Haqil (MNHN collection): Late/Upper Cenomanian, Lebanon

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Cephalopoda - Octobrachia - Trachyteuthididae
Glyphiteuthis libanotica (Fraas 1878)
Jattiot et al. 2015
Chondrichthyes - Trygonidae
Cyclobatis oligodactylus Egerton 1844
Cyclobatis major Davis 1887
Cyclobatis tuberculatus n. sp. Cappetta 1980
Chondrichthyes - Rajiformes - Rajidae
Pararaja expansa (Davis 1887)
Chondrichthyes - Rajiformes - Rhinobatidae
Rhinobatos maronita Pictet and Humbert 1866
Rhinobatos grandis (Davis 1887)
synonym of Rhinobatos maronita
Rhinobatos hakelensis n. sp. Cappetta 1980
Chondrichthyes - Carcharhiniformes - Scyliorhinidae
Scyliorhinus bloti n. sp.
Chondrichthyes - Orectolobiformes - Parascylliidae
Pararhincodon lehami n. sp.
Chondrichthyes - Orectolobiformes
Mesiteia emiliae Gorjanovic-Kramberger 1886
Chondrichthyes - Lamniformes - Cretoxyrhinidae
Plicatolamna semiplicata
Actinopteri - Pycnodontiformes
Akromystax tilmachiton n. gen., n. sp. Poyato-Ariza and Wenz 2005
Poyato-Ariza and Wenz 2005
Actinopteri - Pycnodontiformes - Pycnodontidae
Flagellipinna rhomboides n. gen., n. sp. Cawley and Kriwet 2019
Cawley and Kriwet 2019 4 specimens
MNHN.F.HAK2003, F.HAK2001, F.HAK1972, F.HAK1972b
Actinopteri - Salmoniformes
Clupea lata Agassiz 1839
Pictet and Humbert 1866
recombined as Spaniodon latus
Actinopteri - Aulopiformes - Eurypholidae
Eurypholis boissieri Pictet 1850
Pictet and Humbert 1866
Actinopteri - Aipichthyidae
Aipichthys velifer Woodward 1901
Otero and Gayet 1996
Aipichthys minor (Pictet 1850)
Otero and Gayet 1996
Actinopteri
Clupea gaudryi Pictet and Humbert 1866
Pictet and Humbert 1866
recombined as Scombroclupea gaudryi
Actinopteri - Anguilliformes - Anguillavidae
Anguillavus quadripinnis Hay 1903
Belouze et al. 2003 1 specimen
MNHN HAK 314
Actinopteri - Anguilliformes
Abisaadia hakelensis (Davis 1887)
Belouze et al. 2003
see common names

Geography
Country:Lebanon
Coordinates: 34.2° North, 35.7° East (view map)
Paleocoordinates:13.0° North, 30.4° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Cenomanian 10 m.y. bin:Cretaceous 5
Key time interval:Late/Upper Cenomanian
Age range of interval:99.60000 - 93.50000 m.y. ago
Stratigraphy
Stratigraphic resolution:bed
Stratigraphy comments: STRATIGRAPHIC RELATIONS: Unknown lithostratigraphy. AGE: Cenomanian; no justification provided for age.
Lithology and environment
Primary lithology: lithified calcareous "carbonate"
Includes fossils?Y
Lithology description: SPECIFIC LITHOLOGY: Stated to be calcareous. Appears to be a chalk from figured specimens. LITHIFICATION: Lithified, based on figured specimens.
Environment:carbonate indet.
Geology comments: ENVIRONMENT: No environmental data reported.
Taphonomy
Modes of preservation:body,soft parts
Lagerst�tten type:conservation
Size of fossils:macrofossils
Preservation of anatomical detail:good
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:selective quarrying,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:MNHN
Collection method comments: COLLECTOR: Unknown collector. REPOSITORY: Musée National d'Histoire Naturelle
Taxonomic list comments:COVERAGE: Not exhaustive. NOMENCLATURE: Authoritative publication, modern nomenclature, species resolution data provided.
Metadata
Also known as:Hakel, Haqel
Database number:69118
Authorizer:A. Hendy, M. Clapham, M. Uhen Enterer:A. Hendy, M. Uhen, M. Clapham, J. Mordaunt
Modifier:M. Clapham Research group:marine invertebrate
Created:2007-02-19 10:46:45 Last modified:2020-08-26 10:26:08
Access level:the public Released:2007-02-19 10:46:45
Creative Commons license:CC BY
Reference information

Primary reference:

31066. H. Cappetta. 1980. Les selaciens du Cretace superieur du Liban. II: Batoides. Palaeontolographica Abt. A 168(5-6):149-229 [M. Uhen/M. Uhen/M. Clapham]

Secondary references:

77906 A. Belouze, M. Gayet, and C. Atallah. 2003. Les premiers Anguilliformes : I. Révision des genres cénomaniens Anguillavus Hay, 1903 et Luenchelys nov. gen. Geobios 36:241-273 [M. Clapham/M. Clapham]
77907 A. Belouze, M. Gayet, and C. Atallah. 2003. Les premiers Anguilliformes : II. Paraphylie du genre Urenchelys Woodward, 1900 et relations phylogénétiques. Geobios 36:351-378 [M. Clapham/M. Clapham]
77440 J. J. Cawley and J. Kriwet. 2019. A new genus and species of pycnodontid fish Flagellipinna rhomboides, gen. et sp. nov. (Neopterygii, Pycnodontiformes), from the Upper Cretaceous (Cenomanian) of Lebanon, with notes on juvenile form and ecology. Journal of Vertebrate Paleontology 39(2):1-16 [M. Clapham/J. Mordaunt]
73609 R. Jattiot, A. Brayard, E. Fara and S. Charbonnier. 2015. Gladius-bearing coleoids from the Upper Cretaceous Lebanese Lagerstätten: diversity, morphology, and phylogenetic implications. Journal of Paleontology 89:148-167 [M. Clapham/M. Clapham]
77296 O. Otero and M. Gayet. 1996. Anatomy and phylogeny of the Aipichthyoidea nov. of the Cenomanian Tethys and their place in the Acanthomorpha (Teleostei). Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 202:313-344 [M. Clapham/J. Mordaunt/M. Clapham]
77414 F. J. Pictet and A. Humbert. 1866. Nouvelles Recherches sur les Poissons Fossiles du Mont Liban 1-115 [M. Clapham/M. Clapham]
43282 J. Poyato-Ariza and S. E. K. Wenz. 2005. Akromystax tilmachiton Gen. Et Sp. Nov., a New Pycnodontid Fish from the Lebanese Late Cretaceous of Haqel and En Nammoura. Journal of Vertebrate Paleontology 25(1):27-45 [S. Peters/S. McMullen]