PBDB Collection

Mazon Creek, USNM Lacoe Collection: Westphalian D, Illinois
collected by R.D. Lacoe, Mr. Daniels 1865-1892

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Elonichthyiformes - Elonichthyidae

	Elonichthys hypsilepis Hay 1900	Hay 1900
	USNM 4848
Haplolepidae

	Pyritocephalus comptus Westoll 1944	Westoll 1944
	USNM 4310
Insecta

	Gyrophlebia longicollis n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	nomen dubium belonging to Insecta
	USNM 38150 ("near Cheliphlebia" in Scudder, 1885)

	"Palaeodictyopteron" mazonum n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	nomen dubium belonging to Insecta
	USNM 38831 (1 measurement)
Insecta - Geraridae

	Gerarus collaris Handlirsch 1911	Burnham 1983	1 specimen
	USNM 38835

	Gerarus vetus n. gen., n. sp. Scudder 1885	Handlirsch 1906	1 specimen
	(2 measurements)

	Gerarus mazonus n. sp. Scudder 1885	Scudder 1885	2 specimens
	F.T. Bliss coll, 1 uncertain (2 measurements)

	Genopteryx constricta n. gen., n. sp. Scudder 1885	Scudder 1885	1 specimen
	recombined as Gerarus constrictus
	USNM 38148 (1 measurement)

	Gerarus longus n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	synonym of Gerarus constrictus
	USNM 38822 (1 measurement)

	Gerarus angustus n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	synonym of Gerarus constrictus
	USNM 38811 (1 measurement)

	Genentomum validum n. gen., n. sp. Scudder 1885	Scudder 1885	1 specimen
	(1 measurement)

	Axiologus thoracicus n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38137
Insecta - Caloneurodea - Hapalopteridae

	Endoiasmus reticulatus n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38819
Insecta - Cnemidolestodea - Cnemidolestidae

	Aethophlebia singularis n. gen., n. sp. Scudder 1885	Scudder 1885	1 specimen
	(1 measurement)
Insecta - Cnemidolestodea - Gerapompidae

	Gerapompus blattinoides n. sp. Scudder 1885	Scudder 1885	1 specimen
	F.T. Bliss coll (1 measurement)

	Gerapompus schucherti n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38816 (1 measurement)

	Gerapompus extensus n. sp. Scudder 1885	Scudder 1885	1 specimen
	(1 measurement)
Insecta - Cnemidolestodea - Spanioderidae

	Dieconeura arcuata n. gen., n. sp. Scudder 1885	Scudder 1885	1 specimen
	(1 measurement)

	Spaniodera ambulans n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	synonym of Miamia bronsoni
	USNM 38817 (2 measurements)

	Metryia analis n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	synonym of Miamia bronsoni
	USNM 38834 (1 measurement)
Insecta - Cheliphlebiidae

	Cheliphlebia carbonaria n. gen., n. sp. Scudder 1885	Scudder 1885	1 specimen
	(1 measurement)
Insecta - Blattodea - Archimylacridae

	Paromylacris triangularis n. sp. Scudder 1895	Scudder 1895	1 specimen
	recombined as Oxynoblatta triangularis
	Coll Lacoe 2110 ab (1 measurement)

	Promylacris testudo Scudder 1890	Scudder 1895	1 specimen
	recombined as Metachorus testudo
	Coll Lacoe 2089a (1 measurement)
Insecta - Blattodea - Mylacridae

	Mylacris ampla n. sp. Scudder 1895	Scudder 1895	1 specimen
	Coll Lacoe 2021 ab (1 measurement)

	Dipeltis diplodiscus n. gen., n. sp. Packard 1885	Packard 1885	1 specimen
	recombined as Mylacris diplodiscus
	USNM 38864

	Mylacris antiquum n. sp. Scudder 1884	Scudder 1884	1 specimen
	corrected as Mylacris antiqua
	Coll Lacoe 2036 (1 measurement)

	Mylacris elongata n. sp. Scudder 1895	Scudder 1895	1 specimen
	synonym of Mylacris anthracophila
	Coll Lacoe 2014 ab ; collected by J.C. Carr (1 measurement)

	Paromylacris rotundum n. sp. Scudder 1885		1 specimen
	corrected as Paromylacris rotunda
	a.k.a. rotunda: R. D. Lacoe collection (1 measurement)

	Paromylacris ampla n. sp. Scudder 1890	Scudder 1890	1 specimen
	nomen dubium belonging to Mylacridae
	Lacoe 2088ab (2 measurements)

	Promylacris rigida n. sp. Scudder 1890	Scudder 1890	1 specimen
	recombined as Orthomylacris rigida
	Lacoe 2086ab (2 measurements)
Insecta - Blattodea - Gyroblattidae

	? Archoblattina scudderi n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38105 (Blattina sp. in Scudder, 1895, pl 12, fig 5) (1 measurement)
Insecta - Phyloblattidae

	Etoblattina hilliana n. sp. Scudder 1895	Scudder 1895	1 specimen
	recombined as Phyloblatta hilliana
	Coll Lacoe 2070 ab (1 measurement)
Insecta - Paoliida - Anthracoptilidae

	Adiphlebia lacoana n. gen., n. sp. Scudder 1885	Scudder 1885	1 specimen
	(2 measurements)

	Adiphlebia longitudinalis n. sp. (Scudder 1885)	Handlirsch 1906	1 specimen
	synonym of Adiphlebia lacoana
	USNM 38140 (2 measurements)
Insecta - Protorthoptera

	Chrestotes lapidea Scudder 1868	Handlirsch 1906	1 specimen
Insecta - Protorthoptera - Anthracothremmidae

	Anthracothremma robusta n. gen., n. sp. Scudder 1885	Scudder 1885	2 specimens
	(2 measurements)
Insecta - Protorthoptera

	Megalometer lata n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38825 (1 measurement)

	Danielsiella priscula n. gen., n. sp. Cockerell 1916	Cockerell 1916	1 specimen
	USNM 62000 (coll. Daniels) (1 measurement)

	Oryctoblattina occidua n. sp. Scudder 1885
	recombined as Agogoblattina occidua
	later made the type species of Agogoblattina by Handlirsch 1906 (1 measurement)

	Cheliphlebia elongata n. sp. Scudder 1885	Scudder 1885	1 specimen
	recombined as Metacheliphlebia elongata
	L.M. Umbach coll (1 measurement)
Insecta - Protorthoptera - Eucaenidae

	Eucaenus ovalis n. gen., n. sp. Scudder 1885	Scudder 1885	3 specimens
	USNM 38142; 38810, 38820 mentioned by Handlirsch 1906 (2 measurements)

	Eucaenus attenuatus Melander 1903	Handlirsch 1906	1 specimen
	synonym of Eucaenus ovalis
	USNM 38828 (1 measurement)
Insecta - Protorthoptera

	Polyetes furcifer n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38823 (1 measurement)

	Polyernus complanatus n. gen., n. sp. Scudder 1885	Scudder 1885	1 specimen
	(2 measurements)

	Archaeologus falcatus n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38818 (1 measurement)

	Pseudogerarus scudderi n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38151

	Eucaenus rotundatus n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	nomen dubium belonging to Protorthoptera
	USNM 38153 ("Neuropteroid Fam. Homothetidae" in Scudder, 1885) (1 measurement)
Insecta - Palaeodictyoptera

	Mammia alutacea n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38829 (1 measurement)
Insecta - Palaeodictyoptera - Lithomanteidae

	Acridites priscus Andrée 1864	Scudder 1885	1 specimen
	recombined as Catadyesthus priscus
Insecta - Palaeodictyoptera

	Palaiotaptus mazonus n. gen., n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	USNM 38815
Insecta

	Megathentomum pustulatum Scudder 1868	Scudder 1868	1 specimen
	USNM 38145 (Daniels 2025) (1 measurement)
	= Megathentomum scudderi n. sp. Handlirsch 1906	Handlirsch 1906
Myriapoda - Pleurojulida - Pleurojulidae

	Pleurojulus cf. biornatus Fritsch 1899	Wilson and Hannibal 2005	1 specimen
	USNM 37991
Myriapoda - Euphoberiida - Euphoberiidae

	Acantherpestes inequalis n. sp. Scudder 1890	Scudder 1890	2 specimens
	USNM PAL38042 (Lacoe 1819ab, 1819cd)

	Acantherpestes major (Meek and Worthen 1868)	Scudder 1890
	"several specimens" including Lacoe 1801de; one from W.F.E. Gurley

	Euphoberia sp. Meek and Worthen 1868	Scudder 1890	1 specimen
	Lacoe 1839a

	Euphoberia tracta n. sp. Scudder 1890	Scudder 1890	5 specimens
	USNM PAL37992 (Lacoe 1838ab, 1838cd, 1838ef, 1838gh, 1838ij)

	Euphoberia cuspidata n. sp. Scudder 1890	Scudder 1890	1 specimen
	USNM PAL38017 (Lacoe 1814ab)

	Euphoberia hystricosa n. sp. Scudder 1890	Scudder 1890	1 specimen
	USNM PAL38025 (Lacoe 1802ab)

	Euphoberia spinulosa n. sp. Scudder 1890	Scudder 1890
	USNM PAL37990 (Lacoe 1820a), Lacoe 1820cd, 1820ef, 1820gh

	Euphoberia anguilla Scudder 1882	Scudder 1890	1 specimen
	Lacoe 1807cd

	Euphoberia simplex n. sp. Scudder 1890	Scudder 1890	2 specimens
	USNM PAL38024 (Lacoe 1812ab)

	Euphoberia armigera Meek and Worthen 1868	Scudder 1890	4 specimens
	USNM PAL38019, PAL38020 (Lacoe 1804cd, 1804kl)

	Euphoberia granosa Scudder 1882	Scudder 1890
	Lacoe 1805h, 1805ij, 1805kl, 1805mn
Myriapoda - Xyloiulidae

	Xylobius mazonus n. sp. Scudder 1890	Scudder 1890	2 specimens
	recombined as Xyloiulus mazonus
	USNM PAL38036 (Lacoe 1809ab, 1809cd)

	Xylobius frustulentus n. sp. Scudder 1890	Scudder 1890	1 specimen
	recombined as Xyloiulus frustulentus
	USNM PAL37995 (Lacoe 1842ab)
Myriapoda

	? Archiulus glomeratus Scudder 1890	Scudder 1890	1 specimen
	Lacoe 1823cd

	Palenarthrus impressus n. gen., n. sp. Scudder 1890	Scudder 1890	1 specimen
	USNM PAL38002 (Lacoe 1821ab)

	Eileticus aequalis n. sp. Scudder 1890	Scudder 1890	2 specimens
	USNM PAL37996 (Lacoe 1836ab, 1836cd)

	Eileticus anthracinus Scudder 1882	Scudder 1890	1 specimen
	USNM PAL38004 (Lacoe 1835ab)
Myriapoda - Scutigeromorpha

	Latzelia primordialis n. gen., n. sp. Scudder 1890	Scudder 1890	1 specimen
	USNM PAL38000 (Lacoe 1837ab)
Xiphosura - Xiphosurida - Belinuridae

	Bellinurus lacoei n. sp. Packard 1885	Packard 1885	1 specimen
	recombined as Belinurus lacoei
Arachnida - Phalangiotarbida - Opiliotarbidae

	Architarbus elongatum n. sp. Scudder 1890	Scudder 1890	1 specimen
	recombined as Opiliotarbus elongatus
	Lacoe 1702ab (from Braidwood)
Arachnida - Phalangiotarbida

	Protopilio depressus n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	synonym of Phalangiotarbi
	USNM 37974
Arachnida - Phalangiotarbida - Architarbidae

	Geratarbus lacoei n. gen., n. sp. Scudder 1890	Scudder 1890	1 specimen
	Lacoe 1765ab (USNM 37966)

	Architarbus rotundatus Scudder 1868	Petrunkevitch 1913	8 specimens
	USNM1116,37962, 37969, 37980

	Geraphrynus carbonarius n. sp. Scudder 1884	Scudder 1884	1 specimen
	synonym of Architarbus rotundatus
	USNM 37961 (Lacoe 1701ab)
Arachnida - Ricinulei - Poliocheridae

	Polyochera glabra n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	recombined as Poliochera glabra
	USNM 37981

	Poliochera punctulata n. gen., n. sp. Scudder 1884	Scudder 1884	1 specimen
	USNM 37971 (Lacoe 1745)
Arachnida - Ricinulei - Curculioididae

	Geratarbus scabrum n. sp. Scudder 1890	Scudder 1890	1 specimen
	recombined as Curculioides scaber
	Lacoe 1766ab (USNM 37965)
Arachnida - Opiliones

	Kustarachne tenuipes n. gen., n. sp. Scudder 1890	Scudder 1890	1 specimen
	Lacoe 1756ab (USNM 37967)
Arachnida - Scorpiones - Eoscorpiidae

	Eoscorpius typicus n. sp. Petrunkevitch 1913	Petrunkevitch 1913	2 specimens
	synonym of Eoscorpius carbonarius
	USNM 37986, 37987

	Palaeopisthacanthus mazonensis n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	synonym of Eoscorpius carbonarius
	USNM 37977
Arachnida - Trigonotarbida - Trigonotarbidae

	Trigonotarbus carbonarius n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	USNM 37978
Arachnida - Trigonotarbida - Aphantomartidae

	Anthracomartus pustulatus n. sp. Scudder 1884	Scudder 1884	1 specimen
	recombined as Aphantomartus pustulatus
	Lacoe 1752 (USNM 37984)
Arachnida - Thelyphonida - Thelyphonidae

	Geralinura carbonaria n. sp. Scudder 1884	Scudder 1884	1 specimen
	USNM 37985 (Lacoe 1754a,b)

	Geralinura similis n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	synonym of Geralinura carbonaria
	USNM 37985 (other specimen of G. carbonaria Scudder)

	Geralinura gigantea n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	synonym of Geralinura carbonaria
	USNM 37976 (from Braidwood)
Arachnida - Amblypygi - Weygoldtinidae

	Graeophonus carbonarius (Scudder 1876)	Scudder 1890	1 specimen
	Lacoe 1762 (USNM 37969)
	= Graeophonus scudderi n. sp. Pocock 1911	Pocock 1911
	recombined as Weygoldtina scudderi
Cyclida - Americlidae

	Cyclus americanus n. sp. Packard 1885	Packard 1885	1 specimen
	recombined as Americlus americanus
unclassified

	Ilyodes divisa n. sp. Scudder 1890	Scudder 1890	1 specimen
	USNM PAL38034 (Lacoe 1830ab)

	Ilyodes elongata n. sp. Scudder 1890	Scudder 1890	1 specimen
	USNM PAL38035 (Lacoe 1829ab)
see common names

Geography

Country:	United States	State/province:	Illinois
Coordinates:	41.3° North, 88.4° West (view map)
Paleocoordinates:	4.6° South, 23.4° West
Basis of coordinate:	based on nearby landmark
Geographic resolution:	local area

Time

Period:	Carboniferous	Epoch:	Pennsylvanian
Stage:	Moscovian	10 m.y. bin:	Carboniferous 4
*Local age/stage:	Moscovian (minimum)
Key time interval:	Westphalian D
Age range of interval:	309.80000 - 307.40000 m.y. ago
* legacy (obsolete) database fields

Stratigraphy

Geological group:

Kewanee

Formation:

Carbondale

Member:

Francis Creek Shale

Lithology and environment

Primary lithology:	concretionary,sideritic lithified mudstone
Environment:	interdistributary bay	Tectonic setting:	cratonic basin
Geology comments: Illinois basin

Taphonomy

Modes of preservation:	mold/impression
Size of fossils:	macrofossils

Collection methods and comments

Reason for describing collection:	taxonomic analysis
Museum repositories:	USNM
Collectors:	R.D. Lacoe, Mr. Daniels	Collection dates:	1865-1892
Collection method comments: Mr. Lacoe of Pittston, Pennsylvania, paid local collectors, including Mr. Daniels, to collect specimens from Mazon Creek area. In the plant-bearing- beds of the anthracite and bituminous regions Mr. Lacoe occasionally secured a single insect wing, and when the finds became sufficient to warrant digging for them he would specially detail a collector to examine the shales of a given locality.

Metadata

Also known as:	ETE Locality 802
Database number:	21981
Authorizer:	C. Labandeira, M. Clapham, J. Flannery-Sutherland, D. Smith	Enterer:	C. Labandeira, A. Kinchloe, M. Clapham, J. Flannery-Sutherland, J. Karr
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	1998-06-09 00:00:00	Last modified:	2013-11-07 19:32:33
Access level:	the public	Released:	1998-06-09 00:00:00
Creative Commons license:	CC BY

Reference information

Primary reference:

35478.

S. H. Scudder. 1885. New genera and species of fossil cockroaches, from the older American rocks. Proceedings of the Academy of Natural Sciences of Philadelphia 37:34-39 [J. Alroy/J. Alroy]

Secondary references:

35773		O. Béthoux. 2005. Cnemidolestodea (Insecta): An ancient order reinstated. Journal of systematic Palaeontology 3(4):403-408 [M. Clapham/J. Karr]
35751		O. Béthoux. 2007. Emptying the Paleozoic wastebasket for insects: members of a Carboniferous 'protorthopterous family' assigned to natural groups. Alavesia 1:41-48 [M. Clapham/J. Karr]
39055		L. Burnham. 1983. Studies on Upper Carboniferous insects: 1. The Geraridae (Order Protorthoptera). Psyche 90:1-57 [M. Clapham/M. Clapham]
4983	ETE 695	F. M. Carpenter. 1965. Studies on North American Carboniferous Insects: 4. The Genera Metropator, Eubleptus, Hapaloptera & Hadentomum. Psyche 72(2):175-190 [C. Labandeira/C. Labandeira/M. Kosnik]
40318		T. D. A. Cockerell. 1916. Some American fossil insects. Proceedings of the United States National Museum 51:89-106 [M. Clapham/M. Clapham]
9323		J. A. Dunlop. 1995. Redescription of the Pennsylvanian trigonotarbid arachnid Lissomartus Petrunkevitch 1949 from Mazon Creek, Illinois. The Journal of Arachnology 23(2):118-124 [D. Smith/A. Kinchloe/M. Clapham]
35209		A. Handlirsch. 1906. Die Fossilen Insekten und die Phylogenie der Rezenten Formen, parts I-IV. Ein Handbuch fur Palaontologen und Zoologen 1-640 [M. Clapham/M. Clapham/M. Clapham]
37094		A. Handlirsch. 1906. Revision of American Paleozoic insects. Proceedings of the United States National Museum 29(1441):661-820 [M. Clapham/M. Clapham]
87132		O. Hay. 1900. Descriptions of some vertebrates of the Carboniferous age. Proceedings American Philosophical Society, Philadelphia 39:96-123 [J. Flannery-Sutherland/J. Flannery-Sutherland]
5092	ETE 804	J. Kukalová-Peck. 1985. Ephemeroid wing venation based upon new gigantic Carboniferous mayflies and basic morphology, phylogeny, and metamorphosis of pterygote insects (Insecta, Ephemerida). Canadian Journal of Zoology 63(4):933-955 [C. Labandeira/C. Labandeira/M. Clapham]
48447		A. S. Packard. 1885. Types of Carboniferous Xiphosura new to North America. The American Naturalist 19:291-294 [M. Clapham/M. Clapham]
9398		A. I. Petrunkevitch. 1913. A monograph of the terrestrial Palaeozoic Arachnida of North America. Transactions of the Connecticut Acadamy of Arts and Sciences 18:1-137 [D. Smith/A. Kinchloe/M. Clapham]
47423		R. I. Pocock. 1911. A Monograph of the Terrestrial Carboniferous Arachnida of Great Britain 1-84 [M. Clapham/M. Clapham/M. Clapham]
36688		S. H. Scudder. 1868. Remarks on two new fossil Insects from the Carboniferous formation in America. Proceedings of the Boston Society of Natural History 11:401-403 [M. Clapham/J. Karr/J. Karr]
50024		S. H. Scudder. 1884. The species of Mylacris, a Carboniferous genus of cockroaches. Memoirs of the Boston Society of Natural History 3:299-309 [M. Clapham/M. Clapham]
49579		S. H. Scudder. 1884. A contribution to our knowledge of Paleozoic Arachnida. Proceedings of the American Academy of Arts and Sciences 20:15-22 [M. Clapham/M. Clapham]
37105		S. H. Scudder. 1885. Palaeodictyoptera; or the affinities and classification of Paleozoic Hexapoda. Memoirs of the Boston Society of Natural History 3(11):319-351 [M. Clapham/M. Clapham/J. Karr]
47427		S. H. Scudder. 1890. Illustrations of the Carboniferous Arachnida of North America, of the orders Anthracomarti and Pedipalpi. Memoirs of the Boston Society of Natural History 4:443-456 [M. Clapham/M. Clapham]
39059		S. H. Scudder. 1890. New types of cockroaches from the Carboniferous deposits of the United States. Memoirs of the Boston Society of Natural History 4(9):401-415 [M. Clapham/M. Clapham]
60558		S. H. Scudder. 1890. New Carboniferous Myriapoda from Illinois. Memoirs of the Boston Society of Natural History 4:417-442 [M. Clapham/M. Clapham]
40799		S. H. Scudder. 1895. Revision of the American fossil cockroaches with descriptions of new forms. Bulletin of the United States Geological Survey 124:1-176 [M. Clapham/J. Karr]
48817		P. A. Selden. 1992. Revision of the fossil ricinuleids. Transactions of the Royal Society of Edinburgh: Earth Sciences 83:595-634 [M. Clapham/M. Clapham]
49581		O. E. Tetlie and J. A. Dunlop. 2008. Geralinura carbonaria (Arachnida: Uropygi) from Mazon Creek, Illinois, USA, and the origin of subchelate pedipalps in whip scorpions. Journal of Paleontology 82:299-312 [M. Clapham/M. Clapham]
84231		T. S. Westoll. 1944. The Haplolepidae, a new family of late Carboniferous bony fishes : a study in taxonomy and evolution. Bulletin of the AMNH ; v. 83, article 1 . Bulletin of the American Museum of Natural History 83(1):1-122 [E. Dunne/S. Fasey]
51029		H. M. Wilson and J. T. Hannibal. 2005. Taxonomy and trunk-ring architecture of pleurojulid millipedes (Diplopoda: Chilognatha: Pleurojulida) from the Pennsylvanian of Europe and North America. Journal of Paleontology 79:1105-1119 [M. Clapham/M. Clapham]