Khermeen Tsav II, Middle Red Beds [PMPE]: Middle Campanian - Late/Upper Campanian, Mongolia
collected by PMPE 1970-1971

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Myriapoda - Spirobolida
Diplopoda indet. Gervais 1844
3 specimens
    = Gobiulus sabulosus Dzik 1975
Dzik 1975
ZPAL MgMyr/1, 2, 5
Mammalia - Deltatheroida - Deltatheridiidae
Deltatheridium pretrituberculare Gregory and Simpson 1926
Kielan-Jaworowska 1978 2 specimens
type of "tardum" subspecies; possible incorrectly spelled "pretrituberculare" [check ref. 10462]
Mammalia - Asioryctidae
Asioryctes nemegtensis Kielan-Jaworowska 1975
Kielan-Jaworowska 1975 1 specimen
Mammalia - Zalambdalestidae
Barunlestes butleri Kielan-Jaworowska 1975
Kielan-Jaworowska 1975 4 specimens
ZPAL MgM-I/90 (partial right mandible), I/94 (partial right and left mandibles), I/104 (craniofacial portion of skull with mandibles, humerus and scapula), I/107 (partial skull)
Mammalia - Multituberculata
Nemegtbaatar gobiensis n. gen., n. sp. Kielan-Jaworowska 1974
Kielan-Jaworowska 1974
ZPAL MgM-I/81 (holotype partial skeleton)
Mammalia - Multituberculata - Djadochtatheriidae
Djadochtatherium catopsaloides Kielan-Jaworowska 1974
Kielan-Jaworowska 1974 1 specimen
recombined as Catopsbaatar catopsaloides
skull with partial lower jaws
Mammalia - Multituberculata - Sloanbaataridae
Nessovbaatar multicostatus n. gen., n. sp. Kielan-Jaworowska and Hurum 1997
Kielan-Jaworowska and Hurum 1997 1 specimen
ZPAL MgM-I/103
Mammalia - Multituberculata
Chulsanbaatar vulgaris Kielan-Jaworowska 1974
Kielan-Jaworowska 1974
Reptilia - Polyglyphanodontidae
Erdenetesaurus robinsonae n. gen., n. sp. Sulimski 1975
Sulimski 1975 2 specimens
ZPAL MgR-III/19, 22
Adamisaurus magnidentatus Sulimski 1972
Sulimski 1972 10 individuals
ZPAL MgR-III/1-5, 14, 23, 25-27 (partial-fragmentary skulls)
Reptilia - Slavoiidae
Slavoia darevskii Sulimski 1984
Sulimski 1984 10 specimens
Reptilia
Gobiderma pulchra n. gen., n. sp. Borsuk-Bialynicka 1984
Borsuk-Bialynicka 1984 3 specimens
ZPAL MgR-III/64 (holotype), skull with mandible; ZPAL MgR-III/65, damaged skull roof with frag. right mandible and thick skull osteoderms; ZPAL MgR-III/66, damaged skull and mandibles
Reptilia - Varanidae
Cherminotus longifrons n. gen., n. sp. Borsuk-Bialynicka 1984
Borsuk-Bialynicka 1984 2 individuals
ZPAL MgR-III/59 (holotype), damaged skull; ZPAL MgR-III/67, damaged skull
Reptilia - Protoceratopsidae
? Protoceratopsidae indet. Granger and Gregory 1923
Sabath 1991 8 specimens
ZPAL MgOv-III/1-8, 19; eggs
Protoceratops sp. Granger and Gregory 1923
Reptilia - Spheroolithidae
? Spheroolithidae indet. Zhao 1979
Sabath 1991
ZPAL MgOv-III/18, 17
Reptilia - Ankylosauridae
Ankylosauridae indet. Brown 1908
1 specimen
ZPAL MdG I/114
    = Saichania chulsanensis Maryanska 1977
Maryanska 1977
Reptilia - Dendroolithidae
Dinosauria indet. Owen 1842
Gradzinski and Jerzykiewicz 1972 10 specimens
eggs
    = Dendroolithidae indet. Zhao and Li 1988
Sabath 1991
ZPAL MgOv-III/17, 18a-c, 20
Reptilia - Faveoloolithidae
Faveoloolithidae indet. Zhao and Ding 1976
Sabath 1991
ZPAL MgOv-III/18b
    = Faveoloolithus ningxiaensis Zhao and Ding 1976
Mikhailov et al. 1994
Amphibia - Temnospondyli - Gobiatidae
Eopelobates leptocolaptus n. sp. Borsuk-Bialynicka 1978
Borsuk-Bialynicka 1978 1 specimen
recombined as Gobiates leptocolaptus
ZPAL MgAb-III/l
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 43.3° North, 99.8° East (view map)
Paleocoordinates:40.3° North, 89.4° East
Basis of coordinate:estimated from map
Altitude:1200 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
*Period:Late/Upper Cretaceous *Epoch:Senonian
*International age/stage:Middle Campanian
Key time interval:Middle Campanian - Late/Upper Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Baruungoyot
Stratigraphic resolution:bed
Stratigraphy comments: also "Red Beds of Khermeen Tsav"
Lithology and environment
Primary lithology:tabular,planar lamination,concretionary,red poorly lithified silty,calcareous sandstone
Includes fossils?Y
Lithology description: vivid red to orange, poorly cemented sandstone, with sandy concretions of variable size, small, white calcareous concretions occur. Some silt and clay intercalations. Cross-stratification is rare, but horizontally layered and massive beds are common. Common trace fossils - sediment-filled tubes - within the Red Beds. "In the profile Khermeen Tsav II the thick-bedded sandstones are orange-coloured and numerous intercalations of light-coloured (white, yellow, pale-red, and bluish-green) thin-bedded siltstones and claystones are present." (10453)
Environment:terrestrial indet.
Geology comments: identical description to Djadokhta Fm. (same ref)
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Spatial orientation:life position
Preservation of anatomical detail:excellent
Articulated whole bodies:some
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:occasional
Temporal resolution:snapshot
Spatial resolution:autochthonous
Collection methods and comments
Collection excludes:some genera
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:PMPE Collection dates:1970-1971
Collection method comments: five specimens found at this locality in addition to type specimen
Metadata
Also known as:Polish-Mongolian Palaeontological Expedition, Khermin Tsav
Database number:28243
Authorizer:J. Alroy, P. Barrett, R. Benson, J. Head, M. Carrano Enterer:E. Leckey, J. Head, M. Carrano, T. Cleary, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:2003-02-03 11:14:33 Last modified:2023-09-26 12:49:25
Access level:the public Released:2003-02-03 11:14:33
Creative Commons license:CC BY
Reference information

Primary reference:
9899. Z. Kielan-Jaworowska and R. Barsbold. 1972. Narrative of the Polish-Mongolian Palaeontological Expeditions 1967-1971. Palaeontologia Polonica 27:5-136 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

38112 V. R. Alifanov. 2000. The fossil record of Cretaceous lizards from Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 368-389 [R. Benson/R. Benson/M. Carrano]
66054 V. M. Arbour, P. J. Currie, and D. Badamgarav. 2014. The ankylosaurid dinosaurs of the Upper Cretaceous Baruungoyot and Nemegt formations of Mongolia. Zoological Journal of the Linnean Society 172(3):631-652 [G. Lloyd/G. Lloyd/M. Carrano]
64040 V. M. Arbour and P. J. Currie. 2016. Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs. Journal of Systematic Palaeontology 14(5):385-444 [G. Lloyd/G. Lloyd/M. Carrano]
100875% 34000M. Borsuk-Bialynicka. 1978. Eopelobates leptocolaptus sp. n.- the first upper Cretaceous pelobatid frog from Asia. Palaeontologia Polonica 38:57-63 [J. Head/J. Head/J. Head]
10439 M. Borsuk-Bialynicka. 1984. Anguimorphans and related lizards from the Late Cretaceous of the Gobi Desert, Mongolia. Palaeontologia Polonica 46:5-105 [M. Carrano/M. Carrano/M. Carrano]
14221ETE K. Carpenter and K. Alf. 1994. Global distribution of dinosaur eggs, nests, and babies. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 13-30 [M. Carrano/M. Carrano/M. Carrano]
77782 K. Carpenter, S. Hayashi, Y. Kobayashi, T. Maryanska, R. Barsbold, K. Sato, and I. Obata. 2011. Saichania chulsanensis (Ornithischia, Ankylosauridae) from the Upper Cretaceous of Mongolia. Palaeontographica, Abteilung A: Palaeozoology–Stratigraphy 294(13):1-61 [M. Carrano/M. Carrano/M. Carrano]
10453 R. Gradzinski and T. Jerzykiewicz. 1972. Additional geographical and geological data from the Polish-Mongolian Palaeontological Expeditions. Palaeontologia Polonica 27:17-306 [M. Carrano/M. Carrano/M. Carrano]
77615% 37520Z. Kielan-Jaworowska. 1974. Multituberculate succession in the Late Cretaceous of the Gobi Desert (Mongolia). Palaeontologia Polonica 30:23-44 [J. Alroy/E. Leckey/E. Leckey]
10442 Z. Kielan-Jaworowska. 1975. Preliminary description of two new eutherian genera from the Late Cretaceous of Mongolia. Palaeontologia Polonica 33:5-15 [M. Carrano/M. Carrano/M. Carrano]
10462 Z. Kielan-Jaworowska. 1978. Evolution of the therian mammals in the Late Cretaceous of Asia. Part III. Postcranial skeleton in Zalambdalestidae. Palaeontologia Polonica 58:5-41 [M. Carrano/M. Carrano/M. Carrano]
38300 Z. Kielan-Jaworowska and J. H. Hurum. 1997. Djadochtatheria - a new suborder of multituberculate mammals. Acta Palaeontologica Polonica 42(2):201-242 [R. Benson/R. Benson]
39263 Z. Kielan-Jaworowska and B. A. Trofimov. 1980. Cranial morphology of the Cretaceous eutherian mammal Barunlestes. Acta Palaeontologica Polonica 25(2):167-185 [R. Benson/R. Benson]
9909 T. Maryanska. 1977. Ankylosauridae (Dinosauria) from Mongolia. Palaeontologia Polonica 37:85-151 [M. Carrano/M. Carrano/M. Carrano]
14217ETE K. Mikhailov, K. Sabath, and S. Kurzanov. 1994. Eggs and nests from the Cretaceous of Mongolia. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 88-115 [M. Carrano/M. Carrano/M. Carrano]
41004 K. Sabath. 1991. Upper Cretaceous amniotic eggs from the Gobi Desert. Acta Palaeontologica Polonica 36(2):151-192 [M. Carrano/M. Carrano]
69215 M. A. Shishkin. 2000. Mesozoic amphibians from Mongolia and the Central Asiatic republics. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 297-308 [M. Carrano/M. Carrano]
10451 A. Sulimski. 1972. Adamisaurus magnidentatus n. gen., n. sp. (Sauria) from the Upper Cretaceous of Mongolia. Palaeontologia Polonica 27:33-40 [M. Carrano/M. Carrano/M. Carrano]
10460 A. Sulimski. 1975. Macrocephalosauridae and Polyglyphanodontidae (Sauria) from the Late Cretaceous of Mongolia. Palaeontologia Polonica 33:25-102 [M. Carrano/M. Carrano/M. Carrano]
10444 A. Sulimski. 1984. A new Cretaceous scincomorph lizard from Mongolia. Palaeontologia Polonica 46:143-155 [M. Carrano/M. Carrano/M. Carrano]