PBDB Collection

Oeningen (Carlsruhe collection): Sarmatian, Germany

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Insecta - Hymenoptera - Formicidae

	Formica obesa oeningensis		3 specimens

	Formica pinguis oeningensis		1 specimen
	(1 measurement)

	Ponera fuliginosa oeningensis		2 specimens
	SMNK Inv. 1235, 1236
Insecta - Odonata

	"Aeschna" eudore n. sp. Heer 1849		1 specimen
	larva
Insecta - Odonata - Corduliidae

	Cordulia sp. Leach 1815	Pongrácz 1931	1 specimen
	SMNK Inv. 1219 (Libellula doris by Heer)

	? Cordulia sp. Leach 1815	Pongrácz 1931	2 specimens
	SMNH Inv. 1257, 1263 (Libellula thoe by Heer)
Insecta - Odonata - Aeshnidae

	Aeschna polydore n. sp. Heer 1849		1 specimen
	SMNK Inv. 1265 (2 measurements)
Insecta - Odonata

	"Libellula" thetis Heer 1849		2 specimens
	larvae
Insecta - Hemiptera - Rhyparochromidae

	Pachymerus oblongus n. sp. Heer 1853	Heer 1853	1 specimen
Insecta - Hemiptera - Lygaeidae

	Lygaeites obsoletus n. sp. Heer 1853	Heer 1853	1 specimen
Insecta - Hemiptera - Scutelleridae

	Pachycoris escheri Heer 1853	Heer 1853	1 specimen

	Pachycoris germari Heer 1853	Heer 1853	1 specimen
Insecta - Hemiptera - Pentatomidae

	Eurydema brevicollis n. sp. Heer 1853	Heer 1853	1 specimen
	lower quarry

	Eurydema impudica Heer 1853	Heer 1853	1 specimen

	Eusarcoris pinguis n. sp.	Heer 1853	1 specimen
	lower quarry

	Halys bruckmanni Heer 1853	Heer 1853	1 specimen
	lower quarry

	Pentatoma longiceps n. sp. Heer 1853	Heer 1853	1 specimen
	lower quarry
Insecta - Hemiptera - Reduviidae

	Evagoras impressus n. sp. Heer 1853	Heer 1853	1 specimen
	lower quarry
Insecta - Hemiptera - Belostomatidae

	Diplonychus rotundatus n. sp. Heer 1853	Heer 1853	1 specimen
Insecta - Hemiptera - Membracidae

	Membracites cristatus n. gen., n. sp. Heer 1853	Heer 1853	1 specimen
Insecta - Hemiptera

	Pseudophana amatoria n. sp. Heer 1853	Heer 1853	1 specimen
Insecta - Hemiptera - Cercopidae

	Cercopis oeningensis n. sp. Heer 1853	Heer 1853	1 specimen
	lower quarry
Insecta - Hemiptera - Aphrophoridae

	Aphrophora spumarioides Heer 1853	Heer 1853	1 specimen
Insecta - Coleoptera - Buprestidae

	Ancylochira heydenii n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Ancylochira deleta n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Ancylochira gracilis n. sp. Heer 1847	Heer 1847	1 specimen
	Karlsruhe museum 279/285 (2 measurements)

	Buprestites exstincta n. sp. Heer 1847	Heer 1847	1 specimen
	(1 measurement)

	Capnodis antiqua Heer 1847	Heer 1847	2 specimens
Insecta - Coleoptera - Cantharidae

	Telephorus fragilis n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Telephorus germari n. sp. Heer 1847	Heer 1847	2 specimens
	(1 measurement)
Insecta - Coleoptera - Elateridae

	Diacanthus sutor n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Cardiophorus braunii n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)
Insecta - Coleoptera - Cerambycidae

	Mesosa jasonis n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)
Insecta - Coleoptera - Chrysomelidae

	Bruchus striolatus n. sp. Heer 1847	Heer 1847	1 specimen
	(1 measurement)

	Gonioctena clymene n. sp. Heer 1847	Heer 1847	2 specimens
Insecta - Coleoptera - Curculionidae

	Brachycerus germanus n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Cleonus larinoides n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Cleonus deucalionis n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Cleonus pyrrhae n. sp. Heer 1847	Heer 1847	1 specimen
	(1 measurement)
Insecta - Coleoptera - Anthribidae

	Anthribites moussonii n. sp. Heer 1847	Heer 1847	1 specimen
	(1 measurement)
Insecta - Coleoptera - Trogossitidae

	Trogossita koellikeri n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)
Insecta - Coleoptera - Coccinellidae

	Coccinella hesione n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)
Insecta - Coleoptera - Tenebrionidae

	Helops meissneri n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)
Insecta - Coleoptera - Meloidae

	Lytta aesculapii Heer 1847	Heer 1847	1 specimen
Insecta - Coleoptera

	Malachius vertummni Heer 1847	Heer 1847	2 specimens
Insecta - Coleoptera - Dermestidae

	Dermestes pauper n. sp. Heer 1847	Heer 1847	1 specimen
	(1 measurement)
Insecta - Coleoptera - Hydrophilidae

	Hydrophilus braunii n. sp. Heer 1847	Heer 1847	1 specimen

	Hydrophilus knorrii n. sp. Heer 1847	Heer 1847	1 specimen
	(1 measurement)

	Hydrophilus vexatorius Heer 1847	Heer 1847	1 specimen
	(1 measurement)
Insecta - Coleoptera - Staphylinidae

	Scaphidium deletum n. sp. Heer 1847	Heer 1847	1 specimen
	(1 measurement)
Insecta - Coleoptera - Scarabaeidae

	Rhizotrogus longimanus n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Aphodius meyeri n. sp. Heer 1847	Heer 1847	1 specimen
	(1 measurement)
Insecta - Coleoptera - Carabidae

	Argutor antiquus n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)

	Harpalus sinis n. sp. Heer 1847	Heer 1847	1 specimen
	(2 measurements)
Insecta - Diptera - Asilidae

	Asilus antiquus Heer 1849		1 specimen
	SMNK Inv. 1195 (1 measurement)
Insecta - Diptera - Bibionidae

	Protomyia amoena n. sp. Heer 1849		2 specimens
	SMNK Oe. 170 Inv. 1247; Oe. 159 in Skartveit & Pika 2014

	Bibio elongatus n. sp. Heer 1849		1 specimen
	SMNK Oe. 154 Inv. 1189 (4 measurements)

	Protomyia jucunda n. sp. Heer 1849		2 specimens
	SMNK Oe. 168 Inv. 1279 (2 measurements)

	Bibio fusiformis Heer 1849		6 specimens
	SMNK Oe. 142-145, 148, 161(x2), 194 (2 measurements)

	Bibio obsoletus n. sp. Heer 1849		1 specimen
	SMNK Oe. 161

	Bibio angustatus n. sp. Heer 1849		4 specimens
	SMNK Oe. 179 Inv. 1274, Oe. 151-153

	Bibio brevis Heer 1849		1 specimen
	SMNK Oe. 160 Inv. 1193

	Bibio oblongus Heer 1849	Skartveit and Pika 2014	1 specimen
	SMNK Oe. 156

	Bibio moestus n. sp. Heer 1849		4 specimens
	SMNK Oe. 163, 157
Insecta - Diptera - Chironomidae

	Chironomus obsoletus n. sp. Heer 1849		1 specimen
	(1 measurement)
Insecta - Hymenoptera - Ichneumonidae

	Cryptus antiquus n. sp. Heer 1849		1 specimen

	Anomalon protogaeum n. sp. Heer 1849		1 specimen
Insecta - Hymenoptera - Apidae

	Anthophorites tonsa n. sp. Heer 1849		1 specimen
	(1 measurement)

	Anthophorites titania n. sp. Heer 1849		2 specimens
	Holotype Inv.1273 (Oe.92), paratype now at ETH Zurich Pl. I, 686 (2 measurements)

	Bombus abavus Heer 1867	Zeuner and Manning 1976	1 specimen
	Inv. 1268 (Oe.90) (1 measurement)

	Xylocopa senilis n. sp. Heer 1849		7 specimens
	type and 1 poorly preserved in Heer; Inv.1267, 270 (Pl 5 fig 6 in Pongracz, 1931), 1175 (Oe.89, no. 145), 1175 (Oe.89, no. 146), 1228 (Oe.93) in Zeuner & Manning, 1976 (7 measurements)
Insecta - Hymenoptera - Megachilidae

	Osmia antiqua n. sp. Heer 1849		1 specimen
	(2 measurements)

	Lithurge adamitica (Heer 1864)	Zeuner and Manning 1976	1 specimen
	Inv.1216 (Oe.95) (apparently different from Heer specimens)
Insecta - Hymenoptera - Formicidae

	Formica lignitum Germar 1837		5 specimens
	Includes SMNK Inv. 1182, 1230 (4 measurements)

	Formica immersa Heer 1849	Pongrácz 1931	1 specimen
	SMNK Inv. 1282 (2 measurements)

	Formica macrophthalma Heer 1849		1 specimen
	(1 measurement)

	Formica seuberti Heer 1849	Pongrácz 1931	1 specimen
	SMNK Inv. 1193 (Oe 113) (1 measurement)

	Formica orbata Heer 1849		1 specimen

	Formica longicollis n. sp. Heer 1849		1 specimen
	(2 measurements)

	Formica procera Heer 1849		1 specimen

	Formica pulchella Heer 1849		2 specimens

	Cephites oeningensis n. sp. Heer 1849		1 specimen
	(2 measurements)

	Myrmica macrocephala Heer 1849	Pongrácz 1931	1 specimen
	SMNK Inv. 1215 (2 measurements)
Insecta - Hymenoptera - Pompilidae

	Pompilus induratus n. sp. Heer 1849		1 specimen
	(1 measurement)
Insecta - Orthoptera - Tettigoniidae

	Phaneroptera vetusta Heer 1849		2 specimens
see common names

Geography

Country:	Germany	State/province:	Baden-Württemberg
Coordinates:	47.7° North, 8.9° East (view map)
Paleocoordinates:	45.5° North, 10.6° East (Wright 2013)
Basis of coordinate:	based on nearby landmark
Geographic resolution:	outcrop

Time

Period:	Neogene	Epoch:	Miocene
Stage:	Serravallian	10 m.y. bin:	Cenozoic 5
Key time interval:	Sarmatian	Mammal zone:	MN 7
Age range of interval:	12.8 - 11.63 m.y. ago

Stratigraphy

Formation:	Upper Freshwater-Molasse	Member:	Upper Oehningen beds
Stratigraphic resolution:	group of beds

Lithology and environment

Primary lithology:	lithified "limestone"
Environment:	crater lake	Tectonic setting:	foreland basin
Geology comments: "meromictic maar lake of approximately 1000 m in diameter and perhaps up to 100 m in depth" (Lutz, 1997). "Deposited in the Northern Alpine Foreland basin" (Uhl et al., 2006).

Taphonomy

Modes of preservation:	mold/impression,adpression
Size of fossils:	macrofossils,mesofossils

Collection methods and comments

Collection methods:	survey of museum collection
Reason for describing collection:	taxonomic analysis

Metadata

Database number:	115305
Authorizer:	M. Clapham	Enterer:	M. Clapham
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	2011-09-01 02:15:05	Last modified:	2025-02-22 15:12:02
Access level:	the public	Released:	2011-09-01 02:15:05
Creative Commons license:	CC0

Reference information

Primary reference:

37344.

O. Heer. 1849. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Zweiter Theil: Henschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen 1-264 [M. Clapham/M. Clapham]

Secondary references:

38897		O. Heer. 1847. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Erster Theil: Käfer 1-229 [M. Clapham/M. Clapham]
39159		O. Heer. 1853. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Dritte Theil: Rhynchoten 1-138 [M. Clapham/M. Clapham/M. Uhen]
56453		A. Pongrácz. 1931. Bemerkungen über die Insektenfauna von Oeningen nebst Revision der Heer'schen Typen. Verhandlungen des Naturhistorisch-Medizinischen Vereins zu Heidelberg 17(2):104-125 [M. Clapham/M. Clapham]
53927		J. Skartveit and M. Pika. 2014. Revision of Bibionidae (Diptera) named by Oswald Heer from the Miocene of Öhningen, southern Germany. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 87:103-134 [M. Clapham/M. Clapham]
5180	ETE 892	F. E. Zeuner and F. J. Manning. 1976. A monograph on fossil bees (Hymenoptera: Apoidea). Bulletin of the British Museum (Natural History), Geology 27(3):149-268 [C. Labandeira/C. Labandeira/M. Clapham]