Woodring Loc. 155b - eastern area [middle Gatun Fm]: Tortonian, Panama
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
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Gastropoda
- Turbinidae
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Turbo (Marmarostoma) aff. castaneus
(Gmelin 1791)
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| recombined as Turbo (Senectus) castanea | ||||||||||
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Gastropoda
- Opisthobranchia
- Retusidae
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Volvulella (Volvulella) oxytata
(Bush 1885)
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Woodring 1970 | R category | ||||||||
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Gastropoda
- Heterostropha
- Architectonicidae
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Architectonica (Architectonica) nobilis
(Röding 1798)
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Woodring 1959 | C category | ||||||||
| original and current combination Architectonica nobilis | ||||||||||
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Gastropoda
- Neotaenioglossa
- Calyptraeidae
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Crucibulum (Crucibulum) chipolanum
(Dall 1892)
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Crucibulum (Dispotaea) springvaleense
(Rutsch 1942)
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Gastropoda
- Naticidae
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Polinices brunneus
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| SUBSPECIES: Polinices brunneus subclausus | ||||||||||
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Neverita (Hypterita) helicoides
(Gray 1825)
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| synonym of Amauropsis islandica | ||||||||||
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Stigmaulax guppiana
(Toula 1909)
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Natica (Natica) bolus
Brown and Pilsbry 1913
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Gastropoda
- Neogastropoda
- Cancellariidae
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Cancellaria (Cancellaria) epistomifera
(Guppy 1876)
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Woodring 1970 | C category | ||||||||
| SUBSPECIES: Cancellaria (Cancellaria) epistomifera dariena | ||||||||||
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= Bivetiella dariena
Toula 1908
|
Landau et al. 2012 | |||||||||
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Cancellaria (Charcolleria) terryi
Olsson 1942
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Woodring 1970 | R category | ||||||||
| recombined as Massyla terryi | ||||||||||
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Gastropoda
- Neogastropoda
- Terebridae
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Strioterebrum spiniferum
|
Woodring 1970 | C category | ||||||||
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Strioterebrum gausapatum
(Brown and Pilsbry 1911)
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Woodring 1970 | R category | ||||||||
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Terebra (Oreoterebra) isaacpetiti
(Maury 1925)
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Woodring 1970 | R category | ||||||||
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Terebra (Oreoterebra) subsulcifera
(Brown and Pilsbry 1911)
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Woodring 1970 | F category | ||||||||
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Gastropoda
- Neogastropoda
- Conidae
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Conus imitator
Brown and Pilsbry 1911
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Woodring 1970 | F category | ||||||||
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Conus molis
Brown and Pilsbry 1911
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Woodring 1970 | C category | ||||||||
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Conus aemulator
Brown and Pilsbry 1911
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Woodring 1970 | C category | ||||||||
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Conus burckhardti
Bose 1906
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Woodring 1970 | C category | ||||||||
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Gastropoda
- Neogastropoda
- Turridae
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Gemmula vaningeni
(Brown and Pilsbry 1913)
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Woodring 1970 | R category | ||||||||
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Pleurofusia (Cruziturricula) fusinus
(Brown and Pilsbry 1911)
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Woodring 1970 | F category | ||||||||
| recombined as Cruziturricula fusinus | ||||||||||
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Pleurolira (Polystira) tenagos
Gardner 1947
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Woodring 1970 | C category | ||||||||
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= Polystira sp.
Woodring 1928
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Todd and Johnson 2013 | |||||||||
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Gastropoda
- Neogastropoda
- Drilliidae
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Clathrodrillia gatunensis
(Toula 1909)
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Woodring 1970 | F category | ||||||||
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Crassispira (Hindsiclava) consors
(Sowerby 1850)
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Woodring 1970 | F category | ||||||||
| recombined as Hindsiclava consors | ||||||||||
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Agladrillia (Eumetadrillia) isthmica
(Brown and Pilsbry 1911)
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Woodring 1970 | R category | ||||||||
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Gastropoda
- Neogastropoda
- Melongenidae
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Melongena melongena
(Linnaeus 1758)
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Woodring 1964 | R category | ||||||||
| SUBSPECIES: Melongena melongena consors | ||||||||||
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Gastropoda
- Neogastropoda
- Buccinidae
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Antillophos (Rhipophos) metuloides
(Dall 1896)
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Woodring 1964 | R category | ||||||||
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Antillophos (Antillophos) candei
(d'Orbigny 1842)
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Woodring 1964 | A category | ||||||||
| recombined as Antillophos candeanus | ||||||||||
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Metula gabbi
Brown and Pilsbry 1911
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Woodring 1964 | F category | ||||||||
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Gastropoda
- Neogastropoda
- Fasciolariidae
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Fasciolaria gorgasiana
Brown and Pilsbry 1913
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Woodring 1964 | R category | ||||||||
| recombined as Pleuroploca gorgasiana | ||||||||||
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Gastropoda
- Neogastropoda
- Columbellidae
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Strombina (Strombina) lessepsiana
(Brown and Pilsbry 1911)
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Woodring 1964 | F category | ||||||||
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Gastropoda
- Neogastropoda
- Olividae
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Oliva (Oliva) cristobalcoloni
(Maury 1917)
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Woodring 1964 | R category | ||||||||
| original and current combination Oliva cristobalcoloni | ||||||||||
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Oliva (Oliva) gatunensis
(Toula 1908)
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Woodring 1964 | A category | ||||||||
| original and current combination Oliva gatunensis | ||||||||||
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Gastropoda
- Neogastropoda
- Olivellidae
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Agaronia testacea
(Lamarck 1811)
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Woodring 1964 | R category | ||||||||
| SUBSPECIES: Agaronia testacea mancinella | ||||||||||
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Gastropoda
- Neogastropoda
- Marginellidae
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Volvarina leander
(Brown and Pilsbry 1911)
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Woodring 1970 | R category | ||||||||
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Gastropoda
- Neogastropoda
- Muricidae
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Murex (Chicoreus) brevifrons
(Lamarck 1822)
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Woodring 1959 | R category | ||||||||
| recombined as Chicoreus (Chicoreus) brevifrons | ||||||||||
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Murex (Murex) polynematicus
(Brown and Pilsbry 1911)
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Woodring 1959 | R category | ||||||||
| recombined as Vokesimurex polynematicus | ||||||||||
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Murex (Murex) recurvirostris
(Broderip 1832)
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Woodring 1959 | R category | ||||||||
| recombined as Vokesimurex recurvirostris | ||||||||||
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Paziella (Panamurex) gatunensis
(Brown and Pilsbry 1911)
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Woodring 1959 | R category | ||||||||
| recombined as Poirieria (Panamurex) gatunensis | ||||||||||
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Gastropoda
- Strombidae
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Strombus gatunensis
Toula 1908
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Woodring 1959 | R category | ||||||||
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Gastropoda
- Bursidae
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Bursa (Colubrellina) caelata
(Broderip 1833)
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Woodring 1959 | R category | ||||||||
| SUBSPECIES: Bursa (Colubrellina) caelata amphritites | ||||||||||
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= Bursa rugosa
Sowerby 1835
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Landau and Marques da Silva 2010 | |||||||||
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Gastropoda
- Cassidae
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Sconsia laevigata
(Guppy 1850)
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Woodring 1959 | R category | ||||||||
| SUBSPECIES: Sconsia laevigata sublaevigata | ||||||||||
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= Sconsia grayi
Adams 1855
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Beu 2010 | |||||||||
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Gastropoda
- Personidae
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Distorsio (Rhysema) decussata
(Valenciennes 1832)
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Woodring 1959 | R category | ||||||||
| recombined as Distorsio decussata | ||||||||||
| SUBSPECIES: Distorsio (Rhysema) decussata gatunensis | ||||||||||
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Gastropoda
- Turritellidae
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Turritella (Torcula) mimetes
Brown and Pilsbry 1911
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| recombined as Turritella (Broderiptella) mimetes | ||||||||||
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Turritella (Torcula) gatunensis
Conrad 1857
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| SUBSPECIES: Turritella (Torcula) gatunensis gatunensis | ||||||||||
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Turritella (Torcula) altilira
(Conrad 1857)
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Gastropoda
- Cerithiidae
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Rhinoclavis (Ochetoclava) costaricana
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Woodring 1959 | R category | ||||||||
| SUBSPECIES: Rhinoclavis (Ochetoclava) costaricana canabina | ||||||||||
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Bivalvia
- Arcida
- Glycymerididae
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Glycymeris (Tucetona) pectinata
(Gmelin 1791)
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Woodring 1973 | C category | ||||||||
| recombined as Tucetona pectinata | ||||||||||
| SUBSPECIES: Glycymeris (Tucetona) pectinata canalis | ||||||||||
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Glycymeris (Glycymeris) carbasina
Brown and Pilsbry 1911
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Woodring 1973 | C category | ||||||||
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Bivalvia
- Arcida
- Arcidae
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Anadara (Rasia) actinophora
(Dall 1898)
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Woodring 1973 | R category | ||||||||
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Anadara (Rasia) dariensis
(Brown and Pilsbry 1911)
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Woodring 1973 | C category | ||||||||
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Bivalvia
- Pectinida
- Pectinidae
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Flabellipecten gatunensis
(Toula 1909)
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Woodring 1982 | F category | ||||||||
| recombined as Leopecten gatunensis | ||||||||||
| SUBSPECIES: Flabellipecten gatunensis gatunensis | ||||||||||
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Amusium toulae
(Brown and Pilsbry 1911)
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Woodring 1982 | F category | ||||||||
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Argopecten levicostatus
(Toula 1909)
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Woodring 1982 | F category | ||||||||
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Lepidopecten scissuratus
(Dall 1898)
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Woodring 1982 | C category | ||||||||
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Bivalvia
- Ostreida
- Gryphaeidae
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Hyotissa haitensis
(Sowerby 1850)
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Woodring 1982 | R category | ||||||||
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Bivalvia
- Lucinida
- Lucinidae
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Bellucina actinus
(Dall 1903)
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Woodring 1982 | F category | ||||||||
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Pegophysema janus
(Dall 1903)
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Woodring 1982 | F category | ||||||||
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Bivalvia
- Pholadida
- Corbulidae
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Caryocorbula oropendula
(Olsson 1922)
|
Woodring 1982 | C category | ||||||||
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Bothrocorbula (Hexacorbula) gatunensis
(Toula 1909)
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Woodring 1982 | R category | ||||||||
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Varicorbula islatrinitatis
(Maury 1925)
|
Woodring 1982 | C category | ||||||||
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Bivalvia
- Cardiida
- Veneridae
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Dosinia (Dosinia) delicatissim
|
Woodring 1982 | R category | ||||||||
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Lirophora (Panchione) mactropsis
(Conrad 1855)
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Woodring 1982 | A category | ||||||||
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Chione (Chionopsis) tegulum
(Brown and Pilsbry 1911)
|
Woodring 1982 | R category | ||||||||
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Pitar (Hyphantosoma) centangulatus
(Brown and Pilsbry 1911)
|
Woodring 1982 | R category | ||||||||
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Agriopoma (Pitarella) gatunensis
(Dall 1903)
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Woodring 1982 | C category | ||||||||
| recombined as Callocardia (Agriopoma) gatunensis | ||||||||||
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Cyclinella atacata
|
Woodring 1970 | C category | ||||||||
| SUBSPECIES: Cyclinella atacata stibara | ||||||||||
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Gouldia costaricensis
(Olsson 1922)
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Woodring 1982 | R category | ||||||||
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Bivalvia
- Cardiida
- Chamidae
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Arcinella arcinella
(Linnaeus 1767)
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Woodring 1982 | C category | ||||||||
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Bivalvia
- Cardiida
- Cardiidae
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Lophocardium gurabicum
(Maury 1917)
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Woodring 1982 | R category | ||||||||
| recombined as Nemocardium (Lophocardium) gurabicum | ||||||||||
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Trachycardium (Dallocardia) dominicense
(Gabb 1873)
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Woodring 1982 | R category | ||||||||
| SUBSPECIES: Trachycardium (Dallocardia) dominicense dominicense | ||||||||||
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Trachycardium (Phlogocardia) stiriatum
(Brown and Pilsbry 1911)
|
Woodring 1982 | C category | ||||||||
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Bivalvia
- Cardiida
- Tellinidae
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Psammacoma gatunensis
(Toula 1909)
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Woodring 1982 | R category | ||||||||
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Bivalvia
- Carditida
- Crassatellidae
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Eucrassatella (Eucrassatella) sp.
Iredale 1924
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Woodring 1982 | F category | ||||||||
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Bivalvia
- Carditida
- Carditidae
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Venericardia (Glyptoactis) aversa
(Pilsbry and Johnson 1917)
|
Woodring 1982 | R category | ||||||||
| see common names | ||||||||||
Geography
| Country: | Panama |
| Coordinates: | 9.3° North, 79.9° West (view map) |
| Paleocoordinates: | 9.0° North, 78.5° West |
| Basis of coordinate: | stated in text |
| Geographic resolution: | outcrop |
Time
| Period: | Neogene | Epoch: | Miocene |
| Stage: | Tortonian | 10 m.y. bin: | Cenozoic 6 |
| Key time interval: | Tortonian | ||
| Age range of interval: | 11.63000 - 7.24600 m.y. ago | ||
| Age estimate: | 9.9 Ma (other) |
Stratigraphy
| Formation: | Gatun | Member: | middle Gatun | ||
| Stratigraphic resolution: | bed | ||||
| Stratigraphy comments: Middle part of the Gatun Formation. The Gatun Formation is subdivided into three faunal zones (proposed by Thomson and Keen, 1946): lower, middle, and upper. AGE: Late Miocene, on the basis of microfossil biostratigraphy (see Collins and Coates, 1999), refined by enterer to Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 9.9 Ma (Hendy, unpublished data). | |||||
Lithology and environment
| Primary lithology: | fine unlithified silty sandstone |
| Includes fossils? | Y |
| Lithology description: LITHOLOGY: Soft, silty fine-grained sandstone. | |
| Environment: | marine indet. |
| Geology comments: No specific environmental data presented. | |
Taphonomy
| Modes of preservation: | body |
| Size of fossils: | macrofossils |
Collection methods and comments
| Collection methods: | surface (in situ),field collection |
| Reason for describing collection: | taxonomic analysis |
| Museum repositories: | USNM |
| Collection method comments: COLLECTOR: Not stated in text. REPOSITORY: USNM. | |
| Taxonomic list comments:COVERAGE: Exhaustive for gastropods and bivalves. NOMENCLATURE: Authoritative publications, with modern nomenclature, and species-resolution identifications. | |
Metadata
| Also known as: | USGS 16949 | ||
| Database number: | 93176 | ||
| Authorizer: | A. Hendy | Enterer: | A. Hendy |
| Modifier: | A. Hendy | Research group: | marine invertebrate |
| Created: | 2010-01-06 11:14:52 | Last modified: | 2013-11-25 13:37:23 |
| Access level: | the public | Released: | 2010-01-06 11:14:52 |
| Creative Commons license: | CC BY | ||
Reference information
Primary reference:
| 11306. | W. P. Woodring. 1957. Geology and Paleontology of Canal Zone and adjoining parts of Panama. United States Geological Survey Professional Paper 306(A) [A. Miller/A. Hendy/A. Hendy] |
Secondary references:
| 30037 | A. G. Beu. 2010. Neogene Tonnoidean Gastropods of Tropical and South America; contributions to the Dominican Republic and Panama Paleontology Projects and Uplift of the Central American Isthmus. Bulletins of American Paleontology (377-378)1-550 [A. Hendy/A. Hendy/A. Hendy] | |
| 36423 | B. Landau and C. Marques da Silva. 2010. Early Pliocene gastropods of Cubagua, Venezuela: Taxonomy, palaeobiogeography and ecostratigraphy. Palaeontos 19:1-221 [A. Hendy/A. Hendy] | |
| 43491 | B. Landau, R. E. Petit, and C. M. Silva. 2012. New Cancellariidae (Mollusca, Gastropoda) from the Miocene Gatun Formation of Panama, with eleven new species. Journal of Paleontology 86(6):907-930 [A. Hendy/A. Hendy] | |
| 48647 | J. A. Todd and K. G. Johnson. 2013. Dissecting a marine snail species radiation (Conoidea: Turridae: Polystira) over 12 million years in the southwestern Caribbean. Bulletin of Marine Science 89(4) [A. Hendy/A. Hendy] | |
| 11372 | W. P. Woodring. 1959. Geology and paleontology of Canal Zone and adjoining parts of Panama: Description of Tertiary mollusks (gastropods: Vermetidae to Thaididae). United States Geological Survey Professional Paper 306(B):147-239 [A. Miller/A. Hendy/A. Hendy] | |
| 13232 | W. P. Woodring. 1964. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Columbellidae to Volutidae). United States Geological Survey Professional Paper 306(C):241-297 [A. Miller/A. Hendy/A. Hendy] | |
| 13236 | W. P. Woodring. 1970. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). United States Geological Survey Professional Paper 306(D):299-452 [A. Miller/A. Hendy/A. Hendy] | |
| 13235 | W. P. Woodring. 1973. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae). United States Geological Survey Professional Paper 306(E):453-539 [A. Miller/A. Hendy/A. Hendy] | |
| 13359 | W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy] |