Russell 2012	Hooker and Russell 2012
	Right M1, MNHN.F.I-681; RM2, MNHN.F.I-685; tentatively referred material: RP3, MNHN.F.I-691

	Berrulestes poirieri n. sp. Hooker and Russell 2012	Hooker and Russell 2012
	MNHN.F.I-725 - type (RM1); paratypes: RP4, MNHN.F.I-573; LM2, MNHN.F.BR-17478; two RM2s, MNHN.F.I-688, I-693; RM3, I-261; referred material: RM3, MNHN.F.I-713

	Berrulestes phelizoni n. gen., n. sp. Hooker and Russell 2012	Hooker and Russell 2012
	MNHN.F.BR-29-Ph - holotype (right dentary with P3–M3 and separate condyle); paratypes: RM1, MNHN.F.BR-6-L; LM2, MNHN.F.I-283; RP4, MNHN.F.I-602; RM1, MNHN.F.I-710; LM3, MNHN.F.BR-125-L; three RM3s, MNHN.F.I-580, I-754, I-247; referred material: RM1, MNHN.F.I-606; LM2, MNHN.F.I-692; RM2, MNHN.F.I-689

	Berrulestes pellouini n. sp. Hooker and Russell 2012	Hooker and Russell 2012
	MNHN.F.I-567 - type (LM1); paratypes: LM2, MNHN.F.I-253; two LM3s, MNHN.F.BR-3, BR-6

	Dipavali petri (Russell 1964)	Hooker and Russell 2012
	Associated RM1–3, MNHN.F. BR-15695–7; three RM1, MNHN.F.BR-2-L, BR-4-L, I-272; two LM2s, MNHN.F.I-587, I-690; LM3, MNHN.F.BR-14; two LP3s, MNHN.F.I-668, I-672; LP4, MNHN.F.I-671; twoRP4s, MNHN.F.BR-1-Ph, BR-9-Bn; LM1, MNHN. F.I-667; RM1, MNHN.F.I-673; L dentary, M2–3, MNHN.F.BR-1198; R dentary, M2–3, MNHN.F.BR-101; RM2, MNHN.F.BR-36-L; RM3, MNHN.F.I-669; RDP3, MNHN.F.BR-79

	Walbeckodon girardi Hooker and Russell 2012	Hooker and Russell 2012
	Right dentary with M1–2, MNHN.F.I-207

	Gigarton sigogneauae Hooker and Russell 2012	Hooker and Russell 2012
	LM2, MNHN.F.I-682

	Gigarton meyeri Hooker and Russell 2012	Hooker and Russell 2012
	LM1, MNHN.F.I-251
Mammalia - Condylarthra - Pleuraspidotheriidae

	Orthaspidotherium edwardsi n. gen., n. sp. Lemoine 1885	Ladevèze et al. 2010
Mammalia - Condylarthra - Hyopsodontidae

	Tricuspiodon sobrinus n. sp. Russell 1980	Russell 1980
	BR-2-L - type

	Tricuspiodon magistrae n. sp. Russell 1964	Russell 1980
	BR 4426 - type
Mammalia - Primates - Toliapinidae

	Sarnacius gingerichi (Russell 1981)	Hooker et al. 1999
	MNHN BR-3-Ph
Mammalia - Primates

	Berruvius lasseroni n. gen., n. sp. Russell 1964
	MNHN BR-141 - type; referred material: MNHN BR-8, BR-1109 and BR-22-BN
Mammalia - Primates - Plesiadapidae

	Platychoerops antiquus n. sp. Boyer et al. 2012	Boyer et al. 2012
	NMB Bru4 - type (right lower p3-m3)

	Plesiadapis berruensis n. sp. Jehle et al. 2018	Jehle et al. 2018

	Plesiadapis tricuspidens Gervais 1877	Gingerich 1976
	numerous specimens from the MNHN, YPM, UCMP, and other collections including several skulls

	Plesiadapis remensis Lemoine 1887	Gingerich 1976
	single MNHN specimen from Berru

	Chiromyoides campanicus Stehlin 1916	Gingerich 1973
Mammalia - Lipotyphla

	? Lipotyphla indet. Haeckel 1866	Gheerbrant and Russell 1989
	synonym of Eulipotyphla
	lower molars
Mammalia - Acreodi - Mesonychidae

	Dissacus europaeus Lemoine 1891	Solé et al. 2018
	IRSNB M2096, MNHN.F.2003–2.1121, MNHN.F.BR 13937, 1069, 12524, 1033
Mammalia - Procreodi - Arctocyonidae

	Landenodon phelizoni n. sp. Russell 1980	Russell 1980
	BR 10156 - type

	Landenodon lavocati Russell 1980	Russell 1980
	BR-13-Bn

	Arctocyonides arenae Russell 1964	Smith and Smith 2003
	either Cernay or Berru
Mammalia - Perissodactyla

	cf. Microhyus sp. Teilhard de Chardin 1927	Tabuce et al. 2006
	replaced by Teilhardimys
	MNHN L−3−BR (right M1)

	Teilhardimys brisswalteri Hooker and Russell 2012	Hooker and Russell 2012
	Paratypes: RP4, MNHN.F.I-124; RM1, MNHN.F.I-712; two LM2s, MNHN.F.I-722, I-730; LP3, MNHN.F.I-708; RM1, MNHN.F.I-706; LM2, MNHN.F.I-242; two RM2s, MNHN.F.I-244, I-716; LM3, MNHN.F.I-715; two RM3s, MNHN.F.I-582, I-720; referred material: LM2, MNHN.F.I-726; LM1, BR-39
Mammalia - Adapisoriculidae

	Bustylus cernaysi Gheerbrant and Russell 1991	Gheerbrant and Russell 1991
	MNHN BR 4 Bn, BRL 8 P, BR 9 Ph

	Afrodon sp. Gheerbrant 1988	Gheerbrant and Russell 1989
	e.g. MNHN BR-9-Ph
Mammalia - Pseudorhyncocyonidae

	Phakodon levei (Russell et al. 1967)	Hooker 2013
	MNHN.F.I-601
Mammalia - Multituberculata - Kogaionidae

	Hainina vianeyae n. sp. Peláez-Campomanes et al. 2000	Peláez-Campomanes et al. 2000
	either Cernay or Berru
see common names

Geography

Country:	France	State/province:	Champagne-Ardenne
Coordinates:	49.3° North, 4.2° East (view map)
Paleocoordinates:	43.8° North, 1.7° East
Basis of coordinate:	estimated from map

Time

Period:	Paleogene	Epoch:	Paleocene
Stage:	Thanetian	10 m.y. bin:	Cenozoic 1
Key time interval:	Thanetian
Age range of interval:	59.20000 - 56.00000 m.y. ago

Stratigraphy

Formation:	Sables de Bracheux
Stratigraphic resolution:	bed
Stratigraphy comments: younger than carrière Lemoine, which is late Thanetian (Russell 1981); NP9, MP6 (Smith and Smith 2003) bed 5 (Hooker and Russell 2012)

Lithology and environment

Primary lithology:	conglomerate
Lithology description: conglomerate (Smith and Smith 2003)
Environment:	fluvial indet.
Geology comments: "sédiments fluviatiles"

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils

Collection methods and comments

Reason for describing collection:

taxonomic analysis

Metadata

Also known as:	Berru bed 5; Mouras Quarry; Cernay
Database number:	68428
Authorizer:	M. Uhen, P. Holroyd, L. van den Hoek Ostende, J. Alroy, P. Mannion	Enterer:	M. Uhen, J. Alroy, P. Mannion, P. Holroyd, L. van den Hoek Ostende
Modifier:	P. Mannion	Research group:	vertebrate
Created:	2007-01-22 07:50:17	Last modified:	2020-01-31 04:27:49
Access level:	the public	Released:	2007-01-22 07:50:17
Creative Commons license:	CC BY

Reference information

Primary reference:

29562.

D. E. Russell. 1981. Un primate nouveau du Paléocène supérieur de France. Geobios 14(3):399-405 [J. Alroy/J. Alroy]

Secondary references:

43979		D. M. Boyer, L. Costeur, and Y. Lipman. 2012. Earliest record of Platychoerops (Primates, Plesiadapidae), a new species from Mouras Quarry, Mont de Berru, France. American Journal of Physical Anthropology 149(3):329-346 [P. Mannion/P. Mannion]
60047		F. Broin. 1977. Contribution a l'etude des Cheloniens. Cheloniens continentaux due Cretace et du Tertiare de France. Memoires du Museum National d'Histoire Naturelle, Nouvelle Serie, Serie C, Sciences de la terre 38:1-366 [P. Holroyd/P. Holroyd/P. Holroyd]
63759		M. Delfino, J. E. Martin, F. Lapparent Broin and T. Smith. 2019. Evidence for a pre-PETM dispersal of the earliest European crocodyloids. Historical Biology 31:845-852 [P. Mannion/P. Mannion/P. Mannion]
38121		R. Estes, M. Hecht, and R. Hoffstetter. 1967. Paleocene amphibians from Cernay, France. American Museum Novitates 2295:1-25 [P. Mannion/P. Mannion]
42886		E. Gheerbrant and D. E. Russell. 1989. Presence of the genus Afrodon [Mammalia, Lipotyphla (?), Adapisoriculidae] in Europe; new data for the problem of trans-Tethyan relations between Africa and Europe around the K/T boundary. Palaeogeography, Palaeoclimatology, Palaeoecology 76:1-15 [P. Mannion/P. Mannion]
42888		E. Gheerbrant and D. E. Russell. 1991. Bustylus cernaysi nov. gen., nov. sp., nouvel Adapisoriculidé (Mammalia, Eutheria) Paléocène d'Europe. Geobios 24(4):467-481 [P. Mannion/P. Mannion/P. Mannion]
1632		P. D. Gingerich. 1973. First record of the Palaeocene primate Chiromyoides from North America. Nature 244:517-518 [J. Alroy/J. Alroy/J. Alroy]
1635		P. D. Gingerich. 1976. Cranial Anatomy and Evolution of Early Tertiary Plesiadapidae (Mammalia, Primates). University of Michigan Papers on Paleontology 15:1-141 [J. Alroy/J. Alroy/P. Mannion]
56880		J. J. Hooker. 2013. Origin and evolution of the Pseudorhyncocyonidae, a European Paleogene family of insectivorous placental mammals. Palaeontology 56(4):807-835 [P. Mannion/P. Mannion]
10304		J. J. Hooker, D. E. Russell, and A. Phelzion. 1999. A new family of Plesiadapiformes (Mammalia) from the Old World Lower Paleogene. Palaeontology 42(3):377-407 [J. Alroy/E. Leckey/P. Mannion]
43830		J. J. Hooker and D. E. Russell. 2012. Early Palaeogene Louisinidae (Macroscelidea, Mammalia), their relationships and north European diversity. Zoological Journal of the Linnean Society 164:856-936 [P. Mannion/P. Mannion]
66643		M. Jehle, M. Godinot, D. Delsate, A. Phélizon and J.-L. Pellouin. 2018. Evolution of plesiadapid mammals (Eutheria, Euarchonta, Plesiadapiformes) in Europe across the Paleocene/Eocene boundary: implications for phylogeny, biochronology and scenarios of dispersal. Palaeobiodiversity and Palaeoenvironments 1-59 [P. Holroyd/P. Holroyd]
34057		S. Ladevèze, P. Missiaen, and T. Smith. 2010. First skull of Orthaspidotherium edwardsi (Mammalia, "Condylarthra") from the late Paleocene of Berru (France) and phylogenetic affinities of the enigmatic European family Pleuraspidotheriidae. Journal of Vertebrate Paleontology 30(5):1559-1578 [J. Alroy/J. Alroy]
53670		J. E. Martin, T. Smith, F. Lapparent Broin, F. Escuillié, and M. Delfino. 2014. Late Palaeocene eusuchian remains from Mont de Berru, France, and the origin of the alligatoroid Diplocynodon. Zoological Journal of the Linnean Society 172:867-891 [P. Mannion/P. Mannion]
38160		L. D. Martin. 1992. The status of the Late Paleocene birds Gastornis and Remiornis. Los Angeles: Natural History Museum of Los Angeles County (Sciences series) 36:97-108 [P. Mannion/P. Mannion]
38102		C. Mourer-Chauviré. 1994. A large owl from the Paleocene of France. Palaeontology 37(2):339-348 [P. Mannion/P. Mannion/P. Mannion]
26792		P. Peláez-Campomanes, N. López-Martínez, M.A. Alvarez Sierra and R. Daams. 2000. The earliest mammal of the European Paleocene: the multituberculate Hainina. Journal of Paleontology 74(4):701-711 [L. van den Hoek Ostende/L. van den Hoek Ostende]
44067		A. Pérez-García. 2012. Berruchelus russelli, gen. et sp. nov., a paracryptodiran turtle from the Cenozoic of Europe. Journal of Vertebrate Paleontology 32(3):545-556 [P. Mannion/P. Mannion]
74026		A. Pérez-García. 2020. A European Cenozoic ‘Macrobaenid’: New Data about the Paleocene Arrival of Several Turtle Lineages to Europe. Journal of Vertebrate Paleontology e1795874:1-10 [P. Mannion/P. Mannion]
71657		A. Pérez-García. 2020. Surviving the Cretaceous-Paleogene mass extinction event: A terrestrial stem turtle in the Cenozoic of Laurasia. Scientific Reports 10:1489 [P. Mannion/P. Mannion]
61388		D. E. Russell. 1980. Sur les Condylarthres Cernaysiens Tricuspiodon et Landenodon (Paléocène supérieur de France). Palaeovertebrata 9(Ext):127-166 [P. Mannion/P. Mannion]
19707	5% 57680	T. Smith and R. Smith. 2003. Terrestrial mammals as biostratigraphic indicators in upper Paleocene-lower Eocene marine deposits of the southern North Sea Basin. Geological Society of America Special Paper 369:513-520 [M. Uhen/M. Uhen]
61876		F. Solé, M. Godinot, Y. Laurent, A. Galoyer, and T. Smith. 2018. The European Mesonychid Mammals: Phylogeny, Ecology, Biogeography, and Biochronology. Journal of Mammalian Evolution 25:339-379 [M. Uhen/M. Uhen/P. Mannion]
42854		R. Tabuce, M. T. Antunes, R. Smith and T. Smith. 2006. Dental and tarsal morphology of the European Paleocene/Eocene “condylarth” mammal Microhyus. Acta Palaeontologica Polonica 51(1):37-52 [P. Mannion/P. Mannion/M. Carrano]