Tham Khuyen: Middle Pleistocene, Vietnam
collected by T. K. Le, V. B. Tran 1965, 1966

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Proboscidea - Elephantidae
Stegodon orientalis Owen 1870
Schwartz et al. 1994
synonym of Stegodon insignis
Mammalia - Chiroptera
Chiroptera indet. Blumenbach 1779
Schwartz et al. 1994
Mammalia - Perissodactyla - Rhinocerotidae
Rhinoceros sinensis Owen 1870
Schwartz et al. 1994
Mammalia - Perissodactyla - Tapiridae
Tapirus sp. Brisson 1762
Schwartz et al. 1994
Tapirus (Megatapirus) augustus Matthew and Granger 1923
Schwartz et al. 1994
Mammalia - Cervidae
Muntiacus muntjac (Rafinesque 1815)
Schwartz et al. 1994
original and current combination Muntiacus muntjak
Cervus sp. Linnaeus 1758
Schwartz et al. 1994
Rusa unicolor (Kerr 1792)
Schwartz et al. 1994
original and current combination Cervus unicolor
Mammalia - Bovidae
Bibos gaurus
Schwartz et al. 1994
Bubalus bubalis (Linnaeus 1758)
Schwartz et al. 1994
Mammalia - Suidae
Sus sp. Linnaeus 1758
Schwartz et al. 1994
Sus scrofa Linnaeus 1758
Schwartz et al. 1994
Sus cf. lydekkeri Zdansky 1928
Schwartz et al. 1994
Mammalia - Carnivora - Viverridae
Paradoxurus cf. hermaphroditus (Pallas 1777)
Schwartz et al. 1994
Mammalia - Carnivora - Ursidae
Ursus malayanus Raffles 1821
Schwartz et al. 1994
Ursus thibetanus Cuvier 1823
Schwartz et al. 1994
"Ursus thibetanus kokeni"
Ailuropoda melanoleuca (David 1869)
Schwartz et al. 1994
"Ailuropoda melanoleuca fovealis"
Mammalia - Carnivora - Mustelidae
Arctonyx collaris Cuvier 1825
Schwartz et al. 1994
"Arctonyx collaris cf. rostratus"
Mammalia - Carnivora - Canidae
Cuon sp. Hodgson 1838
Schwartz et al. 1994
Mammalia - Rodentia - Hystricidae
Hystrix subcristata
Schwartz et al. 1994
Hystrix sp. Linnaeus 1758
Schwartz et al. 1994
Atherurus sp. Cuvier 1829
Schwartz et al. 1994
Mammalia - Rodentia - Muridae
Mus sp. Linnaeus 1758
Schwartz et al. 1994
Rattus sp. Fischer 1803
Schwartz et al. 1994
Mammalia - Rodentia - Spalacidae
Rhizomys cf. troglodytes Matthew and Granger 1923
Schwartz et al. 1994
Mammalia - Primates
Hominoidea indet. Gray 1825
Schwartz et al. 1994
said to be a new genus and "large-bodied"
Mammalia - Primates - Hylobatidae
Hylobates cf. concolor (Harlan 1826)
Schwartz et al. 1994
recombined as Nomascus concolor
Mammalia - Primates
Langsonia liquidens n. gen., n. sp. Schwartz et al. 1995
Schwartz et al. 1995
Mammalia - Primates - Hominidae
Gigantopithecus blacki von Koenigswald 1935
Schwartz et al. 1994
Pongo sp. Lacépède 1799
Schwartz et al. 1994
"dentally primitive"
Pongo pygmaeus (Linnaeus 1760)
Schwartz et al. 1994
subspecies kahlkei
Pongo hooijeri n. sp. Schwartz et al. 1995
Schwartz et al. 1995
Homo sp. Linnaeus 1758
Schwartz et al. 1994
Homo erectus (Dubois 1892)
Mammalia - Primates - Cercopithecidae
Macaca sp. Lacépède 1799
Schwartz et al. 1994
Macaca cf. assamensis (MacClelland 1839)
Schwartz et al. 1994
see common names

Geography
Country:Vietnam State/province:Lang Son
Coordinates: 22.0° North, 106.4° East (view map)
Paleocoordinates:21.9° North, 106.4° East
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period:Quaternary Epoch:Pleistocene
Stage:Chibanian 10 m.y. bin:Cenozoic 6
Key time interval:Middle Pleistocene
Age range of interval:0.77400 - 0.12900 m.y. ago
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: from "Two distinct breccia layers, a lower light red unit and an overlying darker red one... the darker one [contains] most of the fossil vertebrate remains... there is no sharp discontinuity between the layers, which were deposited during the same period... faunal analyses... suggest a late middle Pleistocene age (probably around 300-250 Ka) for the entire contained mammalian assemblage"
Lithology and environment
Primary lithology:shelly/skeletal,red lithified breccia
Lithology description: "heavily calcified breccias that are rich in fossil bone and teeth" including "cobbles and small boulders"
Environment:cave
Geology comments: "periodic flooding appears to have washed river sediments ande remains of vertebrates into the cavity... A relatively high energy environment"
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Fragmentation:frequent
Collection methods and comments
Collection methods:bulk,selective quarrying,mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:T. K. Le, V. B. Tran Collection dates:1965, 1966
Collection method comments: collected by "a joint Vietnamese and German (DDR) group... Blocks... were removed... using hammers and chisels, following which manual preparation of the specimens was completed in the laboratories of the Institute of Archaeology in Hanoi"
Metadata
Database number:92781
Authorizer:J. Alroy, M. Uhen Enterer:J. Alroy, M. Uhen
Modifier:M. Uhen Research group:vertebrate
Created:2009-11-28 16:06:45 Last modified:2022-11-14 10:15:23
Access level:the public Released:2009-11-28 16:06:45
Creative Commons license:CC BY
Reference information

Primary reference:

83097. R. Ciochon, V. T. Long, R. Larick, L. González, R. Grün, J. De Vos, C. Yonge, L. Taylor, H. Yoshida and M. Regan. 1996. Dated co-occurrence of Homo erectus and Gigantopithecus from Tham Khuyen Cave, Vietnam. Proceedings of the National Academy of Science, USA 93:3016-3020 [M. Uhen/M. Uhen]

Secondary references:

35877 J. H. Schwartz, V. T. Long, N. L. Cuong, L. T. Kha, and I. Tattersall. 1995. A review of the Pleistocene hominoid fauna of the Socialist Republic of Vietnam (excluding Hylobatidae). Anthropological Paper of the American Museum of Natural History 76:1-23 [M. Uhen/M. Uhen]
31293 J. H. Schwartz, T. H. Vu, L. C. Nguyen, K. T. Le, and I. Tattersall. 1994. A diverse hominoid fauna from the late middle Pleistocene breccia cave of Tham Khuyen, Socialist Republic of Vietnam. Anthropological Papers of the American Museum of Natural History 73:1-11 [J. Alroy/J. Alroy]