Araya Formation, Cubagua Island: Late/Upper Pliocene, Venezuela
collected by Landau 2003-2007

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Cancellaria (Bivetiopsia) pachia Smith 1940
Gastropoda - Heterostropha - Architectonicidae
Architectonica nobilis Röding 1798
Gastropoda - Trochidae
Calliostoma (Calliostoma) laticarinatum (Guppy 1867)
Calliostoma (Elmerlinia) sp. Clench and Turner 1960
Calliostoma (Elmerlinia) olssoni (Maury 1925)
Gastropoda - Turbinidae
Astralium brevispina
Turbo (Marmarostoma) castanea (Gmelin 1791)
recombined as Turbo (Senectus) castanea
Gastropoda - Batillariidae
Pyrazisinus sp. Heilprin 1886
Gastropoda - Turritellidae
Vermicularia cf. woodringi Olsson and Harbison 1953
Turritella (Broderiptella) caronensis (Mansfield 1925)
Turritella (Broderiptella) planigyrata (Guppy 1867)
Turritella (Broderiptella) bifastigata (Nelson 1870)
SUBSPECIES: Turritella (Broderiptella) bifastigata
Turritella (Bactrospira) guppyi (Cossmann 1909)
original and current combination Turritella guppyi
Turritella abrupta (Spieker 1922)
Gastropoda - Modulidae
Modulus tamanensis Maury 1925
Gastropoda - Potamididae
Tympanotonos sp. Schumacher 1817
Gastropoda - Neotaenioglossa - Calyptraeidae
Crepidula cf. maculosa Conrad 1846
Crucibulum (Crucibulum) subsutum (Guppy 1867)
original and current combination Crucibulum subsutum
Crucibulum (Crucibulum) sp. Schumacher 1817
Crucibulum (Dispotaea) sp. (Say 1824)
Calyptraea centralis (Conrad 1841)
Gastropoda - Epitoniidae
Scalina brunneopicta (Dall 1908)
Gastropoda - Naticidae
Polinices (Polinices) sp. Montfort 1810
Polinices (Polinices) stanislasmeunieri (Maury 1917)
original and current combination Polinices stanislasmeunieri
Stigmaulax beaumonti (Rutsch 1934)
Naticarius canrena (Linnaeus 1758)
SUBSPECIES: Naticarius canrena canrena
Sinum gabbi (Brown and Pilsbry 1913)
Gastropoda - Neogastropoda - Columbellidae
Strombina (Strombina) cartagenensis (Pilsbry and Brown 1917)
Gastropoda - Neogastropoda - Fasciolariidae
Fusinus cf. springvalensis (Maury 1925)
Fusinus vonderschmidti (Rutsch 1934)
Fasciolaria goragasiana
Gastropoda - Neogastropoda - Nassariidae
Nassarius trinitatensis Jung 1969
Trajana (Nerva) aff. woodringi Vokes 1969
Gordanops baranoanus (Anderson 1929)
Calophos aff. plicatilis (Bose 1906)
Gastropoda - Neogastropoda - Buccinidae
"Hesperisternia" sp. Gardner 1944
Hesperisternia corrugata (Gabb 1873)
Hesperisternia karinae (Nowell-Usticke 1959)
Hesperisternia tortugera (Olsson 1922)
Solenosteira (Fusinosteira) falconensis (Weisbord 1929)
Solenosteira magdalenensis (Weisbord 1929)
Strombinophos perdoctus
Gastropoda - Neogastropoda - Melongenidae
Melongena consors (Sowerby 1850)
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria (Pyruclia) scheibei (Anderson 1929)
recombined as Pyruclia scheibei
Cancellaria (Cancellaria) sp. Lamarck 1799
Cancellaria (Cancellaria) capeloi Landau et al. 2007
Cancellaria (Euclia) codazzii (Anderson 1929)
recombined as Euclia codazzii
Cancellaria (Euclia) montserratensis (Maury 1925)
recombined as Euclia montserratensis
Cancellaria (Euclia) leuzingeri (Rutsch 1934)
recombined as Euclia leuzingeri
Trigonostoma (Ventrilia) rucksorum (Petuch 1994)
original and current combination Ventrilia rucksorum
Cancellaria (Massyla) cubaguensis
original and current combination Massyla
Cancellaria (Charcolleria) terryi Olsson 1942
recombined as Massyla terryi
Gastropoda - Neogastropoda - Mitridae
Subcancilla longa (Gabb 1873)
SUBSPECIES: Subcancilla longa couvensis
Mitra sanctifrancisci Maury 1925
recombined as Fusimitra sanctifrancisci
Gastropoda - Neogastropoda - Drilliidae
Cerodrillia niaddrina
Crassispira (Crassispira) caroniana (Maury 1925)
recombined as Hindsiclava caroniana
Clathrodrillia cf. tityra Woodring 1928
Clathrodrillia gatunensis (Toula 1909)
Gastropoda - Neogastropoda - Terebridae
Strioterebrum weisbordi Gibson-Smith and Gibson-Smith 1984
Terebra aclinica (Olsson 1967)
Terebra lehneri Rutsch 1934
Gastropoda - Neogastropoda - Conidae
Conus walli
Conus oniscus (Woodring 1928)
Conus imitator Brown and Pilsbry 1911
Conus cf. ultimus (Pilsbry and Johnson 1917)
Conus spurius Gmelin 1791
Gastropoda - Neogastropoda - Turridae
Polystira sp. Woodring 1928
Gastropoda - Neogastropoda - Olividae
Oliva tubaraensis
Oliva informal sp. 1 Bruguière 1789
Oliva immortua (Pilsbry and Brown 1917)
Oliva reticularis Lamarck 1811
Gastropoda - Neogastropoda - Ancillariidae
Eburna caroniana (Maury 1925)
Eburna speciosa (Rutsch 1934)
Gastropoda - Neogastropoda - Costellariidae
Knefastia sp. Dall 1919
Fusiturricula springvaleensis (Mansfield 1925)
Conomitra lavelana Hodson 1931
Gastropoda - Neogastropoda - Turbinellidae
Vasum haitense (Sowerby 1850)
recombined as Vasum haitensis
Turbinella trinitatis (Maury 1925)
Gastropoda - Neogastropoda - Marginellidae
Prunum sp. Swainson 1840
Prunum calypsonis (Maury 1925)
Gastropoda - Neogastropoda - Volutidae
Voluta cubaguanensis
Gastropoda - Neogastropoda - Muricidae
Poirieria (Panamurex) eugeniae Vokes 1992
Poirieria (Panamurex) gatunensis (Brown and Pilsbry 1911)
Vokesimurex messorius (Sowerby 1841)
Haustellum mimiwilsoni Vokes 1990
Chicoreus (Chicoreus) cornurectus (Guppy 1876)
recombined as Chicoreus (Triplex) cornurectus
Stramonita "biserialis" (de Blainville 1832)
recombined as Thais biserialis
Coralliophila aff. meyerdorffii
Eupleura muriciformis (Broderip 1833)
Gastropoda - Neogastropoda - Cystiscidae
Persicula sp. Schumacher 1817
Persicula lavelana (Hodson 1927)
Gastropoda - Strombidae
Strombus (Strombus) aff. floridanus Mansfield 1930
Persististrombus granulatus (Swainson 1822)
Gastropoda - Pediculariidae
Jenneria loxahatcheensis (Smith 1936)
Gastropoda - Triviidae
Niveria pediculus (Linnaeus 1758)
recombined as Pusula pediculus
Pusula radians (Lamarck 1810)
SUBSPECIES: Pusula radians orientalis
    = Pusula orientalis Schilder 1939
Landau and Marques da Silva 2010
Gastropoda - Cypraeidae
Zonaria (Zonaria) cf. spurcoides (Gabb 1873)
recombined as Zonaria spurcoides
Muracypraea caroniensis (Maury 1925)
Zonaria (Zonaria) raymondrobertsi (Pilsbry 1922)
recombined as Pseudozonaria raymondrobertsi
Gastropoda - Tonnidae
Malea sp. Valenciennes 1833
    = Malea beui n. sp. Landau and Marques da Silva 2010
Landau and Marques da Silva 2010
Gastropoda - Personidae
Distorsio macgintyi
Distorsio clathrata (Lamarck 1816)
Gastropoda - Bursidae
Marsupina bufo (Bruguiére 1792)
Bursa rugosa (Sowerby 1835)
Gastropoda - Cassidae
Sconsia sublaevigata (Guppy 1866)
synonym of Sconsia grayi
Gastropoda - Ranellidae
Cymatium (Linatella) cingulatum (Lamarck 1822)
synonym of Linatella caudata
Cymatium (Monoplex) krebsii (Mörch 1877)
recombined as Monoplex krebsii
Cymatium (Monoplex) cercadicum (Maury 1917)
recombined as Monoplex cercadicus
Cymatium (Turritriton) domingense (Gabb 1873)
recombined as Turritriton domingensis
Gastropoda - Ficidae
Ficus carbasea (Guppy 1866)
Gastropoda - Vermetidae
Petaloconchus sculpturatus Lea 1846
SUBSPECIES: Petaloconchus sculpturatus alcimus
Serpulorbis decussatus (Gmelin 1791)
Serpulorbis papulosus (Guppy 1866)
Gastropoda - Pseudomelaniidae
Crassispira (Crassispira) cf. tyloessa Woodring 1970
recombined as Crassispira tyloessa
Crassispira (Crassispira) informal sp. 1 Swainson 1840
see common names

Geography
Country:Venezuela
Coordinates: 10.8° North, 64.2° West (view map)
Paleocoordinates:10.7° North, 63.6° West
Basis of coordinate:based on nearby landmark
Geographic resolution:basin
Time
Period:Neogene Epoch:Pliocene
Stage:Piacenzian 10 m.y. bin:Cenozoic 6
Key time interval:Late/Upper Pliocene
Age range of interval:3.60000 - 2.58000 m.y. ago
Stratigraphy
Geological group:Cubagua Formation:Araya
Stratigraphic resolution:formation
Stratigraphy comments: STRATIGRAPHIC RELATIONS: From the Araya Fm of the Cubagua Gp (Late Miocene-Pliocene), which overlies the La Güica Fm. THICKNESS: From a formation that has a maximum thickness of 164 m at its type section. AGE: Late Pliocene on the basis of planktonic foraminifera. Belongs to the Globoratalia miocenica zone(3.2-2.4 Ma) according to Castro and Mederos, 1997. STRATIGRAPHIC POSITION: A composite list for the formation in this region.
Lithology and environment
Primary lithology:fine poorly lithified sandstone
Includes fossils?Y
Lithology description: SPECIFIC LITHOLOGY: Poorly consolidated fossiliferous fine sand. LITHIFICATION: Poorly lithified (or unlithified) on the basis of facies description.
Environment:coastal indet.
Geology comments: ENVIRONMENT: Tropical shallow-marine environment of normal salinity and with a sandy substrate.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Preservation of anatomical detail:good
Collection methods and comments
Collection methods:surface (float),surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Collectors:Landau Collection dates:2003-2007
Collection method comments: COLLECTOR: B. Landau and colleagues, over "last five years". REPOSITORY: Not stated, but possibly Centro de Geologia da Universidad de Lisboa, Camp Grande, Portugal.
Taxonomic list comments:COVERAGE: Exhaustive for gastropoda. Does not include other mollusca. NOMENCLATURE: Not an authoritative publication, but with modern nomenclature and including systematic revision of other neighboring Caribbean assemblages.
Metadata
Database number:79710
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2008-03-18 22:40:59 Last modified:2013-11-25 13:37:23
Access level:authorizer only Released:2009-03-18 22:40:59
Creative Commons license:CC BY
Reference information

Primary reference:
26791. B. Landau, G. Vermeij, and C. M. Silva. 2008. Southern Caribbean Neogene palaebiogeography revisisted. New data from the Pliocene of Cubagua, Venezuela. Palaeogeography, Palaeoclimatology, Palaeoecology 257:445-461 [A. Hendy/A. Hendy]

Secondary references:

36423 B. Landau and C. Marques da Silva. 2010. Early Pliocene gastropods of Cubagua, Venezuela: Taxonomy, palaeobiogeography and ecostratigraphy. Palaeontos 19:1-221 [A. Hendy/A. Hendy]
16604 W. P. Woodring. 1928. Miocene Molluscs from Bowden, Jamaica. Part 2: Gastropods and discussion of results. Contributions to the Geology and Palaeontology of the West Indies [A. Miller/A. Hendy/M. Uhen]