CSUN Loc. 374, Las Llajas Canyon: Lutetian, California

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Echinoidea - Spatangoida - Schizasteridae
Schizaster diabloensis Kew 1920
Squires 1984
Echinoidea - Spatangoida - Brissidae
Eupatagus stevensi n. sp. Grant and Hertlein 1938
Grant and Hertlein 1938
Chondrichthyes - Lamniformes - Odontaspididae
Odontaspis sp. Agassiz 1838
Squires 1984
Chondrichthyes - Lamniformes - Mitsukurinidae
Striatolamia macrota (Agassiz 1838)
Squires 2001
original and current combination Otodus macrotus
Rhynchonellata - Terebratulida - Terebrataliidae
Terebratalia aff. batequia Sandy et al. 1995
Squires 2001
Polychaeta - Sabellida - Serpulidae
Rotularia (Rotularia) tejonensis Arnold 1909
Squires 1984
recombined as Rotularia (Rotularia) tejonense
Cephalopoda - Nautilida - Aturiidae
Aturia myrlae (Hanna 1927)
Squires 1984
Aturia myrlae (Hanna 1927)
Squires 1984
Cephalopoda - Nautilida - Nautilidae
Kummelonautilus sp. Matsumoto 1984
Squires 2001
Scaphopoda - Dentaliida - Dentaliidae
Dentalium stentor Anderson and Hanna 1925
Squires 1984
Gastropoda - Heterostropha - Architectonicidae
Architectonica (Stellaxis) cognata (Gabb 1864)
Squires 1984
Gastropoda - Opisthobranchia - Akeridae
Akera maga Vokes 1939
Squires 2001
Gastropoda - Opisthobranchia - Retusidae
Cylichnina tantilla (Anderson and Hanna 1925)
Squires 1984
Gastropoda - Opisthobranchia - Philinidae
Megistostoma gabbianum (Stoliczka 1868)
Squires 1984
Gastropoda - Turritellidae
Turritella andersoni Dickerson 1916
Squires 1984
as Turritella andersoni lawsoni
Turritella uvasana Conrad 1855
Squires 1984
as Turritella uvasana applinae
Gastropoda - Brachytrematidae
Benoistia umpquaensis Turner 1938
Squires 1984
Gastropoda - Ampullinidae
Eocernina hannibali (Dickerson 1914)
Squires 1984
Pachycrommium clarki (Stewart 1927)
Squires 1984
Gastropoda - Epitoniidae
Cirsotrema sp. Mörch 1852
Squires 1984
Gastropoda - Naticidae
Polinices (Euspira) nuciformis (Gabb 1864)
Squires 1984
Gastropoda - Neogastropoda - Fasciolariidae
Clavilithes informal n. sp. B
Squires 1984
Clavilithes tabulatus (Dickerson 1913)
Squires 1984
Fusinus teglandae (Hanna 1927)
Squires 1984
Gastropoda - Neogastropoda - Buccinidae
Ancistrolepis carolineae n. sp. Squires 1984
Squires 1984
Gastropoda - Neogastropoda - Melongenidae
Ficopsis cooperiana Stewart 1927
Squires 1984
original and current combination Fusus (Hemifusus) cooperii
Gastropoda - Neogastropoda - Clavatulidae
Fusiturricula (Crenaturricula) crenatospira (Cooper 1894)
Squires 1984
recombined as Crenaturricula crenatospira
also as Fusiturricula (Crenaturricula) crenatospira domenginica
Gastropoda - Neogastropoda - Conidae
Conus hornii Gabb 1864
Squires 1984
as Conus hornii umpquaensis
Gastropoda - Neogastropoda - Turridae
? Cochlespiropsis jenkinsi n. sp. Squires 2001
Squires 2001
Surculites mathewsonii (Gabb 1864)
Squires 1984
Gastropoda - Neogastropoda - Volutidae
Lyrischapa lajollaensis (Hanna 1927)
Squires 1984
Lapparia eomagna (Vokes 1939)
Squires 2001
original and current combination Voluta eomagna
Gastropoda - Neogastropoda - Muricidae
Pseudoperissolax blakei (Conrad 1855)
Squires 1984
as Pseudoperissolax blakei praeblakei
Gastropoda - Neogastropoda - Harpidae
Eocithara mutica (Lamarck 1803)
Squires 1984
original and current combination Harpa mutica
as Eocithara mutica californiensis
Gastropoda - Strombidae
Ectinochilus (Macilentos) macilentus (White 1889)
Squires 1984
Gastropoda - Cypraeidae
Bernaya kaylinae n. sp. Groves and Squires 2023
Groves and Squires 2023
Cypraea castacensis Stewart 1927
Squires 1984
    = Subepona leahae n. sp. Groves and Squires 2023
Groves and Squires 2023
Gastropoda - Cymatiidae
Ranella katherineae n. sp. Squires 1983
Olequahia domenginica (Vokes 1939)
Squires 1984
original and current combination Ranella domenginica
Gastropoda - Cassidae
Phalium (Semicassis) tuberculiformis (Gabb 1864)
Squires 1984
Galeodea (Caliagaleodea) californica Clark 1942
Squires 1984
recombined as Galeodea californica
Gastropoda - Ranellidae
Cymatium janetae n. sp. Squires 1983
Cymatium (Septa) janetae Squires 1983
Squires 1984
Gastropoda - Ficidae
Ficopsis remondii (Gabb 1864)
Squires 1984
recombined as Ficus remondii
as Ficopsis remondii crescentensis
Gastropoda - Xenophoridae
Xenophora stocki Dickerson 1916
Squires 1984
Bivalvia - Mytilida - Mytilidae
Brachidontes (Brachidontes) cowlitzensis (Weaver and Palmer 1922)
Squires 1984
Bivalvia - Pectinida - Spondylidae
Spondylus carlosensis Anderson 1905
Squires 1984
Bivalvia - Ostreida - Malleidae
Nayadina (Exputens) llajasensis (Clark 1934)
Squires 1984
Bivalvia - Ostreida - Pinnidae
Pinna llajasensis n. sp. Squires 1983
Bivalvia - Ostreida - Ostreidae
Ostrea idriaensis Gabb 1869
Squires 1984
recombined as Acutostrea idriaensis
Bivalvia - Ostreida - Gryphaeidae
Pycnodonte (Phygraea) cf. pacifica Squires and Demetrion 1990
Squires 2001
recombined as Phygraea pacifica
Bivalvia - Arcida - Glycymerididae
Glycymeris (Glycymerita) sagittata (Gabb 1864)
Squires 1984
recombined as Glycymerita sagittata
Bivalvia - Cardiida - Cardiidae
Acanthocardia (Agnocardia) sorrentoensis (Hanna 1927)
Squires 2001
recombined as Agnocardia sorrentoensis
Acanthocardia (Schedocardia) brewerii (Gabb 1864)
Squires 1984
recombined as Schedocardia brewerii
Nemocardium linteum (Conrad 1855)
Squires 1984
recombined as Nemocardium (Nemocardium) linteum
Bivalvia - Cardiida - Psammobiidae
Gari texta Gabb 1864
Squires 2001
recombined as Gari (Gobraeus) texta
Gari cf. eoundulata Vokes 1939
Squires 1984
recombined as Gari (Gobraeus) eoundulata
Bivalvia - Cardiida - Veneridae
Marcia (Mercimonia) bunkeri (Hanna 1927)
Squires 1984
Pitar uvasana (Conrad 1855)
Squires 2001
Subspecies: Pitar uvasana coquillensis
Pitar (Calpitaria) uvasanus (Conrad 1855)
Squires 1984
recombined as Pitar uvasana
Bivalvia - Pholadida - Teredinidae
? Teredo sp. Linnaeus 1758
Squires 1984
Bivalvia - Solenida - Solenidae
Solena (Eosolen) navacularis (Anderson and Hanna 1925)
Squires 1984
recombined as Solena (Eosolen) novacularis
Bivalvia - Lucinida - Lucinidae
Claibornites diegoensis (Dickerson 1916)
Squires 1984
recombined as Claibornites (Claibornites) diegoensis
Bivalvia - Carditida - Crassatellidae
Crassatella uvasana Conrad 1855
Squires 1984
Bivalvia - Carditida - Carditidae
Venericardia (Pacificor) hornii (Gabb 1864)
Squires 1984
as Venericardia (Pacificor) hornii calafia
Anthozoa - Scleractinia - Caryophylliidae
? Trochocyathus striatus (Gabb 1864)
Squires 1984
Anthozoa - Scleractinia - Oculinidae
Archohelia clarki Vaughan 1927
Squires 2001
recombined as Petrophyllia clarki
Asterocyclinidae
Pseudophragmina (Proporocyclina) clarki (Cushman 1920)
Squires 1984
recombined as Pseudophragmina clarki
see common names

Geography
Country:United States State/province:California County:Ventura
Coordinates: 34.3° North, 118.7° West (view map)
Paleocoordinates:35.1° North, 104.6° West
Basis of coordinate:estimated from map
Geographic resolution:small collection
Time
Period:Paleogene Epoch:Eocene
Stage:Lutetian 10 m.y. bin:Cenozoic 2
*Period:Early/Lower Tertiary - Middle Tertiary *Epoch:Early/Lower Eocene - Middle Eocene
*International age/stage:Middle Lutetian - Early/Lower Ypresian *Local age/stage:Domengine - Domengine
Key time interval:Lutetian
Age range of interval:47.80000 - 41.20000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Llajas
Stratigraphic resolution:bed
Stratigraphy comments: The Stewart bed is a 1 m thick sandstone unit in the middle of the Llajas formation. Lutetian, chron 21r according to Squires (2001).
Lithology and environment
Primary lithology:bioturbation lithified sandstone
Includes fossils?Y
Lithology description: sandstone.
Environment:transition zone/lower shoreface
Geology comments: The "Stewart bed" is located in a section of transition from shallow-marine to outer slope and shelf facies. A nearshore environment is interpreted for that portion of the shallow-marine (transgressive) facies which consists of laminated beds alternating with bioturbated beds. In such cases, the finely laminated sand represents storm-influenced stratification, and the biogenic reworking is the inter-storm activity. Shell fragments commonly are concentrated as channel lags in surge channels. Fossils were apparently subjected to wave action because the apices of elongate shells such as Turritella are bimodally oriented in opposite directions. The deeper the water, the less amount of laminated sandstone and the more amount of bioturbated sandstone. Such areas, which were farther below effective wave base, existed 1) where the shallow-marine (transgressive) facies grades into the outer shelf to slope facies and 2) in the shallow-marine (regressive) facies. One of the deepest-water shallow-marine beds was the "Stewart bed."
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:bulk,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:LACM
Collection method comments: Repository: formerly CSUN (California State University Northridge)
Metadata
Also known as:LACMIP loc. 7242; LACMIP loc. 40374
Database number:8012
Authorizer:L. Ivany, A. Hendy, J. Marcot, M. Clapham Enterer:E. Kowalski, A. Hendy, J. Marcot, M. Clapham
Modifier:J. Marcot Research group:marine invertebrate
Created:2000-06-12 07:08:25 Last modified:2023-11-19 19:13:00
Access level:the public Released:2000-06-12 07:08:25
Creative Commons license:CC BY
Reference information

Primary reference:
519. R. L. Squires. 1983. New Mollusks from the Lower Middle Eocene Llajas Formation, Southern California. Journal of Paleontology 57(2):354-362 [L. Ivany/E. Kowalski/J. Marcot]

Secondary references:

32966 U. S. Grant and L. G. Hertlein. 1938. The West American Cenozoic Echinoidea. Publications of the University of California at Los Angeles in Mathematical and Physical Sciences 2:1-225 [L. Villier/L. Villier]
86508 L. T. Groves and R. L. Squires. 2023. Revision of northeast Pacific Paleogene cypraeoidean gastropods (Mollusca), including recognition of three new species: Implications for paleobiogeographic distribution and faunal turnover. PaleoBios 40(10):1-52 [J. Marcot/J. Marcot]
59877 R. L. Squires. 1984. Megapaleontology of the Eocene Llajas Formation, Simi Valley, California. Contributions in Science, Natural History Museum of Los Angeles County 350:1-76 [M. Clapham/M. Clapham]
13852 R. L. Squires. 2001. Additions to the Eocene megafossil fauna of the Llajas Formation, Simi Valley, southern California. Contributions in Science (Natural History Museum of Los Angeles County) 489:1-40 [A. Miller/A. Hendy/A. Hendy]