PBDB Collection

Vishnevaya Balka creek, Karagan, Stavropol (PIN collection 254): Middle Miocene, Russian Federation
collected by B.F. Kaspiev 1938

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Insecta - Odonata - Sieblosiidae

	Stenolestes fasciata n. sp. Nel et al. 2005	Nel et al. 2005	1 specimen
	PIN 254/3192 (1 measurement)

	Paraoligolestes stavropolensis n. sp. Nel et al. 2005	Nel et al. 2005	2 specimens
	PIN 254/2171, 2202 (1 measurement)

	Miostenolestes zherikhini n. gen., n. sp. Nel et al. 2005	Nel et al. 2005	1 specimen
	PIN 254/2169 (2 measurements)
Insecta - Odonata - Libellulidae

	Sympetrum sp. Newman 1833	Nel et al. 2005	1 specimen
	PIN 254/2141 (1 measurement)

	Pantalinae indet.	Nel et al. 2005	2 specimens
	PIN 254/2163, 2198 (2 measurements)
Insecta - Odonata - Aeshnidae

	Aeshna stavropolensis n. sp. Nel et al. 2005	Nel et al. 2005	1 specimen
	PIN 254/2148 (1 measurement)
Insecta - Hemiptera - Aphididae

	Eutrichosiphum europaeum Wegierek and Peñalver 2002	Wegierek and Peñalver 2002	1 specimen
	PIN 254/2940 (1 measurement)
Insecta - Hemiptera - Derbidae

	Mysidioides migdisovae n. sp. Emeljanov 2002	Emeljanov 2002	1 specimen
	PIN 254/239 (1 measurement)
Insecta - Orthoptera - Oecanthidae

	Paroecanthus caucasicus n. gen., n. sp. Sharov 1968	Sharov 1968	1 specimen
	PIN 254/2577 (1 measurement)
Insecta - Lepidoptera

	Xyleutites miocenicus n. gen., n. sp. Kozhanchikov 1957	Kozhanchikov 1957	1 specimen
	recombined as Noctuites miocenicus
	PIN 254/182
Insecta - Lepidoptera - Nymphalidae

	Vanessa karaganica n. sp. Nekrutenko 1965		1 specimen
	recombined as Aglais karaganica
	PIN 254/2936a

	Pyrameis fossilis n. sp. Nekrutenko 1965		1 specimen
	recombined as Vanessa fossilis
	PIN 254/2753
Insecta - Trichoptera - Limnephilidae

	Limnephilus kaspievi (Martynova 1939)	Sukatsheva et al. 2018	24 specimens
	PIN 254/256, 651, 813, 958, 1010, 1212, 1262, 1264, 1268, 1269, 1271, 1272, 1273, 1275, 1276, 1277, 1278, 1281, 1313, 1359, 1955, 1351, 3041, 3186; presumably includes specimens from Temnolesskaya
Insecta - Coleoptera - Nitidulidae

	Lasiodactylus caucasicus n. sp. Kirejtshuk 1990	Kirejtshuk and Ponomarenko 1990	1 specimen
	recombined as Phenolia (Lasiodites) caucasicus
	PIN 254/1794 (2 measurements)
Insecta - Coleoptera - Dytiscidae

	? Graphoderus mirabilis n. sp. Říha 1974	Říha 1974	1 specimen
	PIN 254/1607 (1 measurement)

	Methles caucasicus n. sp. Říha 1974	Říha 1974	1 specimen
	PIN 254/1676 (1 measurement)

	Hydroporus lapideus n. sp. Říha 1974	Říha 1974	1 specimen
	PIN 254/1715 (2 measurements)

	Colymbetes sp. Clairville 1806	Říha 1974	1 specimen
	PIN 254/1758 (1 measurement)

	Colymbetes miocaenicus n. sp. Říha 1974	Říha 1974	1 specimen
	PIN 254/2673 (1 measurement)

	Copelatus spp.	Říha 1974	2 specimens
	PIN 254/1654, 1711 (2 measurements)

	Copelatus stavropolitanus n. sp. Říha 1974	Říha 1974	1 specimen
	PIN 254/1743 (2 measurements)

	Copelatus ponomarenkoi n. sp. Říha 1974	Říha 1974	2 specimens
	PIN 254/1734, 1733 (4 measurements)

	Copelatus fossilis n. sp. Říha 1974	Říha 1974	1 specimen
	PIN 254/1623 (1 measurement)

	Cybister fractus n. sp. Říha 1974	Říha 1974	1 specimen
	PIN 254/1626 (1 measurement)
Insecta - Diptera - Limoniidae

	Toxorhina (Ceratocheilus) caucasiensis n. sp. Krzemiński and Freiwald 1991	Krzemiński and Freiwald 1991	2 specimens
	PIN 254/671, 275 (1 measurement)

	Helius stavropolensis n. sp. Krzemiński 2002	Krzemiński 2002	5 specimens
	PIN 254/231, 278, 764, 288, 650 (1 measurement)

	Helius miocenicus n. sp. Krzemiński 2002	Krzemiński 2002	15 specimens
	PIN 254/833, 170a, 170b, 135, 322, 337, 533, 590, 598, 612, 821, 1079, 1442, 1580, 2301 (1 measurement)

	Helius verticillis n. sp. Krzemiński 2002	Krzemiński 2002	3 specimens
	PIN 254/2507, 816, 932 (1 measurement)
Insecta - Diptera - Trichoceridae

	Paracladura caucasiana n. sp. Krzemińska et al. 2009	Krzemińska et al. 2009	1 specimen
	PIN 254/697

	Trichocera sp. Meigen 1803	Krzemińska et al. 2009	1 specimen
	PIN 254/264
Insecta - Diptera - Tephritidae

	? Pseudacidia clotho n. sp. Korneyev 1982	Korneyev 1982	1 specimen
	recombined as Myoleja clotho
	PIN 254/724 (PIN 252/724 is error?) (1 measurement)
see common names

Geography

Country:	Russian Federation
Coordinates:	45.1° North, 42.3° East (view map)
Paleocoordinates:	45.5° North, 41.3° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	outcrop

Time

Period:	Neogene	Epoch:	Miocene
10 m.y. bin:	Cenozoic 5
Key time interval:	Middle Miocene
Age range of interval:	15.98000 - 11.63000 m.y. ago

Stratigraphy

Formation:

Karagan

Lithology and environment

Primary lithology:	lithified carbonaceous "shale"
Environment:	lacustrine indet.	Tectonic setting:	foreland basin

Taphonomy

Modes of preservation:	mold/impression
Size of fossils:	macrofossils,mesofossils

Collection methods and comments

Reason for describing collection:	taxonomic analysis
Museum repositories:	PIN
Collectors:	B.F. Kaspiev	Collection dates:	1938

Metadata

Database number:	105571
Authorizer:	M. Clapham	Enterer:	J. Karr, M. Clapham, L. Edwards
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	2011-02-22 10:36:54	Last modified:	2015-12-15 19:10:04
Access level:	the public	Released:	2011-02-22 10:36:54
Creative Commons license:	CC BY

Reference information

Primary reference:

39211.

Y. P. Nekrutenko. 1965. Tertiary nymphalid butterflies and some phylogenetic aspects of systematic lepidopterology. Journal of Research on the Lepidoptera 4(3):149-158 [M. Clapham/M. Clapham]

Secondary references:

41286		G. M. Dlussky. 1981. Miocenovye muravwi (Hymenoptera, Formicidae) CCCP. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 183:64-83 [M. Clapham/J. Karr/M. Clapham]
49501		A. F. Emeljanov. 2002. A record of Derbidae (Insecta: Homoptera: Fulgoroidea) in the Miocene of the northern Caucasus. Paleontological Journal 36:277-278 [M. Clapham/M. Clapham]
40325		A. G. Kirejtshuk and A. G. Ponomarenko. 1990. Fossil Beetles of the Peltidae and Nitidulidae Families (Coleoptera). Paleontological Journal 24(2):79-90 [M. Clapham/L. Edwards]
50549		V. A. Korneyev. 1982. A new fruit fly from the Miocene of the northern Caucasus. Paleontological Journal 16(4):95-97 [M. Clapham/M. Clapham]
52756		I. V. Kozhanchikov. 1957. Novyy predstavitel' sem. Cossidae iz Miotsenovykh otlozheniy Kavkaza (Lepidoptera, Insecta). Doklady Akademii Nauk SSSR 113:675-677 [M. Clapham/M. Clapham]
51817		E. Krzemińska, W. Krzemiński, and C. Dahl. 2009. Monograph of Fossil Trichoceridae (Diptera): Over 180 Million Years of Evolution 1-171 [M. Clapham/M. Clapham]
48705		W. Krzemiński. 2002. Three new species of the genus Helius Lepeletier & Serville (Diptera, Limoniidae) from the Middle Miocene of Stavropol (northern Caucasus, Russia). Acta Zoologica Cracoviensia 45:317-320 [M. Clapham/M. Clapham/M. Clapham]
44739		W. Krzemiński and A. Freiwald. 1991. Toxorhina (Ceratocheilus) caucasiensis, a new species from the Middle Miocene of Stavropol (northern Caucasus, USSR) (Diptera, Limoniidae). Paläontologische Zeitschrift 65:153-155 [M. Clapham/M. Clapham/M. Clapham]
35266		A. Nel, J. F. Petrulevičius, G. Gentilini and X. Martínez-Delclòs. 2005. Phylogenetic analysis of the Cenozoic family Sieblosiidae (Insecta: Odonata), with description of new taxa from Russia, Italy and France. Geobios 38:219-233 [M. Clapham/J. Karr/M. Clapham]
41734		A. Nel, J. F. Petrulevičius, and E. A. Jarzembowski. 2005. New fossil Odonata from the European Cenozoic (Insecta: Odonata: Thaumatoneuridae, Aeshnidae, Idionychidae, Libellulidae). Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 235(3):343-380 [M. Clapham/L. Edwards/M. Clapham]
48797		P. Říha. 1974. Neue fossile Schwimmkäfer aus dem Tertiär Europas und Westsibiriens (Coleoptera: Dytiscidae). Acta Entomologica Bohemoslovaca 71:398-413 [M. Clapham/M. Clapham/M. Clapham]
31339		A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [K. Beard/K. Beard/J. Karr]
67952		I. D. Sukatsheva, V. D. Ivanov, S. I. Melnitsky and D. S. Aristov. 2018. Caddisflies (Insecta: Trichoptera) from the Middle Miocene of the Stavropol Region, Russia. Paleontological Journal 52:1593-1609 [M. Clapham/M. Clapham]
39219		P. Wegierek and E. Peñalver. 2002. Fossil representatives of the family Greenideidae (Hemiptera, Aphidoidea) from the Miocene of Europe. Geobios 35:745-757 [M. Clapham/M. Clapham]