Woodring Loc. 139e - eastern area [lower Gatun Fm]: Tortonian, Panama

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Lirophora (Parachione) mactropsis Conrad 1855
C category
Scaphopoda - Gadilida - Gadilidae
Cadulus (Platyschides) epetrion
Woodring 1973 C category
Scaphopoda - Dentaliida - Dentaliidae
Dentalium (Dentalium) armillatum
Woodring 1973 R category
Gastropoda - Pseudomelaniidae
Carinodrillia (Carinodrillia) zooki (Brown and Pilsbry 1911)
Woodring 1970 R category
recombined as Carinodrillia zooki
Gastropoda - Turritellidae
Turritella abrupta (Spieker 1922)
Woodring 1970 R category
Gastropoda - Neogastropoda - Marginellidae
Volvarina leander (Brown and Pilsbry 1911)
Woodring 1970 R category
Gastropoda - Neogastropoda - Olividae
Oliva (Oliva) gatunensis (Toula 1908)
Woodring 1964 C category
original and current combination Oliva gatunensis
Olivella (Niteoliva) terryi Olsson 1956
Woodring 1964 R category
Gastropoda - Neogastropoda - Melongenidae
Melongena melongena (Linnaeus 1758)
Woodring 1964 R category
Gastropoda - Neogastropoda - Nassariidae
Nassarius (Nassarius) parapristus (Gardner 1944)
Woodring 1964 R category
Calophos ectyphus Woodring 1964
Woodring 1964 A category
Gastropoda - Neogastropoda - Buccinidae
Cymatophos veatchi (Olsson 1922)
Woodring 1964 R category
SUBSPECIES: Cymatophos veatchi veatchi
Gastropoda - Neogastropoda - Fasciolariidae
Fasciolaria gorgasiana Brown and Pilsbry 1913
Woodring 1964 R category
recombined as Pleuroploca gorgasiana
Gastropoda - Neogastropoda - Drilliidae
Agladrillia (Agladrillia) acaria (Mansfield 1925)
Woodring 1970 R category
SUBSPECIES: Agladrillia (Agladrillia) acaria ectypha
Clathrodrillia gatunensis (Toula 1909)
Woodring 1970 R category
Crassispira (Hindsiclava) consors (Sowerby 1850)
Woodring 1970 R category
recombined as Hindsiclava consors
Gastropoda - Neogastropoda - Turridae
Pleurolira (Polystira) tenagos Gardner 1947
Woodring 1970 C category
    = Polystira sp. Woodring 1928
Todd and Johnson 2013
Gastropoda - Neogastropoda - Terebridae
Strioterebrum oresignum Olsson 1964
Woodring 1970 F category
Terebra (Panaterebra) cucurrupiensis (Oinomikado 1939)
Woodring 1970 F category
Gastropoda - Neogastropoda - Conidae
Conus aemulator Brown and Pilsbry 1911
Woodring 1970 F category
Conus burckhardti Bose 1906
Woodring 1970 F category
Woodring 1970 R category
Conus multiliratus (Olsson 1922)
Woodring 1970 R category
recombined as Conasprelloides multiliratus
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria (Euclia) codazzii (Anderson 1929)
Woodring 1970 R category
recombined as Euclia codazzii
Cancellaria (Cancellaria) epistomifera (Guppy 1876)
Woodring 1970 R category
SUBSPECIES: Cancellaria (Cancellaria) epistomifera dariensis
    = Bivetiella dariena Toula 1908
Landau et al. 2012
Gastropoda - Neogastropoda - Turbinellidae
Xancus validus (Sowerby 1850)
Woodring 1964 R category
recombined as Turbinella valida
SUBSPECIES: Xancus validus falconensis
Xancus dodonaius
Woodring 1973 R category
replaced by Turbinella
SUBSPECIES: Xancus dodonaius praelaeviagtus
Bivalvia - Nuculanida - Nuculanidae
Saccella acrita (Dall 1908)
Woodring 1973 R category
SUBSPECIES: Saccella acrita epacra
Bivalvia - Ostreida - Gryphaeidae
Hyotissa haitensis (Sowerby 1850)
C category
Bivalvia - Ostreida - Pteriidae
Pteria inornata (Gabb 1873)
Woodring 1973 R category
Bivalvia - Arcida - Arcidae
Anadara (Rasia) dariensis (Brown and Pilsbry 1911)
Woodring 1973 A category
Bivalvia - Lucinida - Lucinidae
Bellucina actinus (Dall 1903)
R category
Bivalvia - Pholadida - Corbulidae
Caryocorbula oropendula (Olsson 1922)
F category
SUBSPECIES: Caryocorbula oropendula stena
Varicorbula disparilis (d'Orbigny 1853)
R category
Bivalvia - Cardiida - Kelliellidae
Alveinus rotundus Dall 1900
R category
Bivalvia - Cardiida - Veneridae
Macrocallista (Paradione) maculata (Linnaeus 1758)
F category
recombined as Macrocallista maculata
Clementia (Clementia) dariena (Conrad 1855)
C category
SUBSPECIES: Clementia (Clementia) dariena dariena
Cyclinella cyclica (Guppy 1866)
F category
Lirophora (Lirophora) falconensis (Hodson et al. 1927)
R category
Chione (Chionopsis) tegulum (Brown and Pilsbry 1911)
R category
Agriopoma (Pitarella) gatunensis (Dall 1903)
C category
recombined as Callocardia (Agriopoma) gatunensis
Bivalvia - Cardiida - Chamidae
Arcinella acrinella
F category
Chama berjadinensis Hodson 1927
C category
Bivalvia - Cardiida - Cardiidae
Trigoniocardia (Apiocardia) aminensis (Dall 1900)
R category
recombined as Trigoniocardia (Trigoniocardia) aminensis
Trachycardium (Dallocardia) baiterum Woodring 1982
C category
Bivalvia - Cardiida - Semelidae
Ervilia valhosierr Gardner 1928
R category
Bivalvia - Solenida - Solenidae
Solen (Solena) obliquus (Spengler 1794)
F category
SUBSPECIES: Solen (Solena) obliquus rudis
see common names

Geography
Country:Panama
Coordinates: 9.3° North, 79.8° West (view map)
Paleocoordinates:9.1° North, 78.5° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Tortonian 10 m.y. bin:Cenozoic 6
Key time interval:Tortonian
Age range of interval:11.63000 - 7.24600 m.y. ago
Age estimate:10.9 Ma (other)
Stratigraphy
Formation:Gatun Member:lower Gatun
Stratigraphic resolution:bed
Stratigraphy comments: Lower part of the Gatun Formation. The Gatun Formation is sibdivided into three faunal zones (proposed by Thomson and Keen, 1946): lower, middle, and upper. AGE: Late Miocene, on the basis of microfossil biostratigraphy (see Collins and Coates, 1999), refined by enterer to Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 10.9 Ma (Hendy, unpublished data).
Lithology and environment
Primary lithology:fine unlithified silty sandstone
Includes fossils?Y
Lithology description: LITHOLOGY: Soft, silty fine-grained sandstone.
Environment:marine indet.
Geology comments: No specific environmental data presented.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Museum repositories:USNM
Collection method comments: COLLECTOR: Not stated in text. REPOSITORY: USNM.
Taxonomic list comments:COVERAGE: Exhaustive for gastropods and bivalves. NOMENCLATURE: Authoritative publications, with modern nomenclature, and species-resolution identifications.
Metadata
Also known as:USGS #22019
Database number:93162
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2010-01-06 07:57:03 Last modified:2013-11-25 13:37:23
Access level:the public Released:2010-01-06 07:57:03
Creative Commons license:CC BY
Reference information

Primary reference:

13359. W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

43491 B. Landau, R. E. Petit, and C. M. Silva. 2012. New Cancellariidae (Mollusca, Gastropoda) from the Miocene Gatun Formation of Panama, with eleven new species. Journal of Paleontology 86(6):907-930 [A. Hendy/A. Hendy]
48647 J. A. Todd and K. G. Johnson. 2013. Dissecting a marine snail species radiation (Conoidea: Turridae: Polystira) over 12 million years in the southwestern Caribbean. Bulletin of Marine Science 89(4) [A. Hendy/A. Hendy]
13232 W. P. Woodring. 1964. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Columbellidae to Volutidae). United States Geological Survey Professional Paper 306(C):241-297 [A. Miller/A. Hendy/A. Hendy]
13236 W. P. Woodring. 1970. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). United States Geological Survey Professional Paper 306(D):299-452 [A. Miller/A. Hendy/A. Hendy]
13235 W. P. Woodring. 1973. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae). United States Geological Survey Professional Paper 306(E):453-539 [A. Miller/A. Hendy/A. Hendy]