PBDB Collection

Princeton Quarry: Tiffanian, Wyoming
collected by G. L. Jepsen

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Mammalia - Didelphimorphia - Didelphidae

	Peratherium sp. Aymard 1850
	= Herpetotheriinae indet. Trouessart 1879	Alroy 2002
	= Didelphidae indet. Gray 1821	Secord 2008
Mammalia

	Peradectes elegans Matthew and Granger 1921
Mammalia - Leptictidae

	Prodiacodon cf. concordiarcensis Simpson 1935
	= Prodiacodon paucus n. sp. Secord 2008	Secord 2008

	? Palaeictops sp. Matthew 1899
Mammalia - Proteutheria - Palaeoryctidae

	Palaeoryctes sp. Matthew 1913
	= Palaeoryctes cf. punctatus Van Valen 1966	Gingerich 1982
Mammalia - Apatemyidae

	Unuchinia sp. Simpson 1937
Mammalia - Barylambdidae

	Barylambdidae indet. Patterson 1939	Secord 2008
Mammalia - Macroscelidea

	Adunator abditus n. sp. Secord 2008	Secord 2008

	Diacodon minutus n. sp. Jepsen 1930
	= Diacocherus minutus Jepsen 1930	Alroy 2002
	recombined as Adunator minutus
Mammalia - Macroscelidea - Apheliscidae

	Phenacodaptes sabulosus n. sp. Jepsen 1930
	(1 measurement)

	Phenacodaptes sabulosus Jepsen 1930	Secord 2008
Mammalia - Acreodi - Mesonychidae

	Dissacus cf. navajovius (Cope 1881)
	(1 measurement)
	= Dissacus argenteus n. sp. O'Leary and Rose 1995	O'Leary and Rose 1995
	DUPLICATE REF 2677 or 2678
Mammalia - Phenacodontidae

	Ectocion osbornianus (Cope 1882)
	= Ectocion mediotuber n. sp. Thewissen 1990	Thewissen 1990

	Phenacodus vortmani (Cope 1880)	Secord 2008

	Phenacodus sp. Cope 1873
	= Phenacodus grangeri Simpson 1935	Secord 2008
Mammalia - Procreodi - Arctocyonidae

	Princetonia yalensis Gingerich 1989	Gingerich 1989
	= Princetonia cf. yalensis Gingerich 1989	Secord 2008

	Claenodon sp. Scott 1892
	= Arctocyon sp. Blainville 1841	Alroy 2002
	= Arctocyon corrugatus Cope 1883	Kondrashov and Lucas 2004

	Thryptacodon sp. Matthew and Granger 1915
	= Thryptacodon pseudarctos Simpson 1928	Secord 2008
Mammalia - Viverravidae

	Protictis schaffi Gingerich and Winkler 1985	Gingerich and Winkler 1985
	= Viverravus politus Matthew and Granger 1915	Alroy 2002
	= Viverravus schaffi Gingerich and Winkler 1985	Secord 2008

	Protictis laytoni n. sp. Gingerich and Winkler 1985	Gingerich and Winkler 1985
	(1 measurement)
	= Viverravus laytoni Gingerich and Winkler 1985	Alroy 2002
	"Viverravine sp. A" of Rose

	Protictis dellensis (Dorr 1952)	Gingerich and Winkler 1985
	= Didymictis proteus Polly 1997	Alroy 2002
	"D. sp." said to be new
	= Didymictis dellensis Dorr 1952	Secord 2008
Mammalia - Carnivora - Miacidae

	Viverravinae indet. Matthew 1909
	"Viverravine sp. B" of Rose
Mammalia - Metacheiromyidae

	Propalaeanodon schaffi Rose 1979
Mammalia - Nyctitheriidae

	Leptacodon packi n. sp. Jepsen 1930

	Wyonycteris galensis n. sp. Secord 2008	Secord 2008
Mammalia - Apternodontidae

	Apternodontidae indet. Matthew 1910
	"Apternodus-like" A and B of Rose
Mammalia - Erinaceidae

	Litolestes ignotus n. sp. Jepsen 1930
Mammalia - Primates - Plesiadapidae

	Plesiadapis fodinatus n. sp. Jepsen 1930
	(6 measurements)

	Plesiadapis churchilli Gingerich 1975	Secord 2008
Mammalia - Primates - Carpolestidae

	Carpolestes dubius n. sp. Jepsen 1930
Mammalia - Primates - Paromomyidae

	Phenacolemur pagei n. sp. Jepsen 1930
	= Dillerlemur pagei Jepsen 1930	Alroy 2002
Mammalia - Primates - Micromomyidae

	Micromomys silvercouleei n. sp. Szalay 1973
Mammalia - Multituberculata - Neoplagiaulacidae

	Parectypodus laytoni n. sp. (Jepsen 1940)
	original and current combination Ectypodus laytoni

	Ectypodus powelli n. sp. Jepsen 1940

	Mimetodon churchilli n. sp. Jepsen 1940

	Neoplagiaulax hazeni n. sp. (Jepsen 1940)
Mammalia - Multituberculata

	Neoliotomus conventus Jepsen 1930
Mammalia - Multituberculata - Ptilodontidae

	Prochetodon cavus n. sp. Jepsen 1940
Mammalia - Multituberculata - Microcosmodontidae

	Pentacosmodon pronus n. sp. Jepsen 1940

	Microcosmodon conus n. sp. Jepsen 1930
Reptilia

	Haplodontosaurus excedens (Gilmore 1928)	Gilmore 1942
	nomen dubium belonging to Squamata
Reptilia - Oligodontosauridae

	Oligodontosaurus wyomingensis n. gen., n. sp. Gilmore 1942	Gilmore 1942	1 specimen
	YPM-PU 14246 (holotype), left mandible
Reptilia - Necrosauridae

	Provaranosaurus acutus n. gen., n. sp. Gilmore 1942	Gilmore 1942	3 specimens
	PU 14243 (holotype), left maxilla; 14561, anterior portion of left dentary; 17145, questionably referred fragment of left maxilla of large specimen
Reptilia

	Exostinus rugosus n. sp. Gilmore 1942	Gilmore 1942	4 specimens
	PU 14559, posterior half of right maxilla (holotype); 14560, right maxilla of smaller individual; 17144, left dentary fragment; 14640, disarticulated skull, jaws, and fragments of skeleton, including maxilla, prefrontals, frontal, right jugal, left pterygoid, dentaries, right prearticular- surangular; coronoids, right angular, centrum of axis, several cervical vertebrae, fragmentary left ilium and unidentified fragments.
	= Restes rugosus n. gen., n. sp. Gauthier 1982	Gauthier 1982
	Exostinus rugosus is an objective synonym of Restes rugosus (Gauthier 1982). PU 14559 (holotype), frontals. Additional specimens: "See Estes, 1975, for referred specimens in Princeton’s collection from the Ft. Union Formation"; UC loc. V-74158; UCMP 119434, posterior half of co-ossified frontals; UC loc. V-70246; UCMP 98229, anterior two-thirds of partially cosssified frontals; UCMP 100049, worn right frontal from the level of the post frontal incision; UCMP 100050, fragment from right anterolateral quadrant of frontal showing nasal and prefrontal incisions; UCMP 100051, fragment from right side of frontal near posterior end of frontals; UCMP 100053, fragment from left side of frontal in region of prefrontal incision; UCMP 100054, left frontal from anterior end of postfrontal incision to posterior end of prefrontal incision.
Reptilia - Anguidae

	Pancelosaurus piger (Gilmore 1928)	Estes 1975	10 specimens
	PU 13371, left maxilla; 14562, left maxilla; 14565, right maxilla, posterior part of parietal, one dermal scute; 14574, left maxilla, anterior end of right dentary; 16790, left dentary; 17147, left dentary; 17148, left dentary fragment and coronoid; 18578, left dentary fragment and osteoscutes; 14667, right pterygoid; 16761, parietal.
	= Odaxosaurus piger Gilmore 1928	Meszoely et al. 1978

	Peltosaurus jepseni n. sp. Gilmore 1942	Gilmore 1942	6 specimens
	synonym of Proxestops silberlingii
	Recombined into Proxestops jepseni by Gauthier (1982), synonymised into Proxestops silberlingii by Sullivan (1991). PU 14565 (originally holotype); UCMP 99851, isolated osteoderms; UCMP 100163, right maxilla missing all teeth, premaxillary, posterior and nasal processes; UCMP 100164, worn fragment from posterior end of right maxilla; UCMP 115128, posterior process of left maxilla; UCMP 115130, fragment from middle of right maxilla. Some specimens here removed and reclassified as Xestops savagei by ref. 60296 in coll. 15451.
Reptilia - Testudines - Kinosternoidea

	Ptychogaster sp. Pomel 1847	Estes 1975	1 specimen
	PU 17794, fragments of skull and shell.
	= Emydidae informal indet. C Rafinesque 1815	Holroyd et al. 2001
	= Cardichelyon rogerwoodi Hutchison 2013	Hutchison 2013
Reptilia

	Allognathosuchus sp. Mook 1921	Estes 1975	1 specimen
	PU 16988, anterior end of left dentary.
Amphibia - Temnospondyli

	Anura indet. (Fischer von Waldheim 1813)	Estes 1975	4 specimens
	PU 14662, distal end of right radioulna; 14663, proximal end of left radioulna; 14669 (part), phalanx; 13372, right maxilla.
Amphibia - Temnospondyli - Discoglossidae

	Discoglossidae indet. Günther 1859	Estes 1975	1 specimen
	PU 14670, distal end of right humerus. Undescribed genus and species.
Amphibia - Temnospondyli - Rhinophrynidae

	Eorhinophrynus sp. Hecht 1959	Estes 1975	3 specimens
	PU 14664, left humerus; 14665, distal end of left humerus; 14669 (part), sacral vertebra.
Amphibia - Temnospondyli - Batrachosauroididae

	Opisthotriton kayi Auffenberg 1961	Estes 1975	4 specimens
	PU 14575, articulated skeleton, including poorly-preserved skull, mandibles, atlas, at least 35 presacral vertebrae, many in natural articulation, fragments of hyobranchial skeleton, a humerus, and a single rib; PU 14643, partial skeleton of a somewhat smaller individual, including skull, mandibles, partial hyobranchial skeleton in natural articulation with skull, atlas and five associated vertebrae in natural articulation; 20584, fused articular-prearticular, otoccipitals, associated with material of Amphiuma jepseni noted below; 21781, two vertebral centra associated with a block of dislocated vertebrae, mostly young Amphiuma jepseni.
Amphibia - Temnospondyli - Urodela

	Scapherpeton tectum Cope 1876	Estes 1975	1 specimen
	PU 20583, three broken vertebrae, an atlas, an ilium, and a rib.

	Amphiuma jepseni n. sp. Estes 1969	Estes 1975	3 specimens
	PU 14666 (holotype), partial vertebral column with at least 14 associated but dislocated vertebrae in a single block of matrix, associated left quadrate and left dentary; 14668, partial skull and associated mandibles; 16788, group of dislocated vertebrae from sacral region of young individual mingled with vertebrae of Opisthotriton kayi
see common names

Geography

Country:	United States	State/province:	Wyoming	County:	Park
Coordinates:	44.9° North, 108.9° West (view map)
Paleocoordinates:	50.8° North, 88.4° West
Basis of coordinate:	stated in text
Geographic resolution:	small collection

Time

Period:	Paleogene	Epoch:	Paleocene
10 m.y. bin:	Cenozoic 1
*Period:	Tertiary
Key time interval:	Tiffanian
Age range of interval:	60.90000 - 56.20000 m.y. ago
* legacy (obsolete) database fields

Stratigraphy

Formation:	Fort Union
Local section:	Clark	Local bed:	680 m
Local order:	bottom to top
Stratigraphic resolution:	group of beds
Stratigraphy comments: low within chron C25r according to Butler et al. 1981 but in the middle of C25r according to Secord et al. 2006 "Polecat Bench Fm, Silver Coulee beds" according to Gilmore 1942

Lithology and environment

Primary lithology:	not reported
Environment:	terrestrial indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils
Articulated whole bodies:	none
Associated major elements:	some
Disassociated major elements:	many
Disassociated minor elements:	many

Collection methods and comments

Collection methods:	field collection,survey of museum collection
Collection size:	546 specimens
Reason for describing collection:	general faunal/floral analysis
Museum repositories:	UMMP
Collectors:	G. L. Jepsen
Collection method comments: Princeton University collection
Taxonomic list comments:replaces list of Jepsen and Woodburne 1969

Metadata

Also known as:	Big Sand Coulee; Locality No. 3; SC-187
Database number:	15049
Authorizer:	J. Alroy, E. Vlachos, R. Whatley, K. Beard, J. Marcot, P. Holroyd, P. Barrett	Enterer:	J. Alroy, J. Marcot, E. Vlachos, R. Whatley, K. Beard, T. Cleary, P. Holroyd
Modifier:	P. Mannion	Research group:	vertebrate
Created:	1993-02-18 00:00:00	Last modified:	2021-03-16 09:26:20
Access level:	the public	Released:	1993-02-18 00:00:00
Creative Commons license:	CC BY

Reference information

Primary reference:

2953.

K. D. Rose. 1981. The Clarkforkian Land-Mammal Age and Mammalian Faunal Composition Across the Paleocene-Eocene Boundary. University of Michigan Papers on Paleontology 26:1-197 [J. Alroy/J. Alroy/P. Mannion]

Secondary references:

6294		J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
709		R. Estes. 1975. Lower vertebrates from the Fort Union Formation, Late Paleocene, Big Horn Basin, Wyoming. Herpetologica 31(4):365-385 [J. Alroy/M. Sommers/J. Alroy]
14650		J. A. Gauthier. 1982. Fossil xenosaurid and anguid lizards from the early Eocene Wasatch Formation, southeast Wyoming, and a revision of the Anguidea. Contributions to Geology, University of Wyoming 21(1):7-54 [R. Whatley/R. Whatley/R. Whatley]
28626		C. W. Gilmore. 1942. Paleocene faunas of the Polecat Bench Formation, Park County, Wyoming Part II. Lizards. Proceedings of the American Philosophical Society 85(2):159-167 [J. Alroy/J. Alroy]
1647		P. D. Gingerich. 1982. Second Species of Wyolestes (Condylarthra, Mesonychia) from the Early Eocene of Western North America. Journal of Mammalogy 63(4) [J. Alroy/J. Alroy/J. Ju]
1657		P. D. Gingerich. 1989. New earliest Wasatchian mammalian fauna from the Eocene of northwestern Wyoming: composition and diversity in a rarely sampled high-floodplain assemblage. University of Michigan Papers on Paleontology 28:1-97 [J. Alroy/J. Alroy/J. Alroy]
1676		P. D. Gingerich and D. A. Winkler. 1985. Systematics of Paleocene Viverravidae (Mammalia, Carnivora) in the Bighorn Basin and Clark's Fork Basin, Wyoming. Contributions from the Museum of Paleontology, University of Michigan 27(4):87-128 [J. Alroy/J. Alroy/J. Marcot]
17201		P. A. Holroyd, J. H. Hutchison, and S. G. Strait. 2001. Turtle diversity and abundance through the lower Eocene Willwood Formation of the southern Bighorn Basin. University of Michigan Papers on Paleontology 33:97-108 [J. Alroy/J. Alroy/J. Alroy]
55096		J. H. Hutchison. 2013. New turtles from the Paleogene of North America. In D. B. Brinkman, P. A. Holroyd, J. D. Gardner (eds.), Morphology and Evolution of Turtles 477-497 [P. Holroyd/P. Holroyd]
76629		P. E. Kondrashov and S. G. Lucas. 2004. Arctocyon from the Paleocene of North America. In S. G. Lucas, K. E. Zeigler, and P. E. Kondrashov (eds.), Paleogene Mammals 11-20 [J. Marcot/J. Marcot]
14659		C. A. M. Meszoely, R. Estes, and H. Haubold. 1978. Eocene anguid lizards of Europe and a revision of the genus Xestops. Herpetologica 34(2):87-149 [R. Whatley/R. Whatley/M. Uhen]
2677		M. A. O'Leary and K. D. Rose. 1995. New mesonychian dentitions from the Paleocene of the Bighorn Basin, Wyoming. Annals of Carnegie Museum 64(2):147-172 [J. Alroy/J. Alroy/P. Mannion]
63104		R. Secord. 2008. The Tiffanian Land-Mammal Age (middle and late Paleocene) in the northern Bighorn Basin, Wyoming. University of Michigan Papers on Paleontology 35:1-192 [K. Beard/K. Beard]
3401		J. G. M. Thewissen. 1990. Evolution of Paleocene and Eocene Phenacodontidae (Mammalia, Condylarthra). University of Michigan Papers on Paleontology 29:1-107 [J. Alroy/J. Alroy/P. Mannion]