Mammal Quarry No. 2 (UCMP V-5815): Lancian, Wyoming
collected by J. B. Hatcher, W. A. Clemens 1889, 1958

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gastropoda - Hydrobiidae
Reesidella sp. Yen 1952
Estes 1964
Bivalvia - Unionida - Unionidae
Unionidae indet. Rafinesque 1820
Estes 1964
Mammalia - Pediomyidae
Pediomys florencae Clemens 1966
Clemens 1966
recombined as Protolambda florencae
Pediomys hatcheri (Osborn 1898)
Clemens 1966
original and current combination Protolambda hatcheri
Pediomys krejcii Clemens 1966
Clemens 1966
recombined as Leptalestes krejcii
Pediomys cooki Clemens 1966
Clemens 1966
recombined as Leptalestes cooki
Pediomys elegans Marsh 1889
Clemens 1966
(3 measurements)
Mammalia
Nortedelphys intermedius Case et al. 2004
Case et al. 2004
synonym of Nortedelphys jasoni
Mammalia - Stagodontidae
Didelphodon vorax Marsh 1889
Clemens 1966
Mammalia - Alphadontidae
Alphadon marshi Simpson 1927
Clemens 1966
Alphadon rhaister n. sp. Clemens 1966
Clemens 1964
recombined as Turgidodon rhaister
Mammalia
Alphadon lulli Clemens 1966
Clemens 1964
recombined as Protalphadon lulli
Mammalia - Gypsonictopidae
Gypsonictops hypoconus Simpson 1927
Clemens 1973
Mammalia - Cimolestidae
Cimolestes incisus Marsh 1889
Clemens 1973
Mammalia - Multituberculata - Neoplagiaulacidae
Mesodma formosa (Marsh 1889)
Clemens 1964
    = Mesodma sp. Jepsen 1940
Alroy 2002
Mesodma thompsoni Clemens 1964
Clemens 1964
Mammalia - Multituberculata - Cimolodontidae
Cimolodon nitidus Marsh 1889
Clemens 1964
Mammalia - Multituberculata - Cimolomyidae
Dipriodon robustus n. sp. Marsh 1889
recombined as Meniscoessus robustus
Cimolomys gracilis Marsh 1889
Clemens 1964
present in both UCMP and Hatcher collections
Reptilia - Chamopsiidae
Leptochamops denticulatus (Gilmore 1928)
Estes 1964 1 specimen
Chamops segnis Marsh 1892
Estes 1964 4 specimens
Meniscognathus altmani Estes 1964
Estes 1964 1 specimen
Haptosphenus placodon Estes 1964
Estes 1964 1 specimen
Reptilia
Coniophis precedens Marsh 1892
Longrich et al. 2012 6 specimens
nomen dubium belonging to Serpentes
UCMP 53556‐53561 (vertebrae)
Reptilia - Xenosauridae
Exostinus lancensis Gilmore 1928
Estes 1964
Reptilia
Paraderma bogerti Estes 1964
Estes 1964
Reptilia - Anguidae
Peltosaurus piger Gilmore 1928
Estes 1964 2 specimens
recombined as Odaxosaurus piger
Reptilia
Parasaniwa wyomingensis Gilmore 1928
Estes 1964 2 specimens
Colpodontosaurus cracens Estes 1964
Estes 1964 1 specimen
Palaeosaniwa canadensis Gilmore 1928
Estes 1964 1 specimen
    = Varanoidea indet. Gray 1827
Gao and Fox 1996
Compsemys victa (Leidy 1856)
Estes 1964 1 specimen
Reptilia - Baenidae
Baenidae indet. Cope 1882
Database 2005 1 specimen
Including UCMP 173424, listed by Holroyd & Hutchison (2002)
Reptilia - Testudines
Cryptodira indet. Cope 1868
Database 2005 2 specimens
Kinosternidae indet. (Hay 1892)
Database 2005 1 specimen
spelled with current rank as Kinosternoidea
Including UCMP 173422, listed by Holroyd & Hutchison (2002)
Reptilia - Testudines - Chelydridae
? Chelydridae indet. Swainson 1839
Database 2005 1 specimen
Chelydridae indet. Swainson 1839
Database 2005 3 specimens
Including UCMP 173420, listed by Holroyd & Hutchison (2002)
Reptilia - Testudines - Trionychidae
Trionychidae indet. Gray 1825
Database 2005 1 specimen
Including UCMP 173026, listed by Holroyd & Hutchison (2002)
Reptilia - Testudines - Emydidae
Emydinae indet. Rafinesque 1815
Estes 1964
"near Pseudemys?"
Reptilia - Archosauria
Archosauria indet. Cope 1869
Database 2005 1 specimen
Reptilia
Leidyosuchus sternbergi (Gilmore 1910)
Estes 1964
recombined as Borealosuchus sternbergii
Brachychampsa montana Gilmore 1911
Estes 1964
Thescelosaurus neglectus Gilmore 1913
Estes 1964 2 specimens
    = Thescelosaurus sp. Gilmore 1913
Carpenter 1982
Paronychodon lacustris Cope 1876
Estes 1964 1 specimen
nomen dubium belonging to Paronychodon
cf. Paronychodon lacustris Cope 1876
Sankey 2008 1 specimen
nomen dubium belonging to Paronychodon
UCMP 73076
Reptilia - Troodontidae
Troodontidae indet. Gilmore 1924
Database 2005 2 specimens
Troodon sp. Leidy 1856
Database 2005 1 specimen
Pectinodon bakkeri Carpenter 1982
Carpenter 1982 2 specimens
Reptilia - Dromaeosauridae
Richardoestesia sp. Currie et al. 1990
Database 2005 1 specimen
Saurornitholestinae indet. Longrich and Currie 2009
Larson and Currie 2013 1 specimen
UCMP 187054
Reptilia
Aves indet. Linnaeus 1758
Database 2005 1 specimen
cf. Dryptosaurus sp. Marsh 1877
Estes 1964
Champsosaurus sp. Cope 1876
Estes 1964 1 specimen
Amphibia - Temnospondyli - Pelobatidae
? Pelobatidae indet. Lataste 1879
Estes 1964
    = Eopelobates sp. Parker 1929
Estes and Sanchíz 1982
Amphibia - Temnospondyli - Batrachosauroididae
Opisthotriton kayi Auffenberg 1961
Estes 1964 2 specimens
Prodesmodon copei Estes 1964
Estes 1964 2 specimens
Cuttysarkus mcnallyi Estes 1964
Estes 1964 5 specimens
synonym of Prodesmodon copei
Amphibia - Temnospondyli - Urodela
Scapherpeton tectum Cope 1876
Estes 1964 4 specimens
Habrosaurus dilatus Gilmore 1928
Estes 1964 4 specimens
Actinopteri - Amiiformes - Amiidae
Kindleia fragosa Jordan 1927
Estes 1964 30 elements
recombined as Cyclurus fragosus
Melvius thomasi Bryant 1987
Database 2005 1 element
cf. Protamia sp. (Leidy 1873)
Estes 1964 1 element
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteus occidentalis (Leidy 1856)
Estes 1964 16 elements
original and current combination Lepidotus occidentalis
Actinopteri - Albuliformes - Albulidae
Albulidae indet. Bleeker 1859
Estes 1964 3 elements
Actinopteri - Elopiformes - Elopidae
Elopidae indet. Bonaparte 1846
Estes 1964 2 elements
Actinopteri - Esociformes - Esocidae
Platacodon nanus Marsh 1889
Estes 1964 9 elements
    = Estesesox foxi Wilson et al. 1992
Wilson et al. 1992
Actinopteri - Aspidorhynchiformes - Aspidorhynchidae
Belonostomus longirostris (Lambe 1902)
Estes 1964 2 elements
Acipenseriformes - Acipenseridae
Acipenser eruciferus (Cope 1876)
Estes 1964 1 element
Chondrichthyes - Hybodontiformes - Lonchidiidae
Lonchidion selachos Estes 1964
Estes 1964 1 element
see common names

Geography
Country:United States State/province:Wyoming County:Niobrara
Coordinates: 43.1° North, 104.5° West (view map)
Paleocoordinates:49.6° North, 79.4° West
Basis of coordinate:estimated from map
Geographic resolution:small collection
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
*Epoch:Senonian
Key time interval:Lancian
Age range of interval:72.10000 - 66.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Lance
Stratigraphic resolution:bed
Lithology and environment
Primary lithology:"cross stratification" sandstone
Includes fossils?Y
Lithology description: "cross-stratified sandstone"
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:concentrated
Size of fossils:mesofossils
Preservation of anatomical detail:good
Disassociated major elements:many
Disassociated minor elements:many
Collection methods and comments
Collection methods:bulk,sieve,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:UCMP,YPM
Collectors:J. B. Hatcher, W. A. Clemens Collection dates:1889, 1958
Metadata
Also known as:Lance Quarry 2, Hatcher No. 2
Database number:14566
Authorizer:J. Alroy, R. Benson, M. Carrano, P. Holroyd Enterer:J. Alroy, P. Holroyd, M. Carrano, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:1994-07-26 00:00:00 Last modified:2022-10-18 11:14:33
Access level:the public Released:1994-07-26 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:
14177.ETE R. S. Lull. 1915. The mammals and horned dinosaurs of the Lance Formation of Niobrara County, Wyoming. The American Journal of Science, series 4 40(238):319-348 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
12202ETE K. Carpenter. 1982. Baby dinosaurs from the Late Cretaceous Lance and Hell Creek formations and a description of a new species of theropod. Contributions to Geology, University of Wyoming 20(2):123-134 [M. Carrano/M. Carrano/M. Carrano]
31410 J. A. Case, F. J. Goin, and M. O. Woodburne. 2004. "South American" marsupials from the Late Cretaceous of North America and the origin of marsupial cohorts. Journal of Mammalian Evolution 11(3/4):223-255 [J. Alroy/J. Alroy/J. Alroy]
1239 W. A. Clemens. 1964. Fossil mammals of the type Lance Formation, Wyoming. Part I. Introduction and Marsupialia. University of California Publications in Geological Sciences 48:1-105 [J. Alroy/J. Alroy/J. Alroy]
1241 W. A. Clemens. 1966. Fossil mammals from the type Lance Formation, Wyoming. Part II. Marsupialia. University of California Publications in Geological Sciences 62:1-122 [J. Alroy/J. Alroy/J. Alroy]
1242 W. A. Clemens. 1973. Fossil mammals of the type Lance Formation, Wyoming. Part III. Eutheria and summary. University of California Publications in Geological Sciences 94:1-102 [J. Alroy/J. Alroy/J. Alroy]
13103 U.C.M.P. Database. 2005. UCMP collections database. University of California Museum of Paleontology [M. Carrano/M. Carrano/M. Carrano]
803 R. Estes. 1964. Fossil vertebrates from the Late Cretaceous Lance Formation, eastern Wyoming. University of California Publications in Geological Sciences 49:1-187 [J. Alroy/M. Sommers/M. Carrano]
19691ETE R. Estes and B. Sanchíz. 1982. New discoglossid and palaeobatrachid frogs from the Late Cretaceous of Wyoming and Montana, and a review of other frogs from the Lance and Hell Creek Formations. Journal of Vertebrate Paleontology 2(1):9-20 [M. Carrano/M. Carrano]
37933 K. -Q. Gao and R. C. Fox. 1996. Taxonomy and evolution of Late Cretaceous lizards (Reptilia: Squamata) from western Canada. Bulletin of the Carnegie Museum of natural History 33:1-107 [R. Benson/R. Benson]
42536 P. A. Holroyd and J. H. Hutchison. 2002. Patterns of geographic variation in latest Cretaceous vertebrates: evidence from the turtle component. Geological Society of America Special Paper 361:177-190 [R. Benson/R. Benson]
48570 D. W. Larson and P. J. Currie. 2013. Multivariate analyses of small theropod dinosaur teeth and implications for paleoecological turnover through time. PLoS ONE 8(1):e54329:1-14 [M. Carrano/M. Carrano/M. Carrano]
42713 N. R. Longrich, B.-A. S. Bhullar, and J. A. Gauthier. 2012. A transitional snake from the Late Cretaceous period of North America. Nature 488:205-208 [P. Mannion/P. Mannion]
34479ETE J. T. Sankey. 2008. Diversity of latest Cretaceous (late Maastrichtian) small theropods and birds: teeth from the Lance and Hell Creek formations, USA. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography. Indiana University Press, Bloomington 117-134 [M. Carrano/M. Carrano]
45321 M. V. H. Wilson, D. B. Brinkman, and A. G. Neuman. 1992. Cretaceous Esocoidei (Teleostei): early radiation of the pikes in North American fresh waters. Journal of Paleontology 66(5):839-846 [M. Bell/M. Bell]